Abstract
New diagnostic characters were used to study the genus Litoblatta Hebard: paraprocts, styles due to their shape and distribution of spines, and sclerites of the genitalia of the male with emphasis on the right complex. Male and female genitalia are described in detail. Four new species of cockroaches are described, Litoblatta elisae, Litoblatta lutea, Litoblatta misionerae, and Litoblatta tucumanae. The number of species recorded for Argentina rises to 7 and a key for male determination is provided. The geographic distribution of the studied species of cockroaches is extended.
References
Beccaloni, G.W. (2007) Blattodea Species File Online. Version 1.2/4.0. Available from: http://Blattodea.SpeciesFile.org (accessed 13 June 2019)
Bohn, H. & Chládek, F. (2011) Revision of the maculata-group of Phyllodromica: species from Central Europe (Insecta: Blattodea: Blattellidae: Ectobiinae). Arthropod Systematics & Phylogeny, 69 (11), 3–54.
Brunner von Wattenwyl, C. (1865) Nouveau Système des Blattaires. Braumüller, C., Vienne, 426 pp.
Crespo, F.A. & A.C. Valverde. (2008) Blattaria. In: Claps, L.E., Debandi, G. & Roig-Juñent, S. (Eds.), Biodiversidad de artrópodos argentinos. Vol. 2. Sociedad Entomológica Argentina, Mendoza, pp. 167–179.
Crespo, F.A., Valverde, A.C. & Iglesias, M.S. (2010) Catalogue of Blattaria (Insecta) from Argentina. Zootaxa, 2726 (1), 1–33.
https://doi.org/10.11646/zootaxa.2726.1.1
Crespo, F.A., Valverde, A.C. & Iglesias, M.S. (2015) A new species of Tribonium Saussure, 1862 from the Province of Misiones, Argentina (Blattaria, Blaberidae, Zetoborinae). Zootaxa, 3936 (4), 593–599.
https://doi.org/10.11646/zootaxa.3936.4.9
Gutiérrez, E. (2005) Nueva cucaracha ciega de Litoblatta de Brasil y rediagnosis del género (Dictyoptera: Battaria: Blattellidae). Solenodon, 5, 64–65.
Klass, K.D. (1997) The external male genitalia and the phylogeny of Blattaria and Mantodea. Bonner. Zoologische Monographien, 42, 1–341.
Klass, K.D. (1998) The ovipositor of Dictyoptera (Insecta): homology and ground-plan of the main elements. Zoologister Anzeiger, 236 (2–3), 69–101.
Hebard, M. (1921) South American Blattidae from the Museum National D’Histoire Naturelle, Paris, France. Proceedings Academy of Natural Science of Philadelphia, 73 (2), 193–304.
Mckittrick, F.A. (1964) Evolutionary studies of cockroaches. Cornell University Agriculture Experimental Station Memoir, 389, 1–197.
Princis, K. (1969) Pars 13. Blattaria: subordo Epilamproidea: fam. Blattellidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Junk, The Hague, pp. 713–1038.
Rocha e Silva Albuquerque, I. & Aguiar, G.M. (1975) Revisão do gênero Litoblatta Hebard, 1921 (Blattariae). Revista Brasileira do Biologia, 35 (2), 233–244.
Roth, L.M. (2003) Systematics and Phylogeny of cockroaches (Dictyoptera: Blattaria). Oriental Insects, 37, 1–186.
https://doi.org/10.1080/00305316.2003.10417344
Walker, F. (1868) Catalogue of the specimens of Blattariae in the collection of the British Museum. Printed for the Trustees of the British Museum, London, 239 pp.
https://doi.org/10.5962/bhl.title.8495
Wieczorek, J., Guo, Q. & Hijmans, R.J. (2004) The point-radius method for georeferencing locality descriptions and calculating associated uncertainty. International Journal of Geographical Information Science, 18 (8), 745–767.
https://doi.org/10.1080/13658810412331280211