Skip to main content Skip to main navigation menu Skip to site footer
Type: Monograph
Published: 2020-05-12
Page range: 1–157
Abstract views: 409
PDF downloaded: 214

The marine ichthyofauna of Lebanon: an annotated checklist, history, biogeography, and conservation status

Department of Biology, American University of Beirut, 11-0236 Beirut, Lebanon.
Im Ramstal 76, 97922 Lauda-Königshofen, Germany.
Pisces fishes new records non-indigenous species taxonomy distribution Levant Mediterranean Sea

Abstract

This is an annotated checklist of the marine fish species of Lebanon (Levant coast, eastern Mediterranean). It comprises a total of 367 fish species distributed in 159 families and 27 orders, out of which 70 species are non-indigenous and 28 are recorded for the first time from Lebanon. The checklist includes all names and citations of species found in the literature, as well as references to specimens available in museum collections. It also includes an updated zoogeographic distribution and conservation status for each species. Threats were estimated based on surveys and observations made over the last two decades; broadly categorized conservation actions were also suggested.

 

References

  1. Aguilar, R., Garcia, S., Perry, A.L., Alvarez, H., Blanco, J. & Bitar, G. (2018) 2016 Deep-sea Lebanon Expedition: Exploring Submarine Canyons. Oceana, Madrid, 94 pp.

    Ali, M. (2018) An updated checklist of the marine fishes from Syria with emphasis on alien species. Mediterranean Marine Science, 19 (2), 388–393.

    https://doi.org/10.12681/mms.15850

    Badreddine, A. & Bitar, G. (2019) First record of Heteropriacanthus cruentatus (Lacepède 1801) (Chordata: Priacanthidae) in the Mediterranean Sea from the Lebanese waters. Journal of the Black Sea/Mediterranean Environment, 25 (2), 178–181.

    Ballard, E. (1934) Note on certain problems concerning the variation and abundance of some Palestinian sea-fishes. In: Commission Internationale pour l’Exploration scientifique de la mer Méditerranée (CIESM), Rapport et Procès-Verbaux des Réunions, 8, pp. 31–35.

    Bariche, M. (2005) Age and growth of Lessepsian rabbitfish from the eastern Mediterranean. Journal of Applied Ichthyology, 21 (2), 141–145.

    https://doi.org/10.1111/j.1439-0426.2004.00619.x

    Bariche, M. (2010a) Champsodon vorax (Teleostei: Champsodontidae), a new alien fish in the Mediterranean. aqua, International Journal of Ichthyology, 16 (4), 197–200.

    Bariche, M. (2010b) First record of the angelfish Pomacanthus maculosus (Teleostei: Pomacanthidae) in the Mediterranean. aqua, International Journal of Ichthyology, 16 (1), 31–33.

    Bariche, M. (2011) First record of the cube boxfish Ostracion cubicus (Ostraciidae) and additional records of Champsodon vorax (Champsodontidae) from the Mediterranean. aqua, International Journal of Ichthyology, 17 (4), 181–184.

    Bariche, M. (2012) Recent evidence on the presence of Heniochus intermedius (Teleostei: Chaetodontidae) and Platycephalus indicus (Teleostei: Platycephalidae) in the Mediterranean Sea. BioInvasions Records, 1 (1), 53–57.

    https://doi.org/10.3391/bir.2012.1.1.12

    Bariche, M. (2018) First confirmed record of the white-spotted puffer Arothron hispidus (Linnaeus 1758) in the Mediterranean Sea. BioInvasions Records, 7 (4), 433–436.

    https://doi.org/10.3391/bir.2018.7.4.13

    Bariche, M. & Azzurro, E. (2012) New records and establishment of the Indian Ocean twospot cardinalfish Cheilodipterus novemstriatus (Rüppell 1838) in the Mediterranean Sea. BioInvasions Records, 1 (4), 299–301.

    https://doi.org/10.3391/bir.2012.1.4.10

    Bariche, M. & Bernardi, G. (2009) Lack of a genetic bottleneck in a recent Lessepsian bioinvader, the blue-barred parrotfish, Scarus ghobban. Molecular Phylogenetics and Evolution, 53, 592–595.

    https://doi.org/10.1016/j.ympev.2009.06.017

    Bariche, M. & Di Natale, A. (2019) Preliminary information on the Atlantic bluefin tuna (Thunnus thynnus) fishery in Lebanon. Collective Volumes of Scientific Papers, ICCAT, 75 (6), 1278–1281.

    Bariche, M. & Fricke, R. (2018) Dipterygonotus balteatus (Valenciennes 1830) (Teleostei: Caesionidae), a new alien fish in the Mediterranean Sea. BioInvasions Records, 7 (1), 79–82.

    https://doi.org/10.3391/bir.2018.7.1.12

    Bariche, M. & Heemstra, P. (2012) First record of the blacktip grouper Epinephelus fasciatus (Teleostei: Serranidae) in the Mediterranean Sea. Marine Biodiversity Records, 5 (e1), 1–3.

    https://doi.org/10.1017/S1755267211000509

    Bariche, M. & Saad, M. (2008) Settlement of the lessepsian blue-barred parrotfish Scarus ghobban (Teleostei: Scaridae) in the eastern Mediterranean. Marine Biodiversity Records, 1 (e5), 1–3.

    https://doi.org/10.1017/S1755267205000497

    Bariche, M., Alwan, N., El-Assi, H. & Zurayk, R. (2009) Diet composition of the Lessepsian bluespotted cornetfish Fistularia commersonii in the eastern Mediterranean. Journal of Applied Ichthyology, 25 (4), 460–465.

    https://doi.org/10.1111/j.1439-0426.2008.01202.x

    Bariche, M., Torres, M. & Azzurro, E. (2013a) The presence of the invasive lionfish Pterois miles in the Mediterranean Sea. Mediterranean Marine Science, 14 (2), 292–294.

    https://doi.org/10.12681/mms.428

    Bariche, M., Bilecenoglu, M. & Azzurro, E. (2013b) Confirmed presence of the Red Sea goatfish Parupeneus forsskali (Fourmanoir & Guézé 1976) in the Mediterranean Sea. BioInvasions Records, 2 (2), 173–175.

    https://doi.org/10.3391/bir.2013.2.2.15

    Bariche, M., Harmelin-Vivien, M. & Quignard, J.P. (2003) Reproductive cycles and spawning periods of two Lessepsian siganid fishes on the Lebanese coast. Journal of Fish Biology, 62 (1), 129–142.

    https://doi.org/10.1046/j.0022-1112.2003.00014.x

    Bariche, M., Kazanjian, G. & Azzurro, E. (2014) A lag of 25 years: evidence from an old capture of Fistularia commersonii Rüppell 1838 from Lebanon (Mediterranean Sea). Journal of Applied Ichthyology, 30 (1–2), 535–536.

    https://doi.org/10.1111/jai.12394

    Bariche, M., Kleitou, P., Kalogirou, S. & Bernardi, G. (2017) Genetics reveal the identity and origin of the lionfish invasion in the Mediterranean Sea. Scientific Reports, 7 (1), 6782.

    https://doi.org/10.1038/s41598-017-07326-1

    Bariche, M., Letourneur Y. & Harmelin-Vivien M. (2004) Temporal fluctuations and settlement patterns of native and Lessepsian herbivorous fishes on the Lebanese coast (eastern Mediterranean). Environmental Biology of Fishes, 70 (1), 81–90.

    https://doi.org/10.1023/B:EBFI.0000022928.15148.75

    Bariche, M., Sadek, R., Al-Zein, M.S. & El-Fadel, M. (2007) Diversity of juvenile fish assemblages in the pelagic waters of Lebanon (eastern Mediterranean). Hydrobiologia, 580, 109–115.

    https://doi.org/10.1007/s10750-006-0461-0

    Bariche, M., Torres, M., Smith, C., Sayar, N., Azzurro, E., Baker, R. & Bernardi, G. (2015) Red Sea fishes in the Mediterranean Sea: a preliminary investigation of a biological invasion using DNA barcoding. Journal of Biogeography, 42 (12), 2363–2373.

    https://doi.org/10.1111/jbi.12595

    Belmaker, J., Parravicini, V. & Kulbicki, M. (2013) Ecological traits and environmental affinity explain Red Sea fish introduction into the Mediterranean. Global Change Biology, 19 (5), 1373–1382.

    https://doi.org/10.1111/gcb.12132

    Ben-Tuvia, A. (1953a) Fishes caught off Caesarea, on the Mediterranean coast of Israel. Bulletin of the Research Council of Israel, 2 (4), 439–440.

    Ben-Tuvia, A. (1953b) Mediterranean fishes of Israel. Bulletin of the Sea Fisheries Research Station, Haifa, 8, 1–40.

    Bitar, G. (2013) Sur la présence des poissons exotiques nouveaux de la côte libanaise (Méditerranée orientale). Rapport du Congrès de la Commission Internationale Pour l’Exploration Scientifique de la Mer Méditerranée, 40, 592.

    Bodenheimer, F.S. (1935) Animal life in Palestine. An introduction to the problems of animal ecology and zoogeography. L. Mayer, Jerusalem, 506 pp.

    Bodenheimer, F.S. (1937) Prodomus faunae Palestinae. Mémoires de l’Institut d’Égypte, 233, 271–275.

    Boulos, I. (1961) Océanographie et biologie des pêches au Liban. Université Libanaise, Beyrouth, 50 pp.

    Carpentieri, P., Lelli, S., Colloca, F., Mohanna, C., Bartolino, V., Moubayed, S. & Ardizzone, G.D. (2009) Incidence of lessepsian migrants on landings of the artisanal fishery of south Lebanon. Marine Biodiversity Records, 2, e71.

    https://doi.org/10.1017/S1755267209000645

    CITES, UNEP-WCMC (2017) The Checklist of CITES Species Website. Appendices I, II and III valid from 04 April 2017. CITES Secretariat, Geneva, Switzerland. Compiled by UNEP-WCMC, Cambridge. Available from: https://www.cites.org/eng/app/appendices.php (accessed 10 October 2019)

    Colloca, F. & Lelli, S. (2012) Report of the FAO EastMed support to the fishing trials carried out off the South Lebanese Coast. EastMed technical documents14. FAO, Rome, 38 pp.

    Convention on the Conservation of Migratory Species of Wild Animals (CMS) (2015). Appendices I and II of the Convention on the Conservation of Migratory Species of Wild Animals (CMS). Available from: http://www.cms.int/en/species (accessed 10 October 2019)

    Crocetta, F. & Bariche, M. (2015) Six new records from Lebanon, with general implications for Mediterranean alien fauna. In: Crocetta, F., Agius, A., Balistreri, P., Bariche, M., Bayhan, Y., Çakir, M., Ciriaco, S., Corsini-Foka, M., Deidun, A., Zrelli, R., Ergüden, D., Evans, J., Ghelia, M., Giavasi, M., Kleitou, P., Kondylatos, G., Lipej, L., Mifsud, C., Özvarol, Y., Pagano, A., Portelli, P., Poursanidis, D., Rabaoui, L., Schembri, P., Taşkin, E., Tiralongo, F. & Zenetos, A., New Mediterranean Biodiversity Records (October 2015) Mediterranean Marine Science, 16 (3), 682–702.

    https://doi.org/10.12681/mms.1477

    Crocetta, F. & Bariche, M. (2016) Citizen scientists contribute to better knowledge of the Mediterranean marine biota: records of five alien and a native species from Lebanon. In: Dailianis, T., Akyol, O., Babali, N., Bariche, M., Crocetta, F., Gerovasileiou, V., Ghanem, R., Gökoðlu, M., Hasiotis, T., Izquierdo-Muñoz, A., Julian, D., Katsanevakis, S., Lipej, L., Mancini, E., Mytilineou, C., Ounifi Ben Amor, K., Özgül, A., Ragkousis, M., Rubio-Portillo, E., Servello, G., Sini, M., Stamouli, C., Sterioti, A., Teker, S., Tiralongo, F. & Trkov, D. (Eds.), New Mediterranean Biodiversity Records (July 2016). Mediterranean Marine Science, 17 (2), 608–626.

    https://doi.org/10.12681/mms.1734

    Crocetta, F. & Bariche, M. (2017) Four marine fishes of different origins in Lebanon. In: Gerovasileiou, V., Akel, E.H.K., Akyol, O., Alongi, G., Azevedo, F., Babali, N., Bakiu, R., Bariche, M., Bennoui, A., Castriota, L., Chintiroglou, C.C., Crocetta, F., Deidun, A., Galinou-Mitsoudi, S., Giovos, I., Gökoðlu, M., Golemaj, A., Hadjioannou, L., Hartingerova, J., Insacco, G., Katsanevakis, S., Kleitou, P., Korun, J., Lipej, L., Malegue, M., Michailidis, N., Mouzai Tifoura, A., Ovalis, P., Petrović, S., Piraino, S., Rizkalla, S.I., Rousou, M., Savva, I., Şen, H., Spinelli, A., Vougioukalou, K.G., Xharahi, E., Zava, B. & Zenetos, A. (Eds.), New Mediterranean Biodiversity Records (July 2017). Mediterranean Marine Science, 18 (2), pp. 355–384.

    https://doi.org/10.12681/mms.13771

    Demetropoulos, A. & Neocleous, D. (1969) The fishes and crustaceans of Cyprus. Fishery Bulletin, Cyprus, 1, 1–21.

    Edelist, D., Rilov, G., Golani, D., Carlton, J.T. & Spanier, E. (2013) Restructuring the sea: profound shifts in the world’s most invaded marine ecosystem. Diversity and Distributions, 19 (1), 69–77.

    https://doi.org/10.1111/ddi.12002

    Fanelli, E., Azzurro, E., Battaglia, P., Romeo, T., Bariche, M., Maynou, F. & Andaloro, F. (2014) Preliminary data on the isotopic composition of native and lessepsian pelagic species. Biologia Marina Mediterranea, 21 (1), 115–116.

    Fanelli, E., Azzurro, E., Bariche, M., Cartes, J.E. & Maynou, F. (2015) Depicting the novel Eastern Mediterranean food web: a stable isotopes study following Lessepsian fish invasion. Biological Invasions, 17, 2163–2178.

    https://doi.org/10.1007/s10530-015-0868-5

    Fowler, H.W. (1923) Fishes from Madeira, Syria, Madagascar, and Victoria, Australia. Proceedings of the Academy of Natural Science of Philadelphia, 75, 33–45.

    Fricke, R. (2019) Eschmeyer’s Catalog of Fishes: References. Version 4 September 2019. Available from: http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (accessed 5 Oct. 2019)

    Fricke, R. & Eschmeyer, W.N. (2019) Eschmeyer’s Catalog of Fishes: Guide to fish collections. Version of 4 Sept. 2019. Available from: http://researcharchive.calacademy.org/research/ichthyology/catalog/collections.asp (accessed 5 October 2019)

    Fricke, R., Eschmeyer, W.N. & van der Laan, R. (2019) Eschmeyer’s Catalog of Fishes: Genera, species. Version 4 September 2019. Available from: http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (accessed 5 October 2019)

    Galil, B.S. (2007) Loss or gain? Invasive aliens and biodiversity in the Mediterranean Sea. Marine Pollution Bulletin, 55, 314–322.

    https://doi.org/10.1016/j.marpolbul.2006.11.008

    Galil, B.S. (2012) Truth and consequences: the bioinvasion of the Mediterranean Sea. Integrative Zoology, 7 (3), 299–311.

    https://doi.org/10.1111/j.1749-4877.2012.00307.x

    Gaudillière, J., (1958) Rapport au Gouvernement du Liban sur la pêche. FAO Report No 780. FAO, Rome, 151 pp.

    George, C.J. & Athanassiou, V. (1965) On the occurrence of Scomberomorus commerson (Lacepède) in St George Bay, Lebanon. Doriana, 4 (157), 1–4.

    George, C.J. & Athanassiou, V. (1966a) Additions to the check list of the fishes of the coastal waters of Lebanon. Miscellaneous Papers on Natural Sciences, American University of Beirut, 5, 6–8.

    George, C.J. & Athanassiou, V. (1966b) Observations of Upeneus asymetricus Lachner 1954, (Pisces) in St George Bay, Lebanon. Annali di Museo Civico di Storia Naturale “Giacomo Doria”, Genova, 76, 68–74.

    George, C.J. & Athanassiou, V. (1967) A two year study of the fishes appearing in the seine fishery of St George Bay, Lebanon. Annali di Museo Civico di Storia Naturale “Giacomo Doria”, Genova, 76, 237–294.

    George, C.J., Athanassiou, V.A. & Boulos, I. (1964) The fishes of the coastal waters of Lebanon. Miscellaneous Papers in the Natural Sciences. The American University of Beirut, 4, 1–24.

    George, C.J., Athanassiou, V. & Tortonese, E. (1971) The presence of a third species of the genus Sphyraena (Pisces) in the marine waters of Lebanon. Annali di Museo Civico di Storia Naturale “Giacomo Doria”, Genova, 78, 256–263.

    Golani, D. & Appelbaum-Golani, B. (2010) Fish invasions of the Mediterranean Sea: Change and Renewal. Sofia, Pensoft, 332 pp.

    Golani, D. (1993) Trophic adaption of Red Sea fishes to the eastern Mediterranean environment - Review and new data. Israel Journal of Zoology, 39, 391–402.

    Golani, D. (2005) Checklist of the Mediterranean fishes of Israel. Zootaxa, 947 (1), 1–90.

    https://doi.org/10.11646/zootaxa.947.1.1

    Golani, D. (2010) Colonization of the Mediterranean by Red Sea fishes via the Suez Canal—Lessepsian migration. In: Golani, D. & Appelbaum-Golani, B. (Eds.), Fish invasions of the Mediterranean Sea: Change and renewal. Pensoft Publishers, Sofia, pp. 145–188.

    Golani, D., Orsi-Relini, L., Massuti, E. & Quignard, J.P. (2002) CIESM atlas of exotic species in the Mediterranean. Vol. 1. Fishes. Available from: http://www.ciesm.org/atlas/appendix1.html (accessed 20 September 2019)

    Gruvel, A. (1928) Répartition géographique de quelques crustacés comestibles sur les côtes d’Egypte et de Syrie. Comptes rendus de la société de biogéographie, 5 (39), 45–46.

    Gruvel, A. (1931) Les Etats de Syrie. Richesses marines et fluviales. Exploitation actuelle–Avenir. Société d’Editions Géographiques, Maritimes et Coloniale, 453 pp.

    Harmelin-Vivien, M.L., Bitar, G., Harmelin, J.G. & Monestiez, P. (2005) The littoral fish community of the Lebanese rocky coast (eastern Mediterranean Sea) with emphasis on Red Sea immigrants. Biological Invasions, 7, 625–637.

    https://doi.org/10.1007/s10530-004-5852-4

    Hornell, J. (1935) Report on the fisheries of Palestine. Government of Palestine. Crown Agent for the Colonies, London, 106 pp.

    IUCN (2019) The IUCN Red List of Threatened Species. Version 2019-2. Available from: http://www.iucnredlist.org (accessed 20 October 2019)

    Katsanevakis, S., Coll, M., Piroddi, C., Steenbeek, J., Ben Rais Lasram, F., Zenetos, A. & Cardoso, A.C. (2014) Invading the Mediterranean Sea: biodiversity patterns shaped by human activities. Frontiers in Marine Science, 1 (32), 1–11.

    https://doi.org/10.3389/fmars.2014.00032

    Lakkis, S. & Zeidane, R. (1989) L’ichtyoplancton des eaux côtières libanaises: structure et différentiation écologique. Lebanese Science Bulletin, 5 (2), 17–42.

    Lelli, S., Colloca, F., Carpentieri, P. & Russell, B.C. (2008) The threadfin bream Nemipterus randalli (Perciformes: Nemipteridae) in the eastern Mediterranean Sea. Journal of Fish Biology, 73 (3), 740–745.

    https://doi.org/10.1111/j.1095-8649.2008.01962.x

    Liebman, E. (1934) Contribution to the knowledge of Palestine sea fishes. Commission Internationale pour l’Exploration scientifique de la mer Méditerranée (CIESM), Rapport et Procès-Verbaux des Réunions, 8, 317–327.

    Lteif, M. (2015) Biology, distribution and diversity of cartilaginous fish species along the Lebanese coast, eastern Mediterranean. Ecology, environment. Université de Perpignan, English, 262 pp. [NNT: 2015PERP0026]

    Lteif, M., Mouawad, C., Khalaf, G., Lenfant, P., Seret, B. & Verdoit-Jarraya, M. (2017) Population biology of the little gulper shark Centrophorus uyato in Lebanese waters. Journal of Fish Biology, 91 (5), 1491–1509.

    https://doi.org/10.1111/jfb.13484

    Luksch, J.W. (1892-1894) Physikalische Untersuchungen im östlichen Mittelmeer. Reise des S.M. Schiffes “Pola”. Denkschriften der Akademie der Wissenschaften zu Wien, 59, 61.

    Moazzo, G. (1931) Contribution à la faune malacologique marine des côtes libano-syriennes. In: Gruvel, A. (Ed.), Les Etats de Syrie; Richesses marines et fluviatiles. Exploitation actuelle—Avenir. Société d’Editions Géographiques, Maritimes et Coloniales, pp. 437–453.

    Mouneimné, N. (1977) Liste des poissons de la côte du Liban (Mediterranée orientale). Cybium, 3 (1), 37–66.

    Mouneimné, N. (1979) Poissons nouveaux pour les côtes libanaises. Cybium, 3 (6), 105–110.

    Mouneimné, N. (2002) Poissons marins du Liban et de la Mediterranée orientale. INCAM-EU/CNRS Lebanon, Beyrouth, 271 pp.

    Nader, M., Indary, S. & Boustany, L. (2012) FAO EastMed The Puffer Fish Lagocephalus sceleratus (Gmelin 1789) in the Eastern Mediterranean. EastMed Technical Documents, GCP/INT/041/EC-GRE-ITA/TD-10, 33 pp.

    Nielsen, J.N. (1912) Hydrography of the Mediterranean and adjacent waters. Danish Oceanographic Expedition, 1, 77–192.

    Pajuelo, J.G., González, J.A., Triay-Portella, R., Martín, J.A., Ruiz-Díaz, R., Lorenzo, J.M. & Luque, Á. (2016) Introduction of non-native marine fish species to the Canary Islands waters through oil platforms as vectors. Journal of Marine Systems, 163, 23–30.

    https://doi.org/10.1016/j.jmarsys.2016.06.008

    Pallary, P. (1938) Les mollusques marins de la Syrie. Journal de Conchyliologie, 82 (1), 5–58.

    Philibert, P. (1964) Propositions en vue de l’adoption d’un programme de modernisation et de developpement des pêches maritimes au Liban. Ministère du Plan, Beyrouth, 118 pp.

    Por, F.D. (1978) Lessepsian migration: the influx of Red Sea biota into the Mediterranean by way of the Suez Canal. Vol. 23. Springer-Verlag, Berlin, 228 pp.

    Russell, F.S. (1936) Fishery Research in the Eastern Mediterranean. Nature, 137 (3478), 1080–1080.

    https://doi.org/10.1038/1371080a0

    Sala, E., Kizilkaya, Z., Yildirim, D. & Ballesteros, E. (2011) Alien marine fishes deplete algal biomass in the eastern Mediterranean. PLoS ONE, 6 (2), e17356.

    https://doi.org/10.1371/journal.pone.0017356

    Samaha, C., zu Dohna, H. & Bariche, M. (2016) Analysis of Red Sea fish species’ introductions into the Mediterranean reveals shifts in introduction patterns. Journal of Biogeography, 43 (9), 1797–1807. https://doi.org/10.1111/jbi.12793

    Spicer, I.J. (1931) Fisheries. In: Report of the department of agriculture and forests for the years 1927–30. Printing Office, Russian Building, Jerusalem, pp. 159–160.

    Steinitz, H. (1952) Notes on Fishes from Cyprus. Bulletin de l’Institut Océanographique, Monaco, 1004, 1–12.

    Steinitz, W. (1929) Die Wanderung indopazifischer Arten ins Mittelmeer seit Beginn der Quartär-Periode. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 22 (1), 1–90.

    https://doi.org/10.1002/iroh.19290220102

    Streftaris, N. & Zenetos, A. (2006) Alien marine species in the Mediterranean - the 100 ‘‘worst invasives’’ and their impact. Mediterranean Marine Science, 7 (1), 87–118.

    https://doi.org/10.12681/mms.180

    Toda, T. (1961) On an interim report about Lebanese fisheries. Report to the Ministry of Agriculture and Forestry, Lebanon, 12 pp.

    Zenetos, A., Çinar, M.E., Crocetta, F., Golani, D., Rosso, A., Servello, G., Shenkar, N., Turon, X. & Verlaque, M. (2017) Uncertainties and validation of alien species catalogues: The Mediterranean as an example. Estuarine, Coastal and Shelf Science, 191, 171–187.

    https://doi.org/10.1016/j.ecss.2017.03.031

    Zenetos, A., Gofas, S., Morri, C., Rosso, A., Violanti, D., García Raso, J.E., Çinar, M.E., Almogi-Labin, A., Ateš, A.S., Azzurro, E., Ballesteros, E., Bianchi, C.N., Bilecenoglu, M., Gambi, M.C., Giangrande, A., Gravili, C., Hyams-Kaphzan, O., Karachle, P.K., Katsanevakis, S., Lipej, L., Mastrototaro, F., Mineur, F., Pancucci-Papadopoulou, M.A., Ramos Esplá, A., Salas, C., San Martín, G., Sfriso, A., Streftaris, N. & Verlaque, M. (2012) Alien species in the Mediterranean Sea by 2012. A contribution to the application of European Union’s Marine Strategy Framework Directive (MSFD). Part 2 Introduction trends and pathways. Mediterranean Marine Science, 13 (2), 328–352.

    https://doi.org/10.12681/mms.327

    Zenetos, A., Gofas, S., Verlaque, M., Cinar, M.E., Raso, G., Bianchi, C.N., Morri, C., Azzurro, E., Bilecenoglu, M., Froglia, C., Siokou, I., Violanti, D., Sfriso, A., San Martín, G., Giangrande, A., Kataðan, T., Ballesteros, E., Ramos-Esplá, A., Mastrototaro, F., Ocaña, O., Zingone, A., Gambi, M.C. & Streftaris, N. (2010) Alien species in the Mediterranean Sea by 2010. A contribution to the application of European Union’s Marine Strategy Framework Directive (MSFD). Part I. Spatial distribution. Mediterranean Marine Science, 11 (2), 381–493.

    https://doi.org/10.12681/mms.87