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Issue: Vol. 4789 No. 1: 8 Jun. 2020
Type: Article
Published: 2020-06-08
DOI: 10.11646/zootaxa.4789.1.5
Page range: 171–203
Abstract views: 233
PDF downloaded: 5

A new species of Cyrtodactylus (Squamata: Gekkonidae) from southern Vietnam

Cologne Zoo, Riehler Strasse 173, D-50735 Cologne, Germany. Institute of Zoology, University of Cologne, Zülpicher Street 47b, D-50674 Cologne, Germany.
Phu Yen University, 18 Tran Phu Road, Phu Yen, Vietnam.
Faculty of Environmental Sciences, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam. Central Institute for Natural Resources and Environmental Studies, Vietnam National University Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam Department of Herpetology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024
Central Institute for Natural Resources and Environmental Studies, Vietnam National University Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam 7Faculty of Biology, University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, Vietnam.
Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam.
Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam. Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Cau Giay, Hanoi, Vietnam.
Faculty of Biology, University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, Vietnam.
Cologne Zoo, Riehler Strasse 173, D-50735 Cologne, Germany. Institute of Zoology, University of Cologne, Zülpicher Street 47b, D-50674 Cologne, Germany.
Reptilia Cyrtodactylus phumyensis sp. nov. molecular phylogeny taxonomy Phu My Binh Dinh

Abstract

We describe a new species of Cyrtodactylus on the basis of four specimens collected from Phu My, Binh Dinh Province, southern Vietnam. Cyrtodactylus phumyensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: size small (SVL up to 66.8 mm); two internasals; dorsal tubercle rows 18 or 19 at midbody; ventral scale rows 33–41; ventrolateral folds slightly developed; each thigh with 5–7 enlarged femoral scales; femoral pores absent in males and female; a series of 5–7 precloacal pores plus a pitted, enlarged precloacal scale in males; 6 pitted, enlarged precloacal scales in female; paravertebral tubercles 20–23; lamellae under toe IV 18–21; small subcaudal scales, not transversely enlarged; two irregular dark longitudinal stripes on shoulder. In phylogenetic analyses, the new species is recovered as a member of the Cyrtodactylus irregularis species group, and strongly supported as a sister taxon of C. cucdongensis from Khanh Hoa Province.

 

References

  1. Agarwal, I., Datta-Roy, A., Bauer, A.M. & Giri, V.B. (2012) Rediscovery of Geckoella jeyporensis (Squamata: Gekkonidae), with notes on morphology, coloration and habitat. Hamadryad, 36 (1), 17–24.

    Agarwal, I., Khandekar, A. & Bauer, A.M. (2018) A new bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from the Western Himalayas, Himachal Pradesh, India Zootaxa, 4446 (4), 442–454

    https://doi.org/10.11646/zootaxa.4446.4.2

    Agarwal, I., Mahony, S., Giri, V.B., Chaitanya, R. & Bauer, A.M. (2018) Two new species of bent toed geckos, Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Northeast India with comments on name-bearing types from the region. Zootaxa, 4420 (3), 334–356.

    https://doi.org/10.11646/zootaxa.4420.3.2

    Agarwal, I., Mirza, Z.A., Pal, S., Maddock, S.T., Mishra, A. & Bauer, A.M. (2016) A new species of the Cyrtodactylus (Geckoella) collegalensis (Beddome, 1870) complex (Squamata: Gekkonidae) from Western India. Zootaxa, 4170 (2), 339–354.

    https://doi.org/10.11646/zootaxa.4170.2.7

    Batuwita, S. & Bahir, M.M. (2005) Description of five new species of Cyrtodactylus (Reptilia: Gekkonidae) from Sri Lanka. Raffles Bulletin of Zoology, Supplement 12, 351–380.

    Bauer, A.M. (2002). Two new species of Cyrtodactylus (Squamata: Gekkonidae) from Myanmar. Proceedings of the California Academy of Sciences, 53 (7), 75–88.

    Bauer, A.M. (2003) Descriptions of seven new Cyrtodactylus (Squamata: Gekkonidae) with a key to the species of Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (25), 463–498.

    Bauer, A.M. & Henle, K. (1994) Familia Gekkonidae (Reptilia, Sauria). Part 1 Australia and Oceania. Das Tierreich 109 (part). Walter de Gruyter, Berlin, xiii + 306 pp.

    https://doi.org/10.1515/9783110885958

    Bauer, A.M., Kunya, K., Sumontha, M., Niyomwan, P., Panitvong, N., Pauwels, O.S.G., Chanhome, L. & Kunya, T. (2009) Cyrtodactylus erythrops (Squamata: Gekkonidae), a new cave-dwelling gecko from Mae Hong Son Province, Thailand. Zootaxa, 2124 (1), 51–62.

    https://doi.org/10.11646/zootaxa.2124.1.4

    Bauer, A., Kunya, K., Sumontha, M., Niyomwan, P., Pauwels, O.S.G., Chanhome, L. & Kunya, T. (2010) Cyrtodactylus dumnuii (Squamata: Gekkonidae), a new cave–dwelling gecko from Chiang Mai Province, Thailand. Zootaxa, 2570 (1), 41–50.

    https://doi.org/10.11646/zootaxa.2570.1.2

    Bauer, A.M., Pauwels, O.S.G. & Chanhome, L. (2002) A new species of cave-dwelling Cyrtodactylus (Squamata: Gekkonidae) from Thailand. The Natural History Journal of Chulalongkorn University, 2, 19–29.

    Bauer, A.M., Sumontha, M. & Pauwels, O.S.G. (2003) Two new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Thailand. Zootaxa, 376 (1), 1–18.

    https://doi.org/10.11646/zootaxa.376.1.1

    Boulenger, G.A. (1905) On some batrachians and reptiles from Tibet. Annals and Magazine of Natural History, Series 7, 15 (88), 378–379.

    https://doi.org/10.1080/03745480509443063

    Chan-ard, T. & Makchai, S. (2011) A new insular species of Cyrtodactylus Gray, 1827 (Squamata, Gekkonidae), from the Surin Islands, Phang-nga Province, southern Thailand. The Thailand Natural History Museum Journal, 5 (1), 7–15.

    Chan, K.O. & Norhayati, A. (2010) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa, 2389 (1), 47–56.

    https://doi.org/10.11646/zootaxa.2389.1.2

    Connette, G.M., Oswald, P., Thura, M.K., Connette, K.J.L., Grindley, M.E., Songer, M., Zug, R. G. & Mulcahy, G.D. (2017) Rapid forest clearing in a Myanmar proposed national park threatens two newly discovered species of geckos (Gekkonidae: Cyrtodactylus). PLoS ONE, 12 (4), e0174432.

    https://doi.org/10.1371/journal.pone.0174432

    Darevsky, I.S., Helfenberger, N., Orlov, N. & Shah, K. (1998) Two new species of the genus Gonydactylus (Sauria: Gekkonidae) from eastern Nepal. Rusian. Journal of Herpetology, 4 (2), 89–93. [1997]

    Das, I. (1994) Cnemaspis gordongekkoi, a new gecko from Lombok, Indonesia, and the biogeography of oriental species of Cnemaspis (Squamata: Sauria: Gekkonidae). Hamadryad, 18, 1–9. [1993]

    Das, I. (1997) A new species of Cyrtodactylus from the Nicobar Islands, India. Journal of Herpetology, 31, 375–382.

    https://doi.org/10.2307/1565666

    David, P., Nguyen, T.Q., Schneider, N. & Ziegler, T. (2011) A new species of the genus Cyrtodactylus Gray, 1827 from central Laos (Squamata: Gekkonidae). Zootaxa, 2833 (1), 29–40.

    https://doi.org/10.11646/zootaxa.2833.1.3

    Davis, H.R., Bauer, A.M., Jackman, T.R., Nashriq, I. & Das, I. (2019) Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia. Zootaxa, 4614 (2), 331–352.

    https://doi.org/10.11646/zootaxa.4614.2.4

    Dring, J.C.M. (1979) Amphibians and reptiles from northern Trengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum (Natural History) Zoology, 34 (5), 181–24.

    Duda, P.L. & Sahi, D.N. (1978) Cyrtodactylus himalayanus: a new gekkonid species from Jammu, India. Journal of Herpetology, 12 (3), 351–354.

    https://doi.org/10.2307/1563616

    Grismer, L.L. (2015) New species of bent-toed gecko (Cyrtodactylus Gray 1827) from Pulau Aur, Johor, West Malaysia. Journal of Herpetology, 39 (3), 424–432.

    https://doi.org/10.1670/3-05A.1

    Grismer, L.L. & Ahmad, N. (2008) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Penisular Malaysia. Zootaxa, 1924, 53–68.

    https://doi.org/10.11646/zootaxa.1924.1.3

    Grismer, L. & Grismer, J. (2017) A re-evaluation of the phylogenetic relationships of the Cyrtodactylus condorensis group (Squamata; Gekkonidae) and a suggested protocol for the characterization of rock-dwelling ecomorphology in Cyrtodactylus. Zootaxa, 4300 (4), 486–504.

    https://doi.org/10.11646/zootaxa.4300.4.2

    Grismer, L.L. & Leong, T.M. (2005) New species of Cyrtodactylus (Squamata: Gekkonidae) from Southern Penisular Malaysia. Journal of Herpetology, 39 (4), 584–591.

    https://doi.org/10.1670/43-05A.1

    Grismer, L.L., Wood, P. L.Jr., Anuar, S., Davis, H.R., Burch, B.T., Cobos, A.J. & Murdoch, M. L. (2016) A new species of karst forest Bent-toed Gecko (genus Cyrtodactylus Gray) not yet threatened by foreign cement companies and a summary of Penisular Malaysia´s endemic karst forest herpetofauna and the need for its conservation. Zootaxa, 4061 (1), 1–17.

    https://doi.org/10.11646/zootaxa.4061.1.1

    Grismer, L.L., Wood, P.L. Jr., Ngo, V.T. & Murdoch, M.L. (2015) The systematics and independent evolution of cave ecomorphology in distantly related clades of Bent-toed Geckos (Genus Cyrtodactylus Gray, 1827) from the Mekong Delta and islands in the Gulf of Thailand. Zootaxa, 3980 (1), 106–126.

    https://doi.org/10.11646/zootaxa.3980.1.6

    Grismer, L.L., Wood, P.L. Jr., Quah, E.S.H., Anuar, S., Muin, A.M., Sumontha, M., Ahmad, N., Bauer, A.M., Wangkulangkul, S., Grismer, J.L. & Pauwels, O.S.G. (2012) A phylogeny and taxonomy of the Thai-Malay Peninsula bent-toed geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa, 3520 (1), 1–55.

    https://doi.org/10.11646/zootaxa.3520.1.1

    Grismer, L.L., Wood, P.L.J.R., Quah, E.S.H., Murdoch, M.L., Grismer, M.S., Herr, M.W., Espinoza, R.E., Brown, R.M. & Lin, A. (2018) A phylogenetic taxonomy of the Cyrtodactylus peguensis group (Reptilia: Squamata: Gekkonidae) with descriptions of two new species from Myanmar. PeerJ, 6, e5575.

    https://doi.org/10.7717/peerj.5575

    Grismer, L.L., Wood, P.L. Jr., Thura, M.K., Zin, T., Quah, E.S.H., Murdoch, M.L., Grismer, M.S., Lin, A., Kyaw, H. & Lwin, N. (2017) Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism. Zoological Journal of the Linnean Society, 182 (4), 862–959.

    https://doi.org/10.1093/zoolinnean/zlx057

    Günther, R. & Rösler, H. (2002) Eine neue Art der Gattung Cyrtodactylus Gray, 1827 aus dem Westen von Neuguinea (Reptilia: Sauria: Gekkonidae). Salamandra, 38 (4), 195–212.

    Hartmann, L., Mecke, S., Kieckbusch, Mader, F. & Kaiser, H. (2016) A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa, 4067 (5), 552–568.

    https://doi.org/10.11646/zootaxa.4067.5.2

    Harvey, M.B., O’Connel, K.A., Barraza, G., Riyanto, A., Kurniawan, N. & Smith, E.N. (2015) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the Southern Bukit Barisan Range of Sumatra and an estimation of their phylogeny. Zootaxa, 4020 (3), 495–516.

    https://doi.org/10.11646/zootaxa.4020.3.5

    Heidrich, A., Rösler, H., Vu, T.N., Böhme, W. & Ziegler, T. (2007) Another new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha - Ke Bang National Park, central Truong Son, Vietnam. Zootaxa, 1445 (1), 35–48.

    https://doi.org/10.11646/zootaxa.1445.1.3

    Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology, 13 (3), 91–107.

    https://doi.org/10.5358/hsj1972.13.3_91

    Iskandar, D., Rachmansah, A. & Umilaela (2011) A new bent-toed gecko of the genus Cyrtodactylus Gray, 1827 (Reptilia, Gekkonidae) from Mount Tompotika, eastern peninsula of Sulawesi, Indonesia. Zootaxa, 2838 (1), 65–78.

    https://doi.org/10.11646/zootaxa.2838.1.4

    Ivanova, N.V., Dewaard, J.R. & Hebert, P.D.N. (2006) An inexpensive, automation-friendly protocol for recovering high-quality DNA. Molecular Ecology, 6, 998–1002.

    https://doi.org/10.1111/j.1471-8286.2006.01428.x

    Johnson, C.B., Qoah, E.S.H., Anuar, S., Muin, M.A., Wood, P.L. Jr., Grismer, J.L., Greer, L.F., Chan, K.O., Norhayati, A., Bauer, A.M. & Grismer, L.L. (2012) Phylogeography, geographic variation, and taxonomy of the bent-toed gecko Cyrtodactylus quadrivirgatus Taylor, 1962 from Peninsular Malaysia with the description of a new swamp dwelling species. Zootaxa, 3406 (1), 39–58.

    https://doi.org/10.11646/zootaxa.3406.1.3

    Kathriner, A., Bauer, A.M., O’Shea, M., Sanchez, C. & Kaiser, H. (2014) Hiding in plain sight: A new species of bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from West Timor, collected by Malcolm Smith in 1924. Zootaxa, 3900 (4), 555–568.

    https://doi.org/10.11646/zootaxa.3900.4.6

    Khan, M.S. (1993) A new angular-toed gecko from Pakistan, with remarks on the taxonomy and a key to the species belonging to genus Cyrtodactylus (Reptilia: Sauria: Gekkonidae). Pakistan Journal of Zoology, 25 (1), 67–73.

    King, W. (1962) A new gekkonid lizard of the genus Cyrtodactylus from the Philippine Islands. Fieldiana Zoology, 44, 117–120.

    https://doi.org/10.5962/bhl.title.3779

    Kraus, F. (2008) Taxonomic partitioning of Cyrtodactylus louisiadensis (Lacertilia: Gekkonidae) from Papua New Guinea. Zootaxa, 1883 (1), 1–27.

    https://doi.org/10.11646/zootaxa.1883.1.1

    Kunya, K., Panmongkol, A., Pauwels, O.G.S., Sumontha, M., Meewasana, J., Bunkhwamdi, W. & Dangsri, S. (2014) A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Suthep, Chiang Mai Province, northern Thailand. Zootaxa, 3811 (2), 251–261.

    https://doi.org/10.11646/zootaxa.3811.2.6

    Kunya, K., Sumontha, M., Panitvong, N., Dongkumfu, W., Sirisamphan, T. & Pauwels, O.S.G. (2015) A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Inthanon, Chiang Mai Province, northern Thailand. Zootaxa, 3905 (4), 573–584.

    https://doi.org/10.11646/zootaxa.3905.4.9

    Le, D.T., Nguyen, T.Q., Le, M.D. & Ziegler, T. (2016) A new species of Cyrtodactylus (Squamata: Gekkonidae) from Ninh Binh Province, Vietnam. Zootaxa, 4162 (2), 268–282.

    https://doi.org/10.11646/zootaxa.4162.2.4

    Luu, V.Q., Bonkowski, M., Nguyen, T.Q., Le, M.D., Schneider, N., Ngo, H.T. & Ziegler, T. (2016a) Evolution in karst massifs: Cryptic diversity among bent–toed geckos along the Truong Son Range with descriptions of three new species and one new country record from Laos. Zootaxa, 4107 (2), 101–140.

    https://doi.org/10.11646/zootaxa.4107.2.1

    Luu, V.Q., Dung, T.V., Nguyen, T.Q., Le, M.D. & Ziegler, T. (2017) A new species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from Gia Lai Province, Central Highlands of Vietnam. Zootaxa, 4362 (3), 385-404.

    https://doi.org/10.11646/zootaxa.4362.3.4

    Luu, V.Q., Nguyen, T.Q., Do, H.Q. & Ziegler, T. (2011) A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam. Zootaxa, 3129 (1), 39–50.

    https://doi.org/10.11646/zootaxa.3129.1.3

    Luu, V.Q., Calame, T., Bonkowski, M., Nguyen, T.Q. & Ziegler, T. (2014) A new species of Cyrtodactylus (Squamata: Gekkonidae) from Khammouane Province, Laos. Zootaxa, 3760 (1), 054−066.

    http://dx.doi.org/10.11646/zootaxa.3760.1.3

    Luu, V.Q., Calame, T., Nguyen, T.Q., Bonkowski, M. & Ziegler, T. (2015) A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa, 4058 (3), 388−402.

    https://doi.org/10.11646/zootaxa.4058.3.6

    Mahony, S. (2009) Taxonomic status of Cyrtodactylus khasiensis tamaiensis (Smith, 1940) and description of a new species allied to C. chrysopylos Bauer, 2003 from Myanmar (Reptilia: Gekkonidae). Hamadryad, 34 (1), 62–74.

    Mecke, S., Hartmann, L., Mader, F., Kieckbusch, M. & Kaiser, H. (2016) Redescription of Cyrtodactylus fumosus (Müller, 1895) (Reptilia: Squamata: Gekkonidae), with a revised identification key to the bent-toed geckos of Sulawesi. Acta Herpetologica, 11 (2), 151−160.

    Mecke, S., Kieckbusch, Hartmann, L. & Kaiser, H. (2016) Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa, 4175 (4), 353–365.

    https://doi.org/10.11646/zootaxa.4175.4.5

    Murdoch, M.L., Grismer, L.L., Wood, P.L. Jr., Neang, T., Poyarkov, N.A., Ngo, V.T., Nazarov, R.A., Aowphol, A., Pauwels, O.S.G., Nguyen, H.N. & Grismer, J.L. (2019) Six new species of the Cyrtodactylus intermedius complex (Squamata: Gekkonidae) from the Cardamom Mountains and associated highlands of Southeast Asia. Zootaxa, 4554 (1), 001–062.

    https://doi.org/10.11646/zootaxa.4554.1.1

    Nazarov, R.A., Pauwels, O.S.G., Konstantinov, E.L., Chulisov, A.S., Orlov, N.L. & Poyarkov, N.A. (2018) A new Karst-dwelling bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from Xiengkhoang Province, northeastern Laos. Zoological Research, 39 (3), 197−213.

    Nazarov, R.A., Poyarkov, N.A., Orlov, N.L., Nguyen, S.N., Milto, K.D., Martynov, A.A., Konstantinov, E.L. & Chulisov, A.S. (2014) A review of genus Cyrtodactylus (Reptilia: Sauria: Gekkonidae) in fauna of Laos with description of four new species. Proceedings of the Zoological Institute, Russian Academy of Sciences, 318, 391–423.

    Ngo, T.V. (2008) Two new cave-dwelling species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Southwestern Vietnam. Zootaxa, 1909 (1), 37–51.

    https://doi.org/10.11646/zootaxa.1909.1.4

    Ngo, T.V. (2008) Cyrtodactylus martini, another new karst-dwelling Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Northwestern Vietnam. Zootaxa, 2834 (1), 33−46.

    https://doi.org/10.11646/zootaxa.2834.1.3

    Ngo, T.V. & Bauer, A.M. (2008) Descriptions of two new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) endemic to southern Vietnam. Zootaxa, 1715 (1), 27–42.

    https://doi.org/10.11646/zootaxa.1715.1.2

    Ngo, T.V. & Chan, K.O. (2010) A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa province, southern Vietnam. Zootaxa, 2504 (1), 47–60.

    https://doi.org/10.11646/zootaxa.2504.1.4

    Ngo, T.V. & Chan, K.O. (2011) A new karstic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from northern Vietnam. Zootaxa, 3125, 51–63.

    Ngo, T.V. & Grismer, L.L. (2010) A new karst dwelling Cyrtodactylus (Squamata: Gekkonidae) from Son La Province, northwestern Vietnam. Hamadryad, 35, 84–95.

    Ngo, T.V. & Grismer, L.L. (2012) A new endemic species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Tho Chu Island, southwestern Vietnam. Zootaxa, 3228 (1), 48–60.

    https://doi.org/10.11646/zootaxa.3228.1.2

    Ngo, T.V., Grismer, L.L. & Grismer, J.L. (2008) A new endemic cave dwelling species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Kien Giang Biosphere Reserve, southwestern Vietnam. Zootaxa, 1967 (1), 53–62.

    https://doi.org/10.11646/zootaxa.1967.1.3

    Ngo, T.V., Grismer, L.L. & Grismer, J.L. (2010) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Phu Quoc National Park, Kien Giang Biosphere Reserve, southwestern Vietnam. Zootaxa, 2604 (1), 37–51.

    https://doi.org/10.11646/zootaxa.2604.1.3

    Ngo, H.T., Le, M.D., Pham, N.D., Nguyen, V.T.H., Pham, C.T. & Nguyen, T.Q. (2017) Taxonomy and Phylogenetic relationships of Cyrtodactylus (Squamata: Gekkonidae) in Vietnam. VNU Journal of Science, 33(1S), 182-192.

    Ngo, T. V. & Pauwels, O. S. G. (2010) A new cave-dwelling species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Khammouane Province, southern Laos. Zootaxa, 2730 (1), 44–56.

    https://doi.org/10.11646/zootaxa.2730.1.3

    Nguyen, S.N., Nguyen, L.T., Nguyen, V.D.H. & Murphy, R.W. (2017) Genetic diversity, phylogeny, and altitudinal distribution of geckos genus Cyrtodactylus on Hon Ba Moutain, Khanh Hoa Province, with note on the subcaudals of C. yangbayensis. Journal of Biotechnology, 15 (3A), 55–62.

    Nguyen, L.T., Schmidt, H.A., von Haeseler, A. & Bui, M.Q. (2015) IQ-TREE: A fast and effective stochastic algorithm for estimating maximum likelihood phylogenies. Molecular Biology and Evolution, 32, 268–274.

    https://doi.org/10.1093/molbev/msu300

    Nguyen, S.N., Orlov, N.L. & Darevsky, S.I. (2006) Descriptions of two new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from Southern Vietnam. Russian Journal of Herpetology, 13, 215–226.

    Nguyen, S.N., Yang, J.-X., Le, N.T.T., Nguyen, L.T., Orlov, N.L., Hoang, C.V., Nguyen, T.Q., Jin, J.-Q., Rao, D.-Q., Hoang, T.N., Che, J., Murphy, R.W. & Zhang, Y.-P. (2014) DNA barcoding of Vietnamese bent-toed geckos (Squamata: Gekkonidae: Cyrtodactylus) and the description of a new species. Zootaxa, 3784 (1), 48–66.

    https://doi.org/10.11646/zootaxa.3784.1.2

    Nguyen, T.Q., Ananjeva, B.N., Orlov, N.L., Rybaltovsky, E. & Böhme, W. (2010) A New Species of the Genus Scincella Mittlemann, 1950 (Squamata: Scincidae) from Vietnam. Russian Journal of Herpetology, 17 (4), 269–274.

    Nguyen, T.Q, Pham, A.V., Ziegler, T., Ngo, H.T. & Le, M.D. (2017) A new species of Cyrtodactylus (Squamata: Gekkonidae) and the first record of C. otai from Son La Province, Vietnam. Zootaxa, 4341 (1), 25–40.

    https://doi.org/10.11646/zootaxa.4341.1.2

    Orlov, N.L., Nguyen, Q.T., Nazarov, R.A., Anajeva, N.B. & Nguyen, S.N. (2007) A new species of the genus Cyrtodactylus Gray, 1827 and redescription of Cyrtodactylus paradoxus (Darevsky et Szczerbak, 1997) [Squamata: Sauria: Gekkonidae] from south Vietnam. Russian Journal of Herpetology, 14, 145–152.

    Panitvong, N., Sumontha, M., Tunprasert, J. & Pauwels, O.S.G. (2014) Cyrtodactylus saiyok sp. nov., a new dry evergreen forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Kanchanaburi Province, western Thailand. Zootaxa, 3869 (1), 64–74.

    https://doi.org/10.11646/zootaxa.3869.1.6

    Pauwels, O.S.G., Bauer, A.M., Sumontha, M. & Chanhome, L. (2004) Cyrtodactylus thirakhupti (Squamata: Gekkonidae), a new cave-dwelling gecko from southern Thailand. Zootaxa, 772 (1), 1–11.

    https://doi.org/10.11646/zootaxa.772.1.1

    Pauwels, O.S.G., Nazarov, R.A., Bobrov, V.V. & Poyarkov, N.A. (2018) Taxonomic status of two populations of Bent-toed Geckos of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) with description of a new species from Nui Chua National Park, southern Vietnam. Zootaxa, 4403 (2), 307–335.

    https://doi.org/10.11646/zootaxa.4403.2.5

    Pauwels, O.S.G. & Sumontha, M. (2014) Cyrtodactylus samroiyot, a new limestone-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Prachuap Khiri Khan Province, peninsular Thailand. Zootaxa, 3755 (6), 573–583.

    https://doi.org/10.11646/zootaxa.3755.6.4

    Pauwels, O.S.G., Sumontha, M. & Bauer, A.M. (2016) A new Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Phetchaburi Province, Thailand. Zootaxa, 4088 (3), 409–419.

    https://doi.org/10.11646/zootaxa.4088.3.6

    Pauwels, O.S.G., Sumontha, M., Keeratikiat, K. & Phanamphon, E. (2014a) Cyrtodactylus kunyai (Squamata: Gekkonidae), a new cave-dwelling Bent-toed Gecko from Loei Province, northeastern Thailand. Zootaxa, 3821 (2), 253–264.

    https://doi.org/10.11646/zootaxa.3821.2.5

    Pauwels, O.S.G., Sumontha, M., Latinne, A. & Grismer, L.L. (2013) Cyrtodactylus sanook (Squamata: Gekkonidae), a new cave-dwelling gecko from Chumphon Province, southern Thailand. Zootaxa, 3635 (3), 275–285.

    https://doi.org/10.11646/zootaxa.3635.3.7

    Pauwels, O.S.G., Sumontha, M., Panitvong, N. & Varaguttanonda, V. (2014b) Cyrtodactylus khelangensis, a new cave-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Lampang Province, northern Thailand. Zootaxa, 3755 (6), 584–594.

    https://doi.org/10.11646/zootaxa.3755.6.5

    Pham, A.V, Le, M.D., Ngo, H.T., Ziegler, T. & Nguyen, T.Q. (2019) A new species of Cyrtodactylus (Squamata: Gekkonidae) from northwestern Vietnam. Zootaxa, 4544 (1), 360−380.

    https://doi.org/10.11646/zootaxa.4544.3.3

    Pham, A.V., Pham, N.V., Nguyen, L.D.T., Le, M.D., Do, Q.H., Pham, N.D. & Nguyen, T.Q. (2017) Additional morphological characters and molecular divergence of Cyrtodactylus bichnganae (Reptilia: Gekkonidae). Proceedings of the Seventh National Scientific Conference on Ecology and Biological Resources, 2017, pp. 554−561. [in Vietnamese with English abstract]

    Posada, D. & Crandall, K.A. (1998) MODELTEST: testing the model of DNA substitution. Bioinformatics, 14, 817−818.

    https://doi.org/10.1093/bioinformatics/14.9.817

    Quah, E.S.H, Grismer, L.L., Wood, P.L. Jr. & Sah, S.A.M. (2019) The discovery and description of a new species of Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Penisular Malaysia. Zootaxa, 4668 (1), 51−75.

    https://doi.org/10.11646/zootaxa.4668.1.3

    Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology, 61, 539−542.

    https://doi.org/10.1093/sysbio/sys029

    Schneider, N., Nguyen, T.Q., Schmitz, A., Kingsada, P., Auer, M. & Ziegler, T. (2011) A new species of karst dwelling Cyrtodactylus (Squamata: Gekkonidae) from northwestern Laos. Zootaxa, 2930 (1), 1–21.

    https://doi.org/10.11646/zootaxa.2930.1.1

    Simmons, J.E. (2002) Herpetological Collecting and Collections Management, Revised edition. Society for the Study of Amphibians and Reptiles. Herpetological Circular, 31, 1-153.

    Smith, M.A. (1917) Descriptions of new reptiles and a new batrachian from Siam. Journal of Natural History Society of Siam, 2, 221–225.

    Smith, M.A. (1921a) New or little-known reptiles and batrachians from southern Annam (Indo-China). Proceedings of the Zoological Society of London, 1921, 423–440.

    https://doi.org/10.1111/j.1096-3642.1921.tb03271.x

    Smith, M.A. (1921b) Reptiles and Batrachians collected on Pulo Condore. Journal Natural History Society, 4, 93−97. [1920]

    Smith, M.A. (1935) The Fauna of British India, Including Ceylon and Burma. Vol. II. Sauria. Taylor and Francis, London, viii + 440 pp.

    Sterling, E.J., Hurley, M.H. & Le, M.D. (2006) Vietnam: A Natural History. Yale University Press, New Haven, Connecticut, 460 pp.

    Sumontha, M., Panitvong, N. & Deein, G. (2010) Cyrtodactylus auribalteatus (Squamata: Gekkonidae), a new cave-dwelling gecko from Phitsanulok Province, Thailand. Zootaxa, 2370 (1), 53–64.

    https://doi.org/10.11646/zootaxa.2370.1.3

    Sumontha, M., Pauwels, O.S.G., Kunya, K., Nitikul, A., Samphanthamit, P. & Grismer, L.L. (2012) A new forest-dwelling gecko from Phuket Island, Southern Thailand, related to Cyrtodactylus macrotuberculatus (Squamata: Gekkonidae). Zootaxa, 3522 (1), 61–72.

    https://doi.org/10.11646/zootaxa.3522.1.4

    Sumontha, M., Pauwels, O.S.G., Suwannakarn, N., Nutatheera, N. & Sodob, W. (2014) Cyrtodactylus wangkulangkulae (Squamata: Gekkonidae), a new Bent-toed Gecko from Satun Province, southern Thailand. Zootaxa, 3821 (1), 116–124.

    https://doi.org/10.11646/zootaxa.3821.1.8

    Swofford, D.L. (2001) PAUP*. Phylogenetic Analysis Using Parsimony (* and Other Methods), version 4. Sinauer Associates, Sunderland, Massachusetts. [program]

    Taylor, E.H. (1963) Lizards of Thailand. The Universtiy of Kansas Science Bulletin, 55, 687–1077.

    Thompson, J.D., Gibson, T.J., Plewniak, F., Jeanmougin, F. & Higgins, D.G. (1997) The ClustalX windows interface: Flexible strategies for multiple sequence alignment aided by quality analysis tools. Nucleic Acids Research, 25, 4876−4882.

    https://doi.org/10.1093/nar/25.24.4876

    Tordoff, A.W., Tran, B.Q., Nguyen, T.D. & Le, H.M. (Eds.) (2004) Sourcebook of existing and proposed protected areas in Vietnam, second edition. Birdlife International in Indochina and Ministry of Agriculture and Rural Development, Hanoi.

    Uetz, P. (2020) The Reptile Database. Available from: http://www.reptile-database.org (accessed 1 Feb. 2020)

    Ulber, T. (1993) Bemerkungen über cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitteilungen aus dem Zoologischen Museum in Berlin, 69, 187–200.

    https://doi.org/10.1002/mmnz.19930690202

    Ziegler, T., Nazarov, R., Orlov, N., Nguyen, T.Q., Vu, T.N., Dang, K.N., Dinh, T.H. & Schmitz, A. (2010) A third new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha—Ke Bang National Park, Vietnam. Zootaxa, 2413 (1), 20–36.

    https://doi.org/10.11646/zootaxa.2413.1.2

    Ziegler, T., Phung, T.M., Le, M.D. & Nguyen, T.Q. (2013) A new Cyrtodactylus (Squamata: Gekkonidae) from Phu Yen Province, southern Vietnam. Zootaxa, 3686 (4), 432–446.

    https://doi.org/10.11646/zootaxa.3686.4.2

    Ziegler, T., Rösler, H., Herrmann, H.-W. & Vu, T.N. (2002) Cyrtodactylus phongnhakebangensis sp. n., ein neuer Bogenfingergecko aus dem annamitischen Karstwaldmassiv, Vietnam. Herpetofauna, 24, 11–25.