Abstract
Mexico is a megadiverse country, however its tardigrade fauna is poorly known and to date 44 species have been reported from this region. In the present paper, an update of the reported Mexican tardigrades is presented, increasing the number of recorded tardigrades to 55 species (11 new records). Moreover, we describe Milnesium cassandrae sp. nov. from State of Nuevo León in northern Mexico. The new species is most similar to Mil. krzysztofi Kaczmarek & Michalczyk, 2007 and Mil. beasleyi Kaczmarek, Jakubowska & Michalczyk, 2012, but it differs from them mainly by details of the dorsal sculpture and some morphometric characters. We also discuss the taxonomic value of a recently described feature, ‘pseudoplates’, for differentiating the taxa within the genus Milnesium Doyère, 1840. [Zoobank URL: urn:lsid:zoobank.org:act:0684AE18-0510-4F7B-B75D-AE5177FBF2A2]
References
Arcidiacono, R. (1962) Contributo alla conoscenza dei Tardigradi dei Monti Nebrodi e descrizione di una nuova specie di Itaquascon. Bollettino delle Sedute della Accademia Gioenia di Scienze Naturali in Catania, 3, 123–134.
Bartels, P.J. & Nelson, D.R. (2007) An evaluation of species richness estimators for tardigrades of the Great Smoky Mountains National Park, Tennessee and North Carolina, USA. Journal of Limnology, 66, 104–110.
https://doi.org/10.4081/jlimnol.2007.s1.104
Beasley, C.W. (1972) Some tardigrades from Mexico. The Southwestern Naturalist, 17, 21–29.
https://doi.org/10.2307/3669835
Beasley, C.W., Kaczmarek, Ł. & Michalczyk, Ł. (2008) Doryphoribius mexicanus, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from Mexico (North America). Proceedings of the Biological Society of Washington, 121, 34–40.
https://doi.org/10.2988/07-30.1
Bertolani, R., Bartels, P.J., Guidetti, R., Cesari, M. & Nelson, D.R. (2014a) Aquatic tardigrades in the Great Smoky Mountains National Park, North Carolina and Tennessee, U.S.A., with the description of a new species of Thulinius (Tardigrada, Isohypsibiidae). Zootaxa, 3764 (5), 524–536.
https://doi.org/10.11646/zootaxa.3764.5.2
Bertolani, R., Guidetti, R., Marchioro, T., Altiero, T., Rebecchi, L. & Cesari, M. (2014b) Phylogeny of Eutardigrada: New molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution, 76, 110–126.
https://doi.org/10.1016/j.ympev.2014.03.006
Binda, M.G. & Pilato, G. (1969) Ulteriore contributo alla conoscenza dei Tardigradi di Sicilia con descrizione di due nuove specie. Bollettino delle Sedute Dell’Accdemia Gioenia di Scienze Naturali in Catania, 10, 205–214.
Binda, M.G. & Pilato, G. (1986) Ramazzottius, nuova genere di Eutardigrado (Hypsibiidae). Animalia, 13, 159–166.
Ciobanu D.A., Zawierucha K., Moglan I., Kaczmarek Ł. (2014) Milnesium berladnicorum sp. n. (Eutardigrada, Apochela, Milnesiidae), a new species of water bear from Romania. ZooKeys, 429, 1–11.
https://doi.org/10.3897/zookeys.429.7755
Claps, M.C. & Rossi, G.C. (1988). Tardígrados de Argentina VI. Iheringia, 67, 3–11.
Claps, M.C. & Rossi, G.C. (2002). Tardigrada. In: Llorente–Bousquets, J. & Morrone, J.J. (Eds.), Biodiversidad, taxonomía y biogeografía de artrópodos de México: hacia una síntesis de su conocimiento. Vol. III. Universidad Nacional Autónoma de México, México, D.F., pp. 171–186.
Claxton, S.K. (1998) A revision of the genus Minibiotus (Tardigrada: Macrobiotidae) with descriptions of eleven new species from Australia. Records of the Australian Museum, 50, 125–160.
https://doi.org/10.3853/j.0067-1975.50.1998.1276
Da Cunha, A.X. & du Nascimento R.F. (1962) A fauna de tardígrados da Ilha da Madeira. Memórias e estudos do Museu Zoológico da Universidade de Coimbra, 279, 1–24.
Dastych, H. (1980) Niesporczaki (Tardigrada) Tatrzańskiego Parku Narodowego. Monografie Fauny Polski, 9, 1–232.
Dastych, H. (1988) The Tardigrada of Poland. Monografie Fauny Polski, 16, 1–255.
Dastych, H. (1992) Paradiphascon manningi gen. n. sp. n., a new water–bear from South Africa, with the erecting of a new subfamily Diphasconinae (Tardigrada). Mitteilungen aus dem Zoologischen Museum Hamburg, 89, 125–139.
Daza, A., Caicedo, M., Lisi, O. & Quiroga, S. (2017) New records of tardigrades from Colombia with the description of Paramacrobiotus sagani sp. nov. and Doryphoribius rosanae sp. nov. Zootaxa, 4362 (1), 29–50.
https://doi.org/10.11646/zootaxa.4362.1.2
De Barros, R. (1939) Pseudechiniscus juanitae nova espécie de Tardigrado. Boletim Biológico, New Series, 4, 367–368.
Degma, P., Bertolani, R. & Guidetti, R. (2018) Actual checklist of Tardigrada species. 2009–2018. 34th Edition. 30 June 2018. Available from: http://www.tardigrada.modena.unimo.it/miscellanea/Actual%20checklist%20of%20Tardigrada.pdf (accessed 4 October 2018)
Degma, P. & Guidetti, R. (2007) Notes to the current checklist of Tardigrada. Zootaxa, 1579 (1), 41–53.
https://doi.org/10.11646/zootaxa.1579.1.2
Doyère, L.M.F. (1840) Memoire sur les Tardigrades. Annales des Sciences Naturelles, Zoology, 14, 269–362.
Ehrenberg, C.G. (1848) Fortgesetzte Beobachtungen über jetzt herrschende atmospharische mikroscopische, etc. mit Nachtrag und Novarum Specierum Diagnosis. Akademie der Wissenschaften zu Berlin, 13, 370–381.
Fontoura, P. & Pilato, G. (2007) Diphascon (Diphascon) faialense sp. nov. a new species of Tardigrada (Eutardigrada, Hypsibiidae) from the Azores and a key to the species of the D. pingue group. Zootaxa, 1589 (1), 47–55.
https://doi.org/10.11646/zootaxa.1589.1.4
Fontoura, P., Pilato, G. & Lisi, O. (2008) New records of eutardigrades (Tardigrada) from Faial and Pico Islands, the Azores, with the description of two new species. Zootaxa, 1693 (1), 49–61.
https://doi.org/10.11646/zootaxa.1693.1.4
Guil, N., Jørgensen, A. & Kristensen, R. (2018) An upgraded comprehensive multilocus phylogeny of the Tardigrada tree of life. Zoologica Scripta, 48 (1), 1–18.
https://doi.org/10.1111/zsc.12321
Heinis, F. (1911) Beitrag zur Kenntnis der zentralamericanischen Moosfauna. Revue Suisse de Zoologie, 19, 253–266.
https://doi.org/10.5962/bhl.part.29917
Hinton, J.G., Meyer, H.A., McDaniel, N.D., Bergeron, C.B., Keely, S.J. & Matte, E. (2014) Lone star bears: The Tardigrada of Big Thicket National Preserve, Texas. Southeastern Naturalist, 13, 131–136.
https://doi.org/10.1656/058.013.s512
Hinton, J.G., Meyer, H.A., Soileau, B.N. & Dupuis, A.P. (2013) Tardigrada of the Caribbean island of Dominica (West Indies). Journal of Limnology, 72, 108–112.
https://doi.org/10.4081/jlimnol.2013.s1.e13
Ito, M. (1995) Taxonomic study on the Eutardigrada from the northern slope of Mt. Fuji, Central Japan II. Family Hypsibiidae. Proceedings of the Japanese Society of Systematic Zoology, 53, 18–39.
Ito, M. (1999) Ecological distribution, abundance and habitat preference of terrestrial tardigrades in various forests on the northern slope of Mt. Fuji, Central Japan. Zoologischer Anzeiger, 238, 225–234.
Kaczmarek, Ł. & Michalczyk, Ł. (2004) First record of the genus Doryphoribius Pilato, 1969 from Costa Rica (Central America) and description of a new species Doryphoribius quadrituberculatus (Tardigrada: Hypsibiidae). Genus, 15, 447–453.
Kaczmarek, Ł. & Michalczyk, Ł. (2007) A new species of Tardigrada (Eutardigrada: Milnesiidae): Milnesium krzysztofi from Costa Rica (Central America). New Zealand Journal of Zoology, 34, 297–302.
https://doi.org/10.1080/03014220709510088
Kaczmarek, Ł. & Michalczyk, Ł. (2009) Two new species of Macrobiotidae, Macrobiotus szeptyckii (harmsworthi group) and Macrobiotus kazmierskii (hufelandi group) from Argentina. Acta Zoologica Cracoviensia, 52, 87–99.
https://doi.org/10.3409/azc.52b_1-2.87-99
Kaczmarek, Ł. & Michalczyk, Ł. (2010) The genus Echiniscus Schultze 1840 (Tardigrada) in Costa Rican (Central America) rain forests with descriptions of two new species. Tropical Zoology, 23, 91–106.
Kaczmarek, Ł. & Roszkowska, M. (2016) A new eutardigrade from Costa Rica with taxonomical and zoogeographical remarks on Costa Rican tardigrades. New Zealand Journal of Zoology, 43, 234–245.
https://doi.org/10.1080/03014223.2016.1156553
Kaczmarek, Ł., Diduszko, D. & Michalczyk, Ł. (2011) New records of Mexican Tardigrada. Revista Mexicana de Biodiversidad, 82, 1324–1327.
Kaczmarek, Ł., Jakubowska, N. & Michalczyk, Ł. (2012) Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa, 3589 (1), 49–64.
https://doi.org/10.11646/zootaxa.3589.1.3
Kaczmarek, Ł., Michalczyk, Ł. & McInnes, S.J. (2014) Annotated zoogeography of non–marine Tardigrada. Part I: Central America. Zootaxa, 3763 (1), 1–62.
https://doi.org/10.11646/zootaxa.3763.1.1
Kaczmarek, Ł., Michalczyk, Ł. & McInnes, S.J. (2015) Annotated zoogeography of non–marine Tardigrada. Part II: South America. Zootaxa, 3923 (1), 1–107.
https://doi.org/10.11646/zootaxa.3923.1.1
Kaczmarek, Ł., Michalczyk, Ł. & McInnes, S.J. (2016) Annotated zoogeography of non–marine Tardigrada. Part III: North America and Greenland. Zootaxa, 4203 (1), 1–249.
https://doi.org/10.11646/zootaxa.4203.1.1
Kaczmarek, Ł, Gawlak, M., Bartels, P.J., Nelson, D.R. & Roszkowska, M. (2017) Revision of the genus Paramacrobiotus Guidetti et al., 2009 with the description of a new species, re–descriptions and a key. Annales Zoologici, 67 (4), 627–656.
https://doi.org/10.3161/00034541ANZ2017.67.4.001
Londoño, R., Daza, A., Lisi, O. & Quiroga, S. (2017) New species of waterbear Minibiotus pentannulatus (Tardigrada: Macrobiotidae) from Colombia. Revista Mexicana de Biodiversidad, 88 (4), 807–814.
https://doi.org/10.1016/j.rmb.2017.10.040
Londoño, R., Daza, A., Caicedo, M., Quiroga, S. & Kaczmarek, Ł. (2015) The genus Milnesium (Eutardigrada: Milnesiidae) in the Sierra Nevada de Santa Marta (Colombia), with the description of Milnesium kogui sp. nov. Zootaxa, 3955 (4), 561–568.
https://doi.org/10.11646/zootaxa.3955.4.7
Marcus, E. (1927) Zur Anatomie und Ökologie mariner Tardigraden. Zoologische Jahrbücher Abteilung für Systematik, Ökologie und Geographie der Tiere, 53, 487–558.
Marcus, E. (1936) Tardigrada. In: Schulze, F.E., Kükenthal, W. & Heider, K. (Eds.), Das Tierreich. Vol. 66. Walter de Gruyter, Berlin and Leipzig, pp. 1–340.
Martínez–Meyer, E., Sosa–Escalante, J.E. & Álvarez, F. (2014) El estudio de la biodiversidad en México: ¿Una ruta con dirección? Revista Mexicana de Biodiversidad, 85, 1–9.
https://doi.org/10.7550/rmb.43248
Maucci, W. & Ramazzotti, G. (1981) Cornechiniscus gen. nov.: nuova posizione sistematica per i cosiddetti “Pseudechiniscus gruppo cornutus” con descrizione di una nuova specie (Tardigrada, Echiniscidae). Memorie dell’Istituto Italiano di Idrobiologia, Pallanza, 39, 147–151.
May, R.M. (1948) Nouveau genre et espèce de tardigrade du Mexique: Haplomacrobiotus hermosillensis. Bulletin de la Société Zoologique de France, 73, 95–97.
McInnes, S.J. (1994) Zoogeographic distribution of terrestrial/freshwater tardigrades from current literature. Journal of Natural History, 28, 257–352.
https://doi.org/10.1080/00222939400770131
Mehlen, R.H. (1969) New Tardigrada From Texas. The American Midland Naturalist, 81, 395–404.
https://doi.org/10.2307/2423979
Melo, J.C., Beltrán–Pardo, E., Bernal, J.E. & Kaczmarek, Ł. (2015) New records of tardigrades from Colombia (Guatavita, Cundinamarca Department). Turkish Journal of Zoology, 39, 412–420.
https://doi.org/10.3906/zoo-1405-13
Meyer, H.A. (2013) Terrestrial and freshwater Tardigrada of the Americas. Zootaxa, 3747 (1), 1–71.
https://doi.org/10.11646/zootaxa.3747.1.1
Meyer, H.A. & Domingue, M.N. (2011) Minibiotus acadianus (Eutardigrada: Macrobiotidae), a new species of Tardigrada from southern Louisiana, U.S.A. Western North American Naturalist, 71, 38–43.
https://doi.org/10.3398/064.071.0106
Meyer, H.A. & Hinton, J.G. (2012) Terrestrial Tardigrada of the island of Barbados in the West Indies, with the description of Milnesium barbadosense sp. n. (Eutardigrada: Apochela: Milnesiidae). Caribbean Journal of Science, 46, 194–202.
https://doi.org/10.18475/cjos.v46i2.a8
Meyer, H.A., Domingue, M.N. & Hinton, J.G. (2014) Tardigrada of the West Gulf Coastal Plain, with descriptions of two new species from Louisiana. Southeastern Naturalist, 13, 117–130.
https://doi.org/10.1656/058.013.s511
Meyer, H.A., Hinton, J.G. & Dupré, M.C. (2013) Milnesium lagniappe, a new species of water bear (Tardigrada, Eutardigrada, Apochela, Milnesiidae) from the southern United States. Western North American Naturalist, 73, 295–301.
https://doi.org/10.3398/064.073.0305
Michalczyk, Ł. & Kaczmarek, Ł. (2010) Description of Doryphoribius dawkinsi, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from the Costa Rican highlands, with the key to the genus Doryphoribius. Zootaxa, 2393 (1), 46–58.
https://doi.org/10.11646/zootaxa.2393.1.4
Michalczyk, Ł. & Kaczmarek, Ł. (2013) The Tardigrada Register: a comprehensive online data repository for tardigrade taxonomy. Journal of Limnology, 72, 175–181.
https://doi.org/10.4081/jlimnol.2013.s1.e22
Michalczyk, Ł., Wełnicz, W., Frohme, M. & Kaczmarek, Ł. (2012a) Redescriptions of three Milnesium Doyère, 1840 taxa (Tardigrada: Eutardigrada: Milnesiidae), including the nominal species for the genus. Zootaxa, 3154 (1), 1–20.
https://doi.org/10.11646/zootaxa.3154.1.1
Michalczyk, Ł., Wełnicz, W., Frohme, M. & Kaczmarek, Ł. (2012b) Corrigenda of Zootaxa, 3154:1–20 Redescriptions of three Milnesium Doyère, 1840 taxa (Tardigrada: Eutardigrada: Milnesiidae), including the nominal species for the genus. Zootaxa, 3393 (1), 66–68.
https://doi.org/10.11646/zootaxa.3393.1.6
Miller, W.R. & Mehlen, R.H. (2007) Tardigrades of North America: re-description of Echiniscus tamus Mehlen, 1969 (Tardigrada: Heterotardigrada: Echiniscoidae: Echiniscidae). Proceedings of the Biological Society of Washington, 120, 184–188. https://doi.org/10.2988/0006-324X(2007)120[184:TONARO]2.0.CO;2
Miller, W.R., McInnes, S.J. & Bergstrøm, D.M. (2005) Tardigrades of the Australian Antarctic: Hypsibius heardensis (Eutardigrada: Hypsibiidae: dujardini group) a new species from sub–Antarctic Heard Island. Zootaxa, 1022 (1), 57–64.
https://doi.org/10.11646/zootaxa.1022.1.3
Morek, W., Gąsiorek, P., Stec, D., Blagden, B. & Michalczyk, Ł. (2016) Experimental taxonomy exposes ontogenetic variability and elucidates the taxonomic value of claw configuration in Milnesium Doyère, 1840 (Tardigrada: Eutardigrada: Apochela). Contributions to Zoology, 85, 173–200.
https://doi.org/10.1163/18759866-08502003
Morek, W., Stec, D., Gąsiorek, P., Surmacz, B. & Michalczyk, Ł. (2018) Milnesium tardigradum Doyère, 1840: The first integrative study of interpopulation variability in a tardigrade species. Journal of Zoological Systematic and Evolutionary Research, 57 (1), 1–23.
https://doi.org/10.1111/jzs.12233
Moreno-Talamantes, A., Roszkowska, M., Ríos, G.P., Flores, M.J.J. & Kaczmarek, Ł. (2015) First record of Dactylobiotus parthenogeneticus Bertolani, 1982 (Eutardigrada: Murrayidae) in Mexico. Check List, 11, 1723.
https://doi.org/10.15560/11.4.1723
Nelson, D.R. & Bartels, P.J. (2007) “Smoky Bears”—Tardigrades of the Great Smoky Mountains National Park. Southeastern Naturalist, 6, 229–238.
https://doi.org/10.1656/1528-7092(2007)6[229:SBOGSM]2.0.CO;2
Nelson, D.R. & Bartels, P.J. (2013) Species richness of soil and leaf litter tardigrades in the Great Smoky Mountains National Park (North Carolina/Tennessee, USA). Journal of Limnology, 72, 144–151.
https://doi.org/10.4081/jlimnol.2013.s1.e18
Pérez-Petch, W.A., Anguas–Escalante, A., Cutz-Pool, L.Q. & Guidetti, R. (2017) Doryphoribius chetumalensis sp. nov. (Eutardigrada: Isohypsibiidae) a new tardigrade species discovered in an unusual habitat of urban areas of Mexico. Zootaxa, 4344 (2), 345–456.
https://doi.org/10.11646/zootaxa.4344.2.9
Perry, E., Miller, W.R. & Kaczmarek, Ł (2019) Recommended abbreviations for the names of genera of the phylum Tardigrada. Zootaxa, 4608 (1), 145–154. http://doi.org/10.11646/zootaxa.4608.1.8
Pilato, G. (1969) Schema per una nuova sistemazione delle famiglie e dei genri degli Eutardigrada. Bollettino delle sedute dell’ Accademia Gioenia di Scienze naturale Catania, 10, 181–193.
Pilato, G. (1981) Analisi di nuovi caratteri nello studio degli Eutardigradi. Animalia, 8, 51–57.
Pilato, G. (1987) Revision of the genus Diphascon Plate, 1889, with remarks on the subfamily Itaquasconinae (Eutardigrada, Hypsibiidae). In: Bertolani, R. (Ed.), Biology of Tardigrades. Selected Symposia and Monographs U.Z.I. Vol. 1. Mucchi, Modena, pp. 337–357.
Pilato, G. (1997) Astatumen, a new genus of the Eutardigrada (Hypsibiidae, Itaquasconinae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 12, 205–208.
Pilato, G. (2006) Remarks on the Macrobiotus polyopus group, with the description of two new species (Eutardigrada, Macrobiotidae). Zootaxa, 47 (1), 37–47.
https://doi.org/10.11646/zootaxa.1298.1.4
Pilato, G. & Binda, M.G. (1991) Milnesium tetralamellatum, new species of Milnesiidae from Africa (Eutardigrada). Tropical Zoology, 4, 103–106.
https://doi.org/10.1080/03946975.1991.10539480
Pilato, G. & Binda, M.G. (1998) A comparison of Diphascon (D.) alpinum Murray, 1906, D. (D.) chilense Plate, 1889 and D. (D.) pingue Marcus, 1936 (Tardigrada), and description of a new species. Zoologischer Anzeiger, 263, 181–185.
Pilato, G. & Binda, M.G. (1999) Three new species of Diphascon of the pingue group (Eutardigrada, Hypsibiidae) from Antarctica. Polar Biology, 21, 335–342.
https://doi.org/10.1007/s003000050370
Pilato, G. & Binda, M.G. (2010) Definition of families, subfamilies, genera and subgenera of the Eutardigrada, and keys to their identification. Zootaxa, 2404 (1), 1–52.
https://doi.org/10.11646/zootaxa.2404.1.1
Pilato, G. & Lisi, O. (2006) Notes on some tardigrades from southern Mexico with description of three new species. Zootaxa, 1236 (1), 53–68.
https://doi.org/10.11646/zootaxa.1236.1.4
Pilato, G. & Lisi, O. (2016) Milnesium minutum and Milnesium sandrae, two new species of Milnesiidae (Tardigrada, Eutardigrada, Apochela). ZooKeys, 580, 1–12.
https://doi.org/10.3897/zookeys.580.6603
Pilato, G., Binda, M.G. & Lisi, O. (2002) Notes on tardigrades of the Seychelles with the description of two new species. Bollettino dell’Accademia Gioenia di Scienze Naturali, 35, 503–517.
Pilato, G., D’Urso, V. & Lisi, O. (2013) Ramazzottius thulini (Pilato, 1970) bona species and description of Ramazzottius libycus sp. nov. (Eutardigrada, Ramazzottidae). Zootaxa (3), 3681 (3), 270–280.
https://doi.org/10.11646/zootaxa.3681.3.6
Pilato, G., Sabella, G. & Lisi, O. (2016) Two new species of Milnesium (Tardigrada: Milnesiidae). Zootaxa, 4132 (4), 575–587.
https://doi.org/10.11646/zootaxa.4132.4.9
Pilato, G., Sabella, G., D’Urso, V. & Lisi, O. (2017) Two new species Eutardigrada from Victoria Land, Antartica. Zootaxa, 4317 (3), 541–558.
https://doi.org/10.11646/zootaxa.4317.3.6
Plate, L. (1888) Beiträge zur Naturgeschichte der Tardigraden. Zoologische Jahrbücher, 3, 487–550.
https://doi.org/10.5962/bhl.part.1265
Rahm, G. (1931) Tardigrada of the South of America (esp. Chile). Revista Chilena de Historia Natural, 35, 118–141.
Ramazzotti, G. (1943) Nuova varieta del Tardigrado Pseudechiniscus cornutus. Rivista di Scienze Naturali ‘Natura’, 34, 89–90.
Ramazzotti, G. (1957) Due nuove specie di Tardigradi extra–europei. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 96, 188–191.
Ramazzotti, G. (1962) Tardigradi del Cile, con descrizione di quattro nuove specie e di una nuova variet. Atti della Societa ltaliana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 101, 275–287.
Ramazzotti, G. & Maucci, W. (1983) Il Phylum Tardigrada. Memorie dell’Instituto Italiano di Idrobiologia, 41, 1–1012.
Richters, F. (1926) Tardigrada. In: Kükenthal, W. & Krumbach, T. (Eds.), Handbuch der Zoologie. Vol. 3. Walter de Gruyter & Co., Berlin & Leipzig, pp.1–68.
Roszkowska, M., Ostrowska, M. & Kaczmarek, Ł. (2015) The genus Milnesium Doyère, 1840 (Tardigrada) in South America with descriptions of two new species from Argentina and discussion of the feeding behaviour in the family Milnesiidae. Zoological Studies, 54, 1–17.
https://doi.org/10.1186/s40555-014-0082-7
Roszkowska, M., Stec, D., Ciobanu, D.A. & Kaczmarek, Ł. (2016) Tardigrades from Nahuel Huapi National Park (Argentina, South America) with descriptions of two new Macrobiotidae species. Zootaxa, 4105 (3), 243–260. https://doi.org/10.11646/zootaxa.4105.3.2
Roszkowska, M., Ostrowska, M., Stec, D., Janko, K. & Kaczmarek, Ł. (2017) Macrobiotus polypiformis, a new tardigrade (Macrobiotidae; hufelandi group) from the Ecuadorian Pacific coast, with remarks on the claw abnormalities in eutardigrades. Journal of European Taxonomy, 327, 1–19.
https://doi.org/10.5852/ejt.2017.327
Roszkowska, M., Stec, D., Gawlak, M. & Kaczmarek, Ł. (2018) An integrative description of a new tardigrade species Mesobiotus romani sp. nov. (Macrobiotidae; harmsworthi group) from the Ecuadorian Pacific coast. Zootaxa, 4450 (5), 550–564.
https://doi.org/10.11646/zootaxa.4450.5.2
Rudescu, L. (1964) Tardigrada. Fauna Republicii Populare Romine. Bucuresti, 4, 1–398.
Sands, C.J., McInnes, S.J., Marley, N.J., Goodall-Copestake, W., Convey, P. & Linse, K. (2008) Phylum Tardigrada: an “individual” approach. Cladistics, 24, 1–18.
https://doi.org/10.1111/j.1096-0031.2008.00219.x
Schlabach, S., Donaldson, E., Hobelman K., Miller, W.R. & Lowman, M. (2018) Tardigrades of the Canopy; Milnesium burgessi nov. sp. (Eutardigrada: Apochela: Milnesiidae) a new species from Kansas, U.S.A. Transactions of the Kansas Academy of Science, 121, 39–48.
https://doi.org/10.1660/062.121.0204
Schultze, C.A.S. (1834) Macrobiotus Hufelandii animal e crustaceorum classe novum, reviviscendi post diuturnam asphixiam et aridiatem potens. Berolini, Apud Carolum Curths, 1833, 1–7.
Schultze, C.A.S. (1840) Echiniscus Bellermanni; animal crustaceum. Apud G. Reimer, Berolini, 8 pp.
Schuster, R.O. (1971) Tardigrada from the Barranca del Cobre Sinaloa and Chihuahua, Mexico. Proceedings of the Biological Society of Washington, 84, 213–224.
Schuster, R.O., Nelson, D.R., Grigarick, A.A. & Christenberry, D. (1980) Systematic criteria of Eutardigrada. Transactions of the American Microscopical Society, 99, 284–303.
https://doi.org/10.2307/3226004
Sierra J., C.L., Sosa R., J., Cortés–Calva, P., Solís, C.A.B., Íñiguez, D.L.I. & Ortega-Rubio, A. (2014) México país megadiverso y la relevancia de las áreas naturales protegidas. Investigación y Ciencia, 60, 16–22.
Spiers, A.I., Haefke, B.J., Miller, W.R. & Lowman, M.D. (2013) Tardigrades in the canopy: Doryphoribius dawkinsi Michalczyk and Kaczmarek, 2010—New records from eastern Kansas, U.S.A. Kansas Academy of Science, Transactions, 116, 125–128.
Stec, D., Smolak, R., Kaczmarek, Ł. & Michalczyk, Ł. (2015) An integrative description of Macrobiotus paulinae sp. nov. (Tardigrada: Eutardigrada: Macrobiotidae: hufelandi group) from Kenya. Zootaxa, 4052 (5), 501–526.
https://doi.org/10.11646/zootaxa.4052.5.1
Stec, D., Morek, W., Gąsiorek, P. & Michalczyk, Ł. (2018) Unmasking hidden species diversity within the Ramazzottius oberhaeuseri complex, with an integrative redescription of the nominal species for the family Ramazzottiidae (Tardigrada: Eutardigrada: Parachela), Systematics and Biodiversity, 16 (4), 357–376.
https://doi.org/10.1080/14772000.2018.1424267
Stec, D., Roszkowska, M., Kaczmarek, Ł. & Michalczyk, Ł. (2018) Paramacrobiotus lachowskae, a new species of Tardigrada from Colombia (Eutardigrada: Parachela: Macrobiotidae). New Zealand Journal of Zoology, 45 (1), 43–60.
https://doi.org/10.1080/03014223.2017.1354896
Thulin, G. (1911) Beitrag zur Kenntnis der Tardigradenfauna Schwedens. Arkiv for Zoologi, 7, 1–60.
https://doi.org/10.5962/bhl.part.1270
Thulin, G. (1928) Über die Phylogie und das System der Tardigraden. Heriditas, 11, 207–266.
https://doi.org/10.1111/j.1601-5223.1928.tb02488.x
Tibbs, L., Emanuels, A. & Miller, W.R. (2016) Tardigrades of the canopy: Argentine species Milnesium beatae Roszkowska, Ostrowska and Kaczmarek, 2015 (Eutardigrada, Milnesidae) discovered in the trees of Kansas, U.S.A. Transactions of the Kansas Academy of Science, 119 (2), 173–178.
https://doi.org/10.1660/062.119.0207
Tumanov, D.V. (2006) Five new species of the genus Milnesium (Tardigrada, Eutardigrada, Milnesiidae) Zootaxa, 1122 (1), 1–23.
https://doi.org/10.11646/zootaxa.1122.1.1
Urbanowicz, C. (1925) Sur la variabilité de Macrobiotus oberhaeuseri. Bulletin biologique de la France et de la Belgique, Paris, 59, 124–142.
Węglarska, B. (1959) Tardigraden Polens II. Věstník Československé společnosti Zoologické, 23, 354–357.
Young, A., Chappell, B., Miller, W.R. & Lowman, M.D. (2016) Tardigrades of the tree Canopy: Milnesium swansoni sp. nov. (Eutardigrada: Apochela: Milnesiidae) a new species from Kansas, U.S.A. Zootaxa, 4072 (5), 559–568.
https://doi.org/10.11646/zootaxa.4072.5.3
Zawierucha, K., Dziamięcki, J., Jakubowska, N., Michalczyk, Ł. & Kaczmarek, Ł. (2014) New tardigrade records for the Baltic states with a description of Minibiotus formosus sp. n. (Eutardigrada, Macrobiotidae). ZooKeys, 408, 81–105.