Abstract
Non-studied museum collections are hidden treasures—a source of information for various research fields. The novel taxa presented here were discovered during taxonomic examination of the backlogs of Bryozoa (Cheilostomata) from the Iziko South African Museum. We describe one new genus, Khulisa n. gen., and nine new species of bryozoans from South Africa. The new species are: Biflustra adenticulata n. sp., Aspidostoma sarcophagus n. sp., ?Micropora erecta n. sp., Trypostega richardi n. sp., Khulisa carolinae n. gen. et n. sp., Adeonella assegai n. sp., Hippomonavella lingulata n. sp., Phidolopora chakra n. sp. and Reteporella ilala n. sp. Three genera, Biflustra, Phidolopora and Triphyllozoon, are recorded for the first time from South Africa. This study highlights the importance of examining existing backlogged material lodged in museum collections.
References
Allmann, G.J. (1856) A monograph of the Freshwater Polyzoa, including all the known species, both British and Foreign. The Ray Society, London, 119 pp. https://doi.org/10.5962/bhl.title.9143
Álvarez, J.A. (1990) Membranipora arborescens (Bryozoa: Cheilostomida): a new record in the Bay of Biscay. Journal of the Marine Biological Association of the United Kingdom, 70, 917–919.
https://doi.org/10.1017/S0025315400059154
Álvarez, J.A. (1992a) Briozoos de la Campaña Fauna I. Parte Primera: Ctenostomida y Cheilostomida Anascina. Cahiers de Biologie Marine, 33, 273–297.
Álvarez, J.A. (1992b) Contribución al estudio faunístico del tipo Bryozoa en las costas meridionales de la Península Ibérica. La colección de Briozoos del Proyecto Fauna I. Unpublished Thesis, Universidad del País Vasco, 130 pp.
Amui, A.M. (2005) Adeonellas from the Gulf of Aden including one new species. Zootaxa, 1012 (1), 45–52.
https://doi.org/10.11646/zootaxa.1012.1.5
Arakawa, S. (2016) Taxonomy of some microporids (Bryozoa: Cheilostomata) from the Pacific Coast of Japan. Species Diversity, 21, 9–30.
https://doi.org/10.12782/sd.21.1.009
Audouin, J.V. (1826) Explication sommaire des planches de polypes de l’Egypte et de la Syrie, publiees par Jules-Cesar Savigny. In: Audouin, J.V. (Ed.), Description de l’Egypte, ou recueil des observations et des recherches qui ont été faites en Egypte pendant l’expédition de l’armée française. Histoire naturelle. Vol. Tome 1. 4 Partie. Imprimerie Impériale, Paris, pp. 1–339.
Bassler, R.S. (1934) Notes on Fossil and Recent Bryozoa. Journal of the Washington Academy of Science, 24, 404–408.
Berning, B. (2006) The cheilostome bryozoan fauna from the Late Miocene of Niebla (Guadalquivir Basin, SW Spain): environmental and biogeographic implications. Mitteilungen am dem Geologisch-Palciontologischen Institut der Universität Hamburg, 90, 7–156.
Blainville, de H.M.D. (1830) Zoophytes. In: Cuvier, G.F. (Ed.), Dictionnaire des sciences naturelles, dans lequel on trate méthodiquement des differents êtres de la nature...par plusieur professeurs du Muséum Nationale d’Histoire Naturelle et des autres principales écoles de Paris. F.G. Levrault, Paris, pp. 535–546.
Bock, P.E. (2020) The Bryozoa Home Page. Available from: http://bryozoa.net/ (accessed 30 June 2020)
Bock, P.E. & Cook, P.L. (2001) Revision of Tertiary species of Anaskopora Wass (Bryozoa: Cribrimorpha). Memoirs of Museum Victoria, 58, 179–189.
https://doi.org/10.24199/j.mmv.2001.58.9
Bock, P.E. & Gordon, D.P. (2013) Phylum Bryozoa Ehrenberg, 1831. In: Zhang, Z.-Q. (Ed.), Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013). Zootaxa, 3703 (1), pp. 67–74.
https://doi.org/10.11646/zootaxa.3703.1.14
Boonzaaier, M.K., Florence, W.K. & Spencer Jones, M.E. (2014) Historical review of South African bryozoology: a legacy of European endeavour. In: Wyse Jackson, P.N. & Spencer Jones, M.E. (Eds.), Annals of Bryozoology. 4. Aspects of the history of research on bryozoans. International Bryozoology Association, Dublin, pp. 1–34.
Brown, D.A. (1952) The Tertiary cheilostomatous polyzoa of New Zealand. British Museum, London, 405 pp.
https://doi.org/10.5962/bhl.title.118779
Busk, G. (1852) Catalogue of marine Polyzoa in the collection of the British Museum. I. Cheilostomata. Trustees of the British Museum, London, 54 pp.
https://doi.org/10.5962/bhl.title.20859
Busk, G. (1854) Cheilostomata (part). In: Catalogue of marine Polyzoa in the collection of the British Museum. II. Trustees of the British Museum, London, pp. 55–120.
https://doi.org/10.5962/bhl.title.20859
Busk, G. (1859) A monograph of the fossil Polyzoa of the Crag. The Palaeontographical Society, London, 136 pp.
https://doi.org/10.5962/bhl.title.2037
Busk, G. (1875) Catalogue of the marine Polyzoa in the collection of the British Museum. Part III. Cyclostomata. Trustees of the British Museum, London, 39 pp.
Busk, G. (1884) Report on the Polyzoa collected by H.M.S. Challenger during the years 1873–1876. Part I. The Cheilostomata. Report on the Scientific Results of the Voyage of the H.M.S. “Challenger”, Zoology, 10, 1–216.
Busk, G. (1886) Report on the Polyzoa collected by H.M.S. Challenger during the years 1873-1876. Part II. The Cyclostomata, Ctenostomata and Pedicellinea. Report on the Scientific Results of the Voyage of the H.M.S. “Challenger”, Zoology, 17, 1–47.
Canu, F. (1900) Révision des Bryozoaires du Crétacé figurés par d’Orbigny. II, Cheilostomata. Bulletin de la Société géologique de France, 28, 334–463.
Canu, F. (1911) Bryozoaires in types du prodrome de paleontologie de Alcide d’Orbigny. Annales de Paléontologie, 6, 67 (pl. 9).
Canu, F. & Bassler, R.S. (1917) A synopsis of American Early Tertiary Cheilostome Bryozoa. United States National Museum Bulletin, 96, 1–87.
https://doi.org/10.5479/si.03629236.96.1
Canu, F. & Bassler, R.S. (1920) North American early Tertiary Bryozoa. Bulletin of the U.S. National Museum, 106, 1–879.
https://doi.org/10.5479/si.03629236.106.i
Canu, F. & Bassler, R.S. (1922) Studies on the cyclostomous Bryozoa. Proceedings of the U.S. National Museum, 61, 1–160.
https://doi.org/10.5479/si.00963801.61-2443.1
Canu, F. & Bassler, R.S. (1928) Bryozoaires du Brésil. Bulletin de la Société des Sciences de Seine-et-Oise, 9, 58–110.
Canu, F. & Bassler, R.S. (1929) Bryozoa of the Phillipine Region. Bulletin of the U.S. National Museum, 9, 1–685.
Cook, P.L. (1966) Some “sand fauna” Polyzoa (Bryozoa) from Eastern Africa and the northern Indian Ocean. Cahiers de Biologie Marine, 7, 207–223.
Cook, P.L. (1982) Notes on some African Adeonellidae (Bryozoa, Cheilostomata). Journal of Natural History, 16, 833–846.
https://doi.org/10.1080/00222938200770671
Couch, R.Q. (1844) A Cornish fauna: being a compendium of the natural history of the country, intended to form a companion to the collection in the Royal Institution of Cornwall. Part 3. The zoophytes and calcareous corallines. Royal Institution of Cornwall, Truro, 136 pp.
https://doi.org/10.5962/bhl.title.87783
Di Martino, E. & Rosso, A. (2015) Revision of the bryozoan genus Gephyrotes Norman, 1903 (Cheilostomata, Cribrilinidae) with the description of two new taxa. Zootaxa, 3941 (2), 261–283.
https://doi.org/10.11646/zootaxa.3941.2.7
Esper, E.J.C. (1791–1797) Forsetzungen der Pflanzenthiere in Abbildungen nach der Natur mit Farben erleuchtet hebst Beschreibungen. Theil 1. in der Raspischen Buchhandlung, Nürnberg, 230 pp.
https://doi.org/10.5962/bhl.title.118730
Florence, W.K. (2016) Some deep-water Cheilostome Bryozoa from the south coast of South Africa. African Natural History, 12, 5–11.
https://doi.org/10.17159/2305-7963/2016/v12n1a2
Florence, W.K., Hayward, P.J. & Gibbons, M.J. (2007) Taxonomy of shallow-water Bryozoa from the west coast of South Africa. African Natural History, 3, 1–58.
Gabb, W.M. & Horn, G.H. (1862) The fossil Polyzoa of the Secondary and Tertiary Formations of North America. Journal of the Academy of Natural Sciences of Philadelphia, 5, 111–179.
Gordon, D.P. (1984) The marine fauna of New Zealand: Bryozoa: Gymnolaemata from the Kermadec Ridge. New Zealand Oceanographic Institute Memoir, 91, 1–198.
Gordon, D.P. (1989) The marine fauna of New Zealand: Bryozoa: Gymnolaemata (Cheilostomida Ascophorina) from the western South Island continental shelf and slope. New Zealand Oceanographic Institute Memoir, 97, 1–158.
Gordon, D.P. & Taylor, P.D. (1999) Bryozoa of the Early Eocene Tumaio Limestone, Chatham Island. New Zealand. Bulletin of the Natural History Museum, Geology, 55, 1–45.
Grange, K.R. & Gordon, D.P. (2005) The spread and potential impact of the introduced bryozoan Biflustra grandicella. Final Research Report for Ministry of Fisheries Research Project ZBS2003-07. National Institute of Water and Atmospheric Research, Wellington, 30 pp.
Gray, J.E. (1848) List of the Specimens of British Animals in the Collection of the British Museum. 1. Centroniae or Radiated Animals. British Museum, London, 173 pp.
Hara, U. (2001) Bryozoans from the Eocene of Seymour Island, Antarctic Peninsula. In: Gaździcki, A. (Ed.), Palaeontological Results of the Polish Antarctic Expeditions. Palaeontologia Polonica, Part III, pp. 33–156.
Harmer, S.F. (1933) The genera of Reteporidae. Proceedings of the Zoological Society of London, 1933, 615–627.
https://doi.org/10.1111/j.1096-3642.1933.tb01610.x
Hayward, P.J. (1981) The Cheilostomata (Bryozoa) of the deep sea. Galathea Report. Scientific Results of the Danish Deep-Sea Expedition around the world (1950–52), 15, 21–68.
Hayward, P.J. (1988) The recent species of Adeonella (Bryozoa: Cheilostomata) including descriptions of fifteen new species. Zoological Journal of the Linnean Society, 94, 111–191.
https://doi.org/10.1111/j.1096-3642.1988.tb00105.x
Hayward, P.J. (1995) Antartic cheilostomatous Bryozoa. Oxford University Press, Oxford, 355 pp.
Hayward, P.J. (1999) Taxonomic studies on some Australian Phidoloporidae (Bryozoa: Cheilostomatida). Journal of Zoology of London, 249, 219–240.
https://doi.org/10.1111/j.1469-7998.1999.tb00760.x
Hayward, P.J. & Cook, P.L. (1979) The South African Museum’s Meiring Naudé Cruises. Part 9, Bryozoa. Annals of the South African Museum, 79, 43–130.
Hayward, P.J. & Cook, P.L. (1983) The South African Museum’s Meiring Naudé Cruises. Part 13, Bryozoa II. Annals of the South African Museum, 91, 1–161.
Hayward, P.J. & Ryland, J.S. (1979) British Ascophoran Bryozoans. In: Kermack, D.M. & Barnes, R.S.K. (Eds.), Synopses of the British Fauna. New Series 14. Academic Press for the Linnaean Society, London, pp. 1–312.
Hayward, P.J. & Ryland, J.S. (1991) New and little known Bryozoa from Antarctica and the southwest Atlantic. Bulletin du Muséum national d’histoire naturelle, 4, 241–261.
Helgen, K.M., Pinto, C.M., Kays, R., Helgen, L.E., Tsuchiya, M.T.N. & Quinn, A. (2013) Taxonomic revision of the olingos (Bassaricyon), with description of a new species, the Olinguito. Zookeys, 324, 1–83.
https://doi.org/10.3897/zookeys.324.5827
Hincks, T. (1879) On the classification of the British Polyzoa. Annals and Magazine of Natural History, 3, 153–164.
https://doi.org/10.1080/00222937908682494
Hincks, T. (1880) Contributions towards a general history of the marine Polyzoa. Part I. Madeiran Polyzoa. Annals and Magazine of Natural History, Series 5, 6 (31), 69–80.
https://doi.org/10.1080/00222938009458895
Hincks, T. (1881) Contributions towards a general history of the marine Polyzoa. Part IV. Annals and Magazine of Natural History, 7, 147–161.
https://doi.org/10.1080/00222938109459489
Hutton, F.W. (1873) Polyzoa. In: Catalogue of the marine Mollusca of New Zealand, with diagnoses of species. Government Printer, Wellington, pp. 87–104.
Johnston, G. (1838) A history of British Zoophytes. W.H.Lizars, Edinburgh, London and Dublin, 341 pp.
https://doi.org/10.5962/bhl.title.110844
Jullien, J. (1888) Bryozoaires. Mission Scientifique du Cap Horn 1882–1883, 6, 1–92.
Kluge, G.A. (1914) Die Bryozoen der deutschen Südpolar-Expedition 1901–1903 I. Die Familien Aetidae, Cellularidae, Bicellaridae, Farciminaridae, Flustridae, Membraniporidae und Cribilinidae. Deutsche Südpolar-Expedition ,15 (Zoologie), 7, 599–678.
Lagaaij, R. (1952) The Pliocene Bryozoa of the Low Countries and their bearing on the marine stratigraphy of the North Sea region. Mededelingen van de Geologische Stichting, 5, 6–233.
Lawrence, R.F. (1948) New species of Solifguae in the collection of the Transvaal Museum. Annals of the Transvaal Museum, 21, 201–208.
Lee, H.-J., Kwan, Y.-S., Kong, S.-R., Min, B.S., Seo, J.E. & Won, Y.-J. (2011) DNA barcode examination of Bryozoa (Class: Gymnolaemata) in Korean Seawater. Korean Journal of Systematic Zoology, 27, 159–163.
https://doi.org/10.5635/KJSZ.2011.27.2.159
Levinsen, G.M.R. (1902) Studies on Bryozoa. Videnskabelige Meddelelser fra den naturhistoriske Foreningi København, 54, 1–31.
Levinsen, G.M.R. (1909) Morphological and Systematic Studies on the Cheilostomatous Bryozoa. Nationale Forfatteres Forlag, Copenhagen, 431 pp.
https://doi.org/10.5962/bhl.title.5690
López-Gappa, J., Liuzzi, M.G. & Pereyra, C. (2020) A new species of Hippomonavella (Bryozoa: Cheilostomata) from the Holocene and recent of Argentina and Uruguay (Southwest Atlantic). Zootaxa, 4728 (1), 143–148.
https://doi.org/10.11646/zootaxa.4728.1.8
López de la Cuadra, C.M. & García-Gómez, J.C. (2000) The cheilostomate Bryozoa (Bryozoa: Cheilostomatida) collected by the Spanish “Antártida 8611” Expedition to the Scotia Arc and South Shetland Islands. Journal of Natural History, 34, 755–772.
https://doi.org/10.1080/002229300299408
Louis, S. & Menon, N.R. (2009) Biflustra perambulata n. sp. (Cheilostomata: Bryozoa), a new alien species from Cochin Harbour, Kerala, India. Zootaxa, 2066 (1), 59–68.
https://doi.org/10.11646/zootaxa.2066.1.3
Liu, X. (1992) On the genus Membranipora (Anasca: Cheilostomata: Bryozoa) from south Chinese seas. Raffles Bulletin of Zoology, 40, 103–144.
MacGillivray, P.H. (1860) On some new Australian Polyzoa. Transactions of the Philosophical Society of Victoria, 4, 97–98.
MacGillivray, P.H. (1881) Polyzoa. In: McCoy, F. (Ed.), Prodromus of the Zoology of Victoria Decade VI. Government Printer, Melbourne, pp. 27–46.
MacGillivray, P.H. (1883) Descriptions of new or little-known Polyzoa. Parts 2, 3 and 4. Transactions and Proceedings of the Royal Society of Victoria, 19, 130–138 + 191–195 + 287–293.
MacGillivray, P.H. (1887) Descriptions of new or little-known Polyzoa, Parts 10, 11 and 12. Transactions and Proceedings of the Royal Society of Victoria, 19, 34–38 + 64–72 + 179–186.
MacGillivray, P.H. (1895) A monograph of the Tertiary Polyzoa of Victoria. Transactions of the Royal Society of Victoria, 4, 1–166.
https://doi.org/10.5962/bhl.title.6076
Marcus, E. (1922) Südafrikanische Bryozoen aus der Sammlung des Gothenburger Museums. Gotenburgs Kongligen Vetenskaps och Vitterhets Samhalles Handlingar, 25, 1–45.
Mead, A., Carlton, J.T., Griffiths, C.L. & Rius, M. (2011) Revealing the scale of marine bioinvasions in developing regions: a South African re-assessment. Biological Invasions, 13, 1991–2008.
https://doi.org/10.1007/s10530-011-0016-9
Moratelli, R. & Wilson, D.E. (2014) A new species of Myotis (Chiroptera, Vespertilionidae ) from Bolivia. Journal of Mammalogy, 95, 17–25.
https://doi.org/10.1644/14-MAMM-149
Norman, A.M. (1864) On undescribed British Hydrozoa, Actinozoa and Polyzoa. Annals and Magazine of Natural History, 13, 82–90.
https://doi.org/10.1080/00222936408681578
Norman, A.M. (1903) Notes on the natural history of East Finmark, Polyzoa. Annals and Magazine of Natural History, Series 7, 11, 567–598.
https://doi.org/10.1080/00222930308678818
O’Donoghue, C.H. (1924) The Bryozoa (Polyzoa) collected by the S.S. “Pickle.” Report No. 3 of the Fisheries and Marine Biological Survey. For the year 1922, Special Reports, 10, 1–63.
O’Donoghue, C.H. (1957) Some South African Bryozoa. Transactions of the Royal Society of South Africa, 35, 71–95.
https://doi.org/10.1080/00359195709519012
O’Donoghue, C.H. & de Watteville, D. (1935) A collection of Bryozoa from South Africa. Journal of the Linnean Society, Zoology, 39, 203–218.
https://doi.org/10.1111/j.1096-3642.1935.tb00070.x
O’Donoghue, C.H. & de Watteville, D. (1937) Notes on South African Bryozoa. Zoologischer Anzeiger, 117, 12–22.
O’Donoghue, C.H. & de Watteville, D. (1944) Additional notes on Bryozoa from South Africa. Annals of the Natal Museum, 10, 407–432.
Oliver, J.-C. & Florence, W.K. (2016) A new species of Taylorius (Bryozoa: Escharinidae) from the east coast of South Africa. African Natural History, 12, 1–4.
https://doi.org/10.17159/2305-7963/2016/v12n1a1
Orbigny, d’, A. (1851–54) Bryozoaires. Paléontologie Française, Description des Mollusques et Rayonnés Fossiles, Terrains Crétacés. Victor Masson, Paris, 1192 pp. [pp. 1–188 (1851) + pp. 185 bis–472 (1852) + pp. 473–984 (1853) + pp. 985–1192 (1899)]
Orr, R.J.S., Haugen, M.N., Berning, B., Bock, P.E., Cumming, R.L., Florence, W.K., Hirose, M., Di Martino, E., Ramsfjell, M.H., Sannum, M.M., Smith, A.M., Vieira, L.M., Waeschenbach, A. & Liow, L.H. (2019) A genome-skimmed phylogeny of a widespread bryozoan family, Adeonidae. BMC Evolutionary Biology, 19, 1–10. https://doi.org/10.1186/s12862-019-1563-4
Ortmann, A. (1890) Die Japanische Bryozoenfauna. Bericht über die von Herrn Dr.L.Döderlein in Jahre 1880-81, gemachten Sammlungen. Archiv für Naturgeschichte, 56, 1–74.
Ostrovsky, A.N. (2013) Evolution of sexual reproduction in marine invertebrates: example of Gymnolaemate Bryozoans. Springer-Verlag, Dordrecht, 356 pp.
https://doi.org/10.1007/978-94-007-7146-8
Ostrovsky, A.N., O’Dea, A., & Rodríguez, F. (2009) Comparative anatomy of internal incubational sacs in cupuladriid bryozoans and the evolution of brooding in free‐living cheilostomes. Journal of Morphology, 270, 1413–1430.
https://doi.org/10.1002/jmor.10767
Ramalho, L.V., Muricy, G. & Taylor, P.D. (2008) Two new species of Bitectiporidae (Bryozoa, Ascophora) from Rio de Janeiro State, Brazil. In: Hageman, S.J., Key, M.M.J. & Winston, J.E. (Eds.), Proceedings of the 14th International Bryozoology Association Conference, Boone, North Carolina, July 1–8, 2007, Virginia Museum of Natural History. Special Publication No. 15. Virginia Museum of Natural History, Martinsville, Virginia, pp. 243–250.
Rosso, A. & Novosel, M. (2010) The genus Adeonella (Bryozoa, Ascophora) in the Mediterranean, with description of two new living species and rediscovery of a fossil one. Journal of Natural History, 44, 1697–1727.
https://doi.org/10.1080/00222931003760061
Sheridan, J.A. & Stuart, B.L. (2018) Hidden species diversity in Sylvirana nigrovittata (Amphibia: Ranidae) highlights the importance of taxonomic revisions in biodiversity conservation. PLoS One, 13, e0196242.
https://doi.org/10.1371/journal.pone.0192766
Shimada, K., Welton, B.J. & Long, D.J. (2014) A New Fossil Megamouth Shark (Lamniformes, Megachasmidae) from the Oligocene—Miocene of the Western United States. Journal of Vertebrate Paleontology, 34, 281–290.
https://doi.org/10.1080/02724634.2013.803975
Silén, L. (1941) Cheilostomata Anasca (Bryozoa) collected by Prof. Dr. Sixten Bock’s expedition to Japan and the Bonin Islands 1914. Arkiv för zoologi, 33A, 1–130.
Smitt, F.A. (1868) Bryozoa marina in regionibus arcticis et borealibus viventia recensuit. Öfversigt af Kongliga Vetenskaps- Akademiens Förhandlingar, 24, 443–487.
Souto, J., Reverter-Gil, O., De Blauwe, H. & Fernández Pulpeiro, E. (2014) New records of bryozoans from Portugal. Cahiers de Biologie Marine, 55, 129–150.
Taylor, P.D. & Tan, S.A. (2015) Cheilostome Bryozoa from Penang and Langkawi, Malaysia. European Journal of Taxonomy, 149, 1–34.
https://doi.org/10.5852/ejt.2015.149
Thornely, L.R. (1924) Polyzoa. Scientific reports of the Australian Antarctic Expedition 1911-1914, Series C, Zoology and Botany, 6, 1–23.
Tilbrook, K.J. (2006) Cheilostomatous Bryozoa from the Solomon Islands. Santa Barbara Museum of Natural History Monographs, 4 (Studies in Biodiversity Number 3), 1–386.
Tilbrook, K.J. (2012) Bryozoa , Cheilostomata: First records of two invasive species in Australia and the northerly range extension for a third. Checklist, 8, 181–183.
https://doi.org/10.15560/8.1.181
Tilbrook, K.J., Hayward, P.J. & Gordon, D.P. (2001) Cheilostomatous Bryozoa from Vanuatu. Zoological Journal of the Linnean Society, 131, 35–109.
https://doi.org/10.1006/zjls.2000.0249
Uttley, G.H. (1949) The Recent and Tertiary Polyzoa (Bryozoa) in the collection of the Canterbury Museum. Part 1. Records of the Canterbury Museum, 5, 167–192.
Velazco, P.M. & Patterson, B.D. (2014) Two new species of yellow-shouldered bats, genus Sturnira Gray, 1842 (Chiroptera, Phyllostomidae ) from Costa Rica, Panama and western Ecuador. Zookeys, 402, 43–66.
https://doi.org/10.3897/zookeys.402.7228
Waters, A.W. (1904) Bryozoa. Resultats du voyage du S.Y. Belgica. Zoologie, 4, 1–114.
Waters, A.W. (1907) Tubucellaria; its species and ovicells. Journal of the Linnean Society, Zoology, 30, 126–133.
https://doi.org/10.1111/j.1096-3642.1907.tb02127.x
Waters, A.W. (1909) Reports on the marine biology of the Sudanese Red Sea, from collections made by Cyril Crossland, M.A., B.Sc., F.Z.S.; together with collections made in the Red Sea by Dr R. Hartmeyer, 7. The Bryozoa, Part 1, Cheilostomata. Journal of the Linnean Society, Zoology, 31, 123–181.
https://doi.org/10.1111/j.1096-3642.1909.tb00458.x
Waters, A.W. (1916) Some species of Crisia. Annals and Magazine of Natural History, 8, 469–477.
https://doi.org/10.1080/00222931609486901
Waters, A.W. (1918) Some Mediterranean Bryozoa. Annals and Magazine of Natural History, 2, 96–102.
https://doi.org/10.1080/00222931808562345
White, W.T., Kyne, P.M. & Harris, M. (2019) Lost before found: A new species of whaler shark Carcharhinus obsolerus from the Western Central Pacific known only from historic records. PLoS One, 14, e0209387.
https://doi.org/10.1371/journal.pone.0209387
Winston, J.E. (2005) Re-description and revision of Smitt’s “Floridan Bryozoa” in the collection of the Museum of Comparative Zoology, Harvard University (with an annotated catalogue of specimens collected by L.F. de Pourtalès in Florida and Carolina, housed at the Museum of Comparative Zoology, Harvard University). Virginia Museum of Natural History Memoir, 7, 1–147.
WoRMS Editorial Board (2020) World Register of Marine Species. VLIZ. Available from https://www.marinespecies.org (accessed 30 May 2020)
Yang, H.J, Seo, J.E., Min, B.S., Grischenko, A.V. & Gordon, D.P. (2018) Cribrilinidae (Bryozoa: Cheilostomata) of Korea. Zootaxa, 4377 (2), 216–234.