Abstract
Middle Ordovician (Darriwilian) species representing early Laurentian occurrences of the Subfamily Calymeninae Milne Edwards, 1840 (=Flexicalymeninae Siveter, 1977) are assigned to Atlanticalymene n. gen. (type species: A. bardensis n. sp. from the Table Cove Formation, western Newfoundland, Canada). They have routinely been confused with the older (Dapingian) calymenoidean taxon Protocalymene Ross, 1967. Revision of the type species of Protocalymene, P. mcallisteri Ross, 1967, from the Antelope Valley Formation, Funeral Mountains, California, indicates that it is not a calymenine, and that while it is clearly a calymenoidean its close affinity is otherwise difficult to determine. A single genuine calymenine species is known from the Laurentian Dapingian, and revised here as “Calymeninae n. gen.? n. sp. A” from the Antelope Valley Formation, Nevada, USA. A species from the Dapingian of Tarim, known from a single partial cranidium, appears to represent an older, extra-Laurentian species of Atlanticalymene.
References
Adrain, J.M. (2013) A synopsis of Ordovician trilobite distribution and diversity. In: Harper, D.A.T. & Servais, T. (Eds.), Early Palaeozoic Biogeography and Palaeogeography. Geological Society, London, Memoirs, 38, pp. 297–336.
https://doi.org/10.1144/M38.20
Adrain, J.M. & Edgecombe, G.D. (1997) Silurian (Wenlock) calymenid trilobites from the Cape Phillips Formation, central Canadian Arctic. Journal of Paleontology, 71, 657–682.
https://doi.org/10.1017/S0022336000040130
Adrain, J.M. & Fortey, R.A. (1997) Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the Natural History Museum, London, Geology Series, 53, 79–115.
Adrain, J.M. & Westrop, S.R. (2006) Notchpeakia, a new genus of Upper Cambrian (Sunwaptan) “entomaspidid” trilobites. Journal of Paleontology, 80, 1152–1171.
https://doi.org/10.1666/0022-3360(2006)80[1152:NANGOU]2.0.CO;2
Andrawis, S.F., El Afify, F. & El Hameed, A.T.A. (1983) Lower Paleozoic trilobites from subsurface rocks of the Western Desert, Egypt. Neues Jahrbuch fuer Geologie und Palaeontologie, Monatshefte, 1983, 65–68.
Angelin, N.P. (1854) s.n. In: Palaeontologica Scandinavica. II. Crustacea formationis transitionis. Samson and Wallin, Lund, pp. 21–9.
Baldis, B.A.J. & Pöthe de Baldis, E.D. (1995) Trilobites Ordovicicos de la Formacion Las Aguaditas (San Juan, Argentina) y consideraciones estratigraficas. Boletin de la Academia Nacional de Ciencias. Córdoba, 60, 409–448.
Basse, M. (2012) Fossilium Catalogus. I: Animalia. Pars 150: Trilobites Africae: Catalogus typorum. Backhuys, Leiden, 311 pp.
Bridge, J. (1931) Geology of the Eminence and Cardareva quadrangles. Missouri Bureau of Geology and Mines, Series 2, 24, 1–228. [for 1930]
Bruton, D.L., Wright, A.D. & Hamedi, M.A. (2004) Ordovician trilobites from Iran. Palaeontographica, Abteilung A, 271, 111–149.
Černyševa, N.E. (1960) [Trilobita]. In: Orlov, Û.A. (Ed.), Osnovy paleontologii; spravočnik dlâ paleontologov i geologov SSSR. 8. Členistonogie, trilobitoobraznye i rakoobraznye. Akademiâ Nauk SSSR, Moscow, pp. 17–194. [in Russian]
Chapman, E.J. (1890) Some remarks on the classification of the trilobites as influenced by stratigraphical relations: with outline of a new grouping of these forms. Transactions of the Royal Society of Canada, 7 (Section 4), 113–120. [for 1889]
https://doi.org/10.5962/bhl.title.38719
Chatterton, B.D.E. (1971) Taxonomy and ontogeny of Siluro-Devonian trilobites from near Yass, New South Wales. Palaeontographica, Abteilung A, 137, 1–108.
Chatterton, B.D.E. & Campbell, K.S.W. (1980) Silurian trilobites from near Canberra and some related forms from the Yass Basin. Palaeontographica, Abteilung A, 167, 77–119.
Chatterton, B.D.E., Siveter, D.J., Edgecombe, G.D. & Hunt, A.S. (1990) Larvae and relationships of the Calymenina (Trilobita). Journal of Paleontology, 64, 255–277.
https://doi.org/10.1017/S0022336000018424
Clarkson, E.N.K. & Howells, Y. (1981) Upper Llandovery trilobites from the Pentland Hills, Scotland. Palaeontology, 24, 507–536.
Courtessole, R., Marek, L., Pillet, J., Ubaghs, G. & Vizcaïno, D. (1983) Calymenina, Echinodermata et Hyolitha de l’Ordovicien inférieur de la Montagne Noire (France méridionale). Mémoires de la Société des Études Scientifiques de l’Aude, 1983, 1–62.
Dalman, J.W. (1827) Om Palaeaderna eller de så kallade Trilobiterna. Kungliga Svenska Vetenskapsakademiens Handlingar, 1826, 113–152 + 226–294. [for 1826]
Dean, W.T. (1965) A revision of the Ordovician trilobite genus Bathycheilus Holub. Sborník Národního Muzea v Praze. Rada B. Prírodní Vedy, 21, 1–10.
Edgecombe, G.D. (2007) Acaste (Trilobita: Phacopina) from the Early Devonian of Tasmania. Alcheringa, 31, 59–66.
https://doi.org/10.1080/03115510601123627
Edgecombe, G.D. & Wright, A.J. (2004) Silicified Early Devonian trilobites from Brogans Creek, New South Wales. Proceedings of the Linnean Society of New South Wales, 124, 177–188.
Edgecombe, G.D., Chatterton, B.D.E., Waisfeld, B.G. & Vaccari, N.E. (1998) Ordovician (Whiterock) calymenid and encrinurid trilobites from the Precordillera of Argentina. Journal of Paleontology, 72, 678–697.
https://doi.org/10.1017/S0022336000040397
Fortey, R.A. (1975) Early Ordovician trilobite communities. Fossils and Strata, 4, 331–352.
Fortey, R.A. (1983) Cambrian-Ordovician trilobites from the boundary beds in western Newfoundland and their phylogenetic significance. In: Briggs, D.E.G. & Lane, P.D. (Eds.), Trilobites and other early arthropods. Papers in honour of Professor H. B. Whittington, F. R. S. Special Papers in Palaeontology, 30, pp. 179–211.
Fortey, R.A. (1990) Ontogeny, hypostome attachment and trilobite classification. Palaeontology, 33, 529–576.
Fortey, R.A. & Cocks, L.R.M. (2003) Palaeontological evidence bearing on global Ordovician-Silurian continental reconstructions. Earth-Science Reviews, 61, 245–307.
https://doi.org/10.1016/S0012-8252(02)00115-0
Fortey, R.A. & Droser, M.L. (1996) Trilobites at the base of the Middle Ordovician, western United States. Journal of Paleontology, 70, 73–99.
https://doi.org/10.1017/S002233600002312X
Fortey, R.A. & Droser, M.L. (1999) Trilobites from the base of the type Whiterockian (Middle Ordovician) in Nevada. Journal of Paleontology, 73, 182–201.
https://doi.org/10.1017/S0022336000027712
Geyer, G. (1990) Die marokkanischen Ellipsocephalidae (Trilobita: Redlichiida). Beringeria, 3, 3–363.
Hammann, W. (1983) Calymenacea (Trilobita) aus dem Ordovizium von Spanien; ihre Biostratigraphie, Ökologie und Systematik. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 542, 1–177.
Hammann, W. & Leone, F. (1997) Trilobites of the post-Sardic (Upper Ordovician) sequence of southern Sardinia. Part 1. Beringeria, 20, 1–217.
Hicks, H. (1873) On the Tremadoc rocks in the neighbourhood of St. David’s, South Wales. Quarterly Journal of the Geological Society of London, 29, 39–52.
https://doi.org/10.1144/GSL.JGS.1873.029.01-02.11
Holloway, D.J. (1980) Middle Silurian trilobites from Arkansas and Oklahoma, U.S.A. Part 1. Palaeontographica, Abteilung A, 170, 1–85.
Holloway, D.J. (1994) Early Silurian trilobites from the Broken River area, north Queensland. Memoirs of the Museum of Victoria, 54, 243–269.
https://doi.org/10.24199/j.mmv.1994.54.12
Hupé, P. (1953a) Classe des Trilobites. In: Piveteau, J. (Ed.), Traité de Paléontologie. Tome 3. Les Formes Ultimes d’Invertébrés. Morphologie et Évolution. Onycophores. Arthropodes. Échinoderms. Stomocordés. Masson et Cie, Paris, pp. 44–246.
Hupé, P. (1953b) Contribution à l’étude du Cambrien inférieur et du Précambrien III de l’Anti-Atlas marocain. Service géologique du Maroc (Rabat), Notes et Mémoires, 103, 1–402. (for 1952)
Hupé, P. (1955) Classification des Trilobites. Annales de Paléontologie, 41, 111–345.
Jell, P.A. & Adrain, J.M. (2003) Available generic names for trilobites. Memoirs of the Queensland Museum, 48, 331–553.
Klappa, C.F., Opalinski, P.R. & James, N.P. (1980) Middle Ordovician Table Head Group of western Newfoundland; a revised stratigraphy. Canadian Journal of Earth Sciences, 17, 1007–1019.
https://doi.org/10.1139/e80-101
Kobayashi, T. (1956) On the Kaolishaniinae. Japanese Journal of Geology and Geography, 27, 9–20.
Kobayashi, T. & Hamada, T. (1977) Devonian trilobites of Japan in comparison with Asian, Pacific and other faunas. Special Papers of the Palaeontological Society of Japan, 20, 1–202.
Kolobova, I.M. (1978) [Trilobites]. In: Sokolov, B.S. & Yolkin, E.A. (Eds.), Progranichnye sloi ordovika i silura Altae-Sayanskoj blasti i Tyan-Shanya. Trudy Instituta Geologii i Geofiziki. Sibirskoe Otdelenie, Akademiya Nauk SSSR, pp. 126–144. [in Russian]
Li, S.-J. (1978) [Trilobita]. In: [Paleontological atlas of southwest China. Sichuan Province. Volume 1. Sinian to Devonian]. Geological Publishing House, Beijing, pp. 179–284. [in Chinese]
Lin, T.-R. (2008) Silurian. In: Z.-Y., Z. & Zhen, Y.-Y. (Eds.), Trilobite Record of China. Science Press, Beijing, pp. 275–284.
Lindström, G. (1885) Förteckning på Gotlands Siluriska Crustacéer. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar. Stockholm, 6, 37–100.
Loch, J.D. & Ethington, R.L. (2017) An integrated trilobite and conodont biostratigraphy across the base of the Laurentian Whiterockian Series (lower Middle Ordovician) at its stratotype, Whiterock Canyon Narrows, Nevada. Journal of Paleontology, 91, 294–317.
https://doi.org/10.1017/jpa.2016.127
Ludvigsen, R., Westrop, S.R. & Kindle, C.H. (1989) Sunwaptan (Upper Cambrian) trilobites of the Cow Head Group, western Newfoundland, Canada. Palaeontographica Canadiana, 6, 1–175.
Maletz, J., Egenhoff, S., Böhme, M., Asch, R., Borowski, K., Höntzsch, S., Kirsch, M. & Werner, M. (2011) A tale of both sides of Iapetus—upper Darriwilian (Ordovician) graptolite faunal dynamics on the edges of two continents. Canadian Journal of Earth Sciences, 48, 841–859.
https://doi.org/10.1139/e10-105
Milne Edwards, H. (1840) Histoire naturelle des Crustacés, comprenant l’anatomie, la physiologie et la classification de ces animaux. Vol. 3. Roret, Paris, 638 pp.
Prantl, F. & Přibyl, A. (1949) O novych nebo malo známych trilobitech ceského ordoviku. Rozpravy Ceské Akademie Ved a Umeni, 58, 1–22. [for 1948; published in English as: Some new or imperfectly known Ordovician trilobites from Bohemia. Bulletin International de l’Académie Tchèque des Sciences, 49 (8), 1–23]
Přibyl, A. (1953) Seznam ceskych trilobitovych rodu. Knihovna Ústředního Ústavu Geologického, 25, 1–80.
Rasetti, F. (1944) Upper Cambrian trilobites from the Lévis Conglomerate. Journal of Paleontology, 18, 229–258.
Raymond, P.E. (1913) A revision of the species which have been referred to the genus Bathyurus. Bulletin of the Victoria Memorial Museum, 1, 51–69.
https://doi.org/10.4095/104944
Raymond, P.E. (1920) Some new Ordovician trilobites. Bulletin of the Museum of Comparative Zoology, Harvard, 64, 273–296.
Reed, F.R.C. (1935) The Lower Palaeozoic trilobites of Girvan. Supplement No. 3. Monographs of the Palaeontographical Society, 88 (400), 1–64.
https://doi.org/10.1080/02693445.1935.12035635
Ross, R.J., Jr. (1967) Some Middle Ordovician brachiopods and trilobites from the Basin Ranges, western United States. United States Geological Survey Professional Paper, 523-D, 1–43.
https://doi.org/10.3133/pp523D
Ross, R.J., Jr. (1970) Ordovician brachiopods, trilobites, and stratigraphy in eastern and central Nevada. United States Geological Survey Professional Paper, 639, 1–103.
Ross, R.J., Jr. (1972) Fossils from the Ordovician bioherm at Meiklejohn Peak, Nevada. United States Geological Survey Professional Paper, 685, 1–47.
Schmidt, F. (1894) Revision der ostbaltischen silurischen Trilobiten. Abteilung IV: Calymmeniden, Proetiden, Bronteiden, Harpeiden, Trinucleiden, Remopleuriden und Agnostiden. Mémoires de l’Academie Impériale des Sciences de St.-Petersbourg, VII’e Série, 42 (5), 1–93.
Schuchert, C. & Dunbar, C.O. (1934) Stratigraphy of western Newfoundland. Geological Society of America Memoir, 1, 1–123.
https://doi.org/10.1130/MEM1-p1
Sdzuy, K. (1957) Bemerkungen zur Familie Homalonotidae (mit Beschreibung einer neuen Art von Calymenella). Senckenbergiana lethaea, 38, 275–290.
Shaw, F.C. & Bolton, T.E. (2011) Ordovician trilobites from the Romaine and Mingan formations (Ibexian–Late Whiterockian), Mingan Islands, Quebec. Journal of Paleontology, 85, 406–441.
https://doi.org/10.1666/10-062.1
Sheng, X.-F. (1974) [Ordovician trilobites from western Yunnan and its stratigraphical significance]. In: [Subdivision and correlation of the Ordovician System in China]. Geological Publishing House, Beijing, pp. 96–140. [in Chinese]
Shirley, J. (1936) Some British trilobites of the family Calymenidae. Quarterly Journal of the Geological Society of London, 92, 384–422.
https://doi.org/10.1144/GSL.JGS.1936.092.01-04.19
Siveter, D.J. (1977) The Middle Ordovician of the Oslo Region, Norway, 27. Trilobites of the family Calymenidae. Norsk Geologisk Tidsskrift, 56, 335–396. [for 1976]
Siveter, D.J. & Chatterton, B.D.E. (1996) Silicified calymenid trilobites from the Mackenzie Mountains, northwest Canada. Palaeontographica Abteilung A, 239, 43–60.
Stenzel, S.R., Knight, I. & James, N.P. (1990) Carbonate platform to foreland basin; revised stratigraphy of the Table Head Group (Middle Ordovician), western Newfoundland. Canadian Journal of Earth Sciences, 27, 14–26.
https://doi.org/10.1139/e90-002
Stouge, S. (1984) Conodonts of the Middle Ordovician Table Head Formation, western Newfoundland. Fossils and Strata, 16, 1–145.
Swinnerton, H.H. (1915) Suggestions for a revised classification of trilobites. Geological Magazine, Series 6, 2, 487–496, 538–545.
https://doi.org/10.1017/S0016756800203634
Tripp, R.P. (1962) Trilobites from the confinis Flags (Ordovician) of the Girvan district, Ayrshire. Transactions of the Royal Society of Edinburgh, 65, 1–40.
https://doi.org/10.1017/S0080456800012291
Tripp, R.P. (1976) Ordovician trilobites from the basal superstes Mudstones (Ordovician) at Aldons Quarry, near Girvan, Ayrshire. Transactions of the Royal Society of Edinburgh, 69, 369–423.
https://doi.org/10.1017/S0080456800015386
Turvey, S.T. (2002) Phylogeny of the Reedocalymeninae (Trilobita): implications for Early Ordovician biogeography of Gondwana. In: Crame, J.A. & Owen, A.W. (Eds.), Palaeobiogeography and Biodiversity Change: the Ordovician and Mesozoic-Cenozoic Radiations. Geological Society Special Publications 194. Geological Society, London, 53–68.
https://doi.org/10.1144/GSL.SP.2002.194.01.05
Turvey, S.T. (2005) Reedocalymenine trilobites from the Ordovician of central and eastern Asia, and a review of species assigned to Neseuretus. Palaeontology, 48, 549–575.
https://doi.org/10.1111/j.1475-4983.2005.00469.x
Waisfeld, B.G. & Vaccari, N.E. (2003) Trilobites. In: Benedetto, J.L. (Ed.), Ordovician Fossils from Argentina. Secretaría de Ciencia y Tecnología, Universidad Nacional de Córdoba, Cordoba, pp. 295–409.
Westrop, S.R. (1995) Sunwaptan and Ibexian (Upper Cambrian-Lower Ordovician) trilobites of the Rabbitkettle Formation, Mountain River region, northern Mackenzie Mountains, northwest Canada. Palaeontographica Canadiana, 12, 1–75.
Whittington, H.B. (1963) Middle Ordovician trilobites from Lower Head, western Newfoundland. Bulletin of the Museum of Comparative Zoology, Harvard, 129, 1–118.
Whittington, H.B. (1965) Trilobites of the Ordovician Table Head Formation, western Newfoundland. Bulletin of the Museum of Comparative Zoology, Harvard, 132, 275–442.
Whittington, H.B. (1966) Phylogeny and distribution of Ordovician trilobites. Journal of Paleontology, 40, 696–737.
Whittington, H.B. (1971) Silurian calymenid trilobites from the United States, Norway and Sweden. Palaeontology, 14, 455–477.
Whittington, H.B. & Kindle, C.H. (1963) Middle Ordovician Table Head Formation, western Newfoundland. Geological Society of America Bulletin, 74, 745–758.
https://doi.org/10.1130/0016-7606(1963)74[745:MOTHFW]2.0.CO;2
Zhou, Z.-Y. & Fortey, R.A. (1986) Ordovician trilobites from north and northeastern China. Palaeontographica, Abteilung A, 192, 157–210.
Zhou, Z.-Y. & Zhou, Z.-Q. (2008) Latest Cambrian and Ordovician. In: Zhou, Z.-Y. & Zhen, Y.-Y. (Eds.), Trilobite Record of China. Science Press, Beijing, pp. 208–274.
Zhou, Z.-Y., Yin, G.-Z. & Zhou, Z.-Q. (2014) Ordovician (Darriwilian-early Katian) trilobite faunas of northwestern Tarim, Xinjiang, China. Memoirs of the Association of Australasian Palaeontologists, 46, 1–142.
Zhou, Z.-Y., Zhou, Z.-Q. & Yin, G.-Z. (2016) Ordovician trilobites from the uppermost Zhuozishan Formation (early Darriwilian) at Zhuozishan, Wuhai, Inner Mongolia. Memoirs of the Association of Australasian Palaeontologists, 49, 289–328.