Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2021-07-05
Page range: 201-252
Abstract views: 773
PDF downloaded: 32

A Reexamination of the Herpetofauna of Oaxaca, Mexico: Composition Update, Physiographic Distribution, and Conservation Commentary

Department of Biological Sciences, The University of Texas at El Paso, El Paso, Texas 79968-0500, USA
Oaxaca de Juárez, Oaxaca 68023, MEXICO.
Department of Biological Sciences, The University of Texas at El Paso, El Paso, Texas 79968-0500, USA
Department of Biological & Environmental Sciences, Georgia College & State University, Milledgeville, Georgia 31061-0490, USA.
Department of Biological Sciences, The University of Texas at El Paso, El Paso, Texas 79968-0500, USA
Laboratorio de Ecología de Poblaciones, Centro de Investigaciones Biológicas, Instituto de Ciencias Básicas e Ingeniería, Universidad Autónoma del Estado de Hidalgo, Km 4.5 Carretera Pachuca-Tulancingo, 42184 Mineral de La Reforma, Hidalgo, MEXICO
Centro Zamorano de Biodiversidad, Escuela Agrícola Panamericana Zamorano, Departamento de Francisco Morazán, HONDURAS; 1350 Pelican Court, Homestead, Florida 33035-1031, USA.
Amphibia Conservation status endemism herpetofaunal update patterns of distribution taxonomic and nomenclatural changes

Abstract

A substantial number of changes to the composition of the herpetofauna of the Mexican state of Oaxaca, including taxonomic additions and deletions, have occurred in the five years since our original assessment of this region. These changes now establish a herpetofauna of 480 species for the state. A number of taxonomic and nomenclatural changes involving the Oaxacan herpetofauna also are discussed. Updated patterns of physiographic distribution, endemism, and conservation status of the members of the state herpetofauna are examined.

 

References

  1. Aguilar-López, J.L., Canseco-Márquez, L., Pineda, E. & Luría-Manzano, R. (2014) Aporte al conocimiento de la distribución de la culebra de cola corta de Linton, Tantillita lintoni en México. Revista Mexicana de Biodiversidad, 85, 1292–1294.
    https://doi.org/10.7550/rmb.46781
    Aguilar-López, J.L. & Canseco-Márquez, L. (2006) Herpetofauna del municipio de Las Choapas, Veracruz, México. Boletín de la Sociedad Herpetológica Mexicana, 14, 20­–37.
    Bocourt, M.F. (1876) Note sur quelques reptiles de l’isthme de Tehuantepec (Mexique) donnés par M. Sumichrast au Muséum. Journal de Zoologie, 5, 386‒411.
    Bogert, C.M. & Porter, A.P. (1966) A new species of Geophis (Serpentes: Colubridae) from the state of Colima, Mexico. American Museum Novitates, 2260, 1­–10.
    Breuil, M., Vuillaume, B., Schikorski, D., Krauss, U., Morton, M.N., Haynes, P., Daltry, J.C., Corry, E., Gaymes, G., Gaymes, J., Bech, N., Jelic, M. & Grandean, F. (2019) A story of nasal horns: two new subspecies of Iguana Laurenti, 1768 (Squamata, Iguanidae) in Saint Lucia, St Vincent and the Grenadines, and Grenada (southern Lesser Antilles). Zootaxa, 4608 (2), 201–232.
    https://doi.org/10.11646/zootaxa.4608.2.1
    Campbell, J.A. (2015) A new species of Rhadinella (Serpentes: Colubridae) from the Pacific versant of Oaxaca, Mexico. Zootaxa, 3918 (3), 397–405.
    https://doi.org/10.11646/zootaxa.3918.3.3
    Campbell, J.A., Ford, L.S. & Karges, J.P. (1983) Resurrection of Geophis anocularis Dunn with comments on its relationships and natural history. Transactions of the Kansas Academy of Sciences, 86, 38–47.
    https://doi.org/10.2307/3628422
    Campbell, J.A., Smith, E.N. & Hall, A.S. (2018) Caudals and calyces: the curious case of a consumed Chiapan colubroid. Journal of Herpetology, 52, 458–471.
    https://doi.org/10.1670/18-042
    Campbell, J.A., Solano-Zavaleta, I., Flores-Villela, O., Caviedes-Solís, I.W. & Frost, D.R. (2016) A new species of Abronia (Squamata: Anguidae) from the Sierra Madre del Sur of Oaxaca, Mexico. Journal of Herpetology, 50, 149–156.
    https://doi.org/10.1670/14-162
    Campbell, J.A., Brodie Jr., E.D., Caviedes-Solís, I.W., Nieto-Montes de Oca, A., Luja, V.H., Flores-Villela, O., García-Vázquez, U.O., Chandra Sarker, G., Wostl, E. & Smith, E.N. (2018) Systematics of the frogs allocated to Sarcohyla bistincta sensu lato (Cope, 1877), with description of a new species from western Mexico. Zootaxa, 4422 (3), 366–384.
    https://doi.org/10.11646/zootaxa.4422.3.3
    Campbell, J.A., Frost, D.A. & Castoe, T.A. (2019) New generic name for jumping pitvipers (Serpentes:Viperidae). Nota científica. Revista Latinoamericana de Herpetología, 2, 52–53.
    Canseco-Márqquez, L., Aguilar-López, J.L., Luría-Manzano, R., Pineda, E. & Caviedes-Solís, I.W. (2017) A new species of treefrog of the genus Ptychohyla (Anura: Hylidae) from southern Mexico. Zootaxa, 4317 (2), 279–290.
    https://doi.org/10.11646/zootaxa.4317.2.5
    Canseco-Márquez, L. & Ramírez-González, C.G. (2015) New herpetofaunal records for the state of Oaxaca, Mexico. Mesoamerican Herpetology, 2, 363–367.
    Canseco-Márquez, L., Ramírez-González, C.G. & González-Bernal, E. (2017) Discovery of another new species of Charadrahyla (Anura, Hylidae) from the cloud forest of northern Oaxaca, México. Zootaxa, 4329 (1), 64–72.
    https://doi.org/10.11646/zootaxa.4329.1.2
    Canseco-Márquez, L., Ramírez-González, C.G. & Campbell, J.A. (2018) Taxonomic review of the rare Mexican snake genus Chersodromus (Serpentes: Dipsadidae), with the description of two new species. Zootaxa, 4399 (2), 151–169.
    https://doi.org/10.11646/zootaxa.4399.2.1
    Carbajal-Márquez, R.A., Cedeño-Vázquez, J.R., Martínez-Arce, A., Neri-Castro, E. & Machkour-M’Rabet, S.C. (2020) Accessing cryptic diversity in Neotropical rattlesnakes (Serpentes: Viperidae: Crotalus) with the description of two new species. Zootaxa, 4729 (4), 451–481.
    https://doi.org/10.11646/zootaxa.4729.4.1
    Card, D.C., Schield, D.R., Adams, R.H., Corbin, A.B., Perry, B.W., Andrew, A.L., Pasquesi, G.I.M., Smith, E.N., Jezkova, T., Boback, S.M., Booth, W. & Castoe, T.A. (2016) Phylogeographic and population genetic analyses reveal multiple species of Boa and independent origins of insular dwarfism. Molecular Phylogenetics and Evolution, 102, 104‒116.
    https://doi.org/10.1016/j.ympev.2016.05.034
    Casas-Andreu, G., Méndez-de la Cruz, F. & Camarillo, J.L. (1996) Anfibios y reptiles de Oaxaca. Lista, distribución y conservación. Acta taxonómica Mexicana, Nueva Serie, 69, 1–35.
    Casas-Andreu, G., Méndez de la Cruz, F. & Aguilar- Miguel, X. (2004) Anfibios y Reptiles. In: García-Mendoza, A., Ordoňez, J. & Briones-Salas, M.A. (Eds.), Biodiversidad de Oaxaca. Instituto de Biología, UNAM-Fondo Oaxaqueňo para la conservación de la Naturaleza-World Wildlife Fund, México, D.F., Mexico, pp. 375–390.
    Castañeda-Hernández, C., Canseco-Márquez, L. & Vargas-Orrego, M.E. (2015) Additional distributional records for the state of Oaxaca, Mexico. Mesoamerican Herpetology, 2, 368‒370.
    Chambers, E.A. & Hillis, D.M. (2020) The multispecies coalescent over-splits species in the case of geographically widespread taxa. Systematic Biology, 69, 184‒193.
    https://doi.org/10.1093/sysbio/syz042
    Cisneros-Bernal, A.Y., Palacios-Aguilar, R., Medina-Rangel, G., Campillo, G.G. & Riva-Reyes, R. (2020) New distributional records of amphibians and reptiles from the Mixteca region of Oaxaca (Oaxaca, Mexico). North-Western Journal of Zoology, 16, 220‒224.
    Clause, A.G., Pavón-Vázquez, C.J., Scott, P.A., Murphy, C.M., Schaad, E.W. & Gray, L.N. (2016) Identification uncertainty and proposed best-practices for documenting herpetofaunal geographic distributions, with applied examples from southern Mexico. Mesoamerican Herpetology, 3, 977‒1000.
    del Moral-Flores, L.F., Vázquez-Núñez, L.G. & Hernández-Arellano, T. (2019) Primer registro del cocodrilo de pantano, Crocodylus moreletii Duméril & Bibron 1851, en la selva de los Chimalapas, Oaxaca. Revista Latinoamericana de Herpetología, 2 (1), 52–54
    De La Torre-Loranca, M.A., Grünwald, C.I., Valdenegro-Brito, A.E., Cervantes-Burgos, R.I. & García-Vázquez, U.O. (2019) New distributional records for the rare mexican pitviper Cerrophidion petlalcalensis (Squamata: Viperidae) From Veracruz and Oaxaca, México, with comments on the natural history. Revista Latinoamericana de Herpetología, 2, 71–77.
    Downs, F.L. (1967) Intrageneric relationships among colubrid snakes of the genus Geophis Wagler. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 131, 1–193.
    Duellman, W.E., Marion, A.B. & Hedges, S.B. (2016) Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae). Zootaxa, 4104 (1), 1–109.
    https://doi.org/10.11646/zootaxa.4104.1.1
    Ehrlich, P.R. & Ehrlich, A.H. (1996) Betrayal of Science and Reason: How Anti-Environmental Rhetoric Threatens Our Future. Islands Press, Washington, D.C., 335 pp.
    Escobedo-Galván, A., Casas-Andreu, G. & Barrios-Quiroz, G. (2015) On the occurrence of Caiman crocodilus in Oaxaca, Mexico: a misunderstanding for over 140 years. Miscellaneous notes. Mesoamerican Herpetology, 2, 221–223.
    Faivovich, J., Pereyra, M.O., Luna, M.C., Hertz, A., Blotto, B.L., Vásquez-Almazán, C.R., McCranie, J.R., Sánchez, D.A., Baêta, D., Araujo-Vieira, K., Köhler, G., Kubicki, B., Campbell, J.A., Frost, D.R., Wheeler, W.C. & Haddad, C.F.B. (2018) On the monophyly and relationships of several genera of Hylini (Anura: Hylidae: Hylinae), with comments on recent taxonomic changes in hylids. South American Journal of Herpetology, 13, 1–32.
    https://doi.org/10.2994/SAJH-D-17-00115.1
    Gadow, H. (1905) The distribution of Mexican amphibians and reptiles. Proceedings of the Zoological Society of London, 1905, 191‒244.
    https://doi.org/10.1111/j.1469-7998.1905.tb08388.x
    García-Grajales, J., Arrazola Bohórquez, R., Penguilly Macías, M.A. & Buenrostro Silva, A. (2020) New records of Heloderma alvarezi (Wiegmann, 1829) (Sauria: Helodermatidae) on the coast of Oaxaca and increases to its distribution in Mexico. Journal of Threatened Taxa, 12, 15495–15498.
    https://doi.org/10.11609/jott.5691.12.4.15495-15498
    García-Padilla, E., Figel, J.J., Wilson, L.D. & Mata-Silva, V. (2019) Geographic distribution. Bolitoglossa mexicana (Mexican Mushroom-tongued Salamander). Herpetological Review, 50, 744.
    García-Vázquez, U.O., Pavón-Vázquez, C.J., Feria-Ortiz, M. & Nieto-Montes de Oca, A. (2021) A new species of Blue-tailed Skink (Scincidae: Plestiodon) from the Sierra Madre del Sur, Mexico. Herpetologica, 77, 85–93.
    https://doi.org/10.1655/0018-0831-77.1.85
    Gray, L., Meza-Lázaro, R., Poe, S. & Nieto-Montes de Oca, A. (2016) A new species of semiaquatic Anolis (Squamata: Dactyloidae) from Oaxaca and Veracruz, Mexico. Herpetological Journal, 26, 253–262.
    Grünwald, C.I., Jones, J.M., Franz-Chávez & Ahumada-Carrillo, I.T. (2015) A new species of Ophryacus (Serpentes: Viperidae: Crotalinae) from eastern Mexico, with comments on the taxonomy of related pitvipers. Mesoamerican Herpetology, 1, 387–416.
    Grünwald, C.I., Pérez-Rivera, N., Ahumada-Carrillo, I.T., Franz-Chávez, H. & LaForest B.T. (2016) New distributional records for the herpetofauna of Mexico. Herpetological Review, 47, 85–90.
    Gutiérrez-Rodríguez, J., Zaldívar-Riverón, A., Solano-Zavaleta, I., Campbell, J.A., Meza-Lázaro, R.N., Flores-Villela, O. & Nieto-Montes de Oca, A. (2020) Phylogenomics of the Mesoamerican alligator-lizard genera Abronia and Mesaspis (Anguidae: Gerrhonotinae) reveals multiple independent clades of arboreal and terrestrial species. Molecular Phylogenetics and Evolution, 154, 106963.
    https://doi.org/10.1016/j.ympev.2020.106963
    Heimes, P. (2016) Herpetofauna Mexicana Vol. 1. Snakes of Mexico. Edition Chimaira, Frankfurt am Main, 572 pp.
    Hernández-Jiménez, C.A., Flores-Villela, O. & Campbell, J.A. (2019) A new species of patch-nosed snake (Colubridae: Salvadora Baird and Girard, 1853) from Oaxaca, Mexico. Zootaxa, 4564 (2), 588–600.
    https://doi.org/10.11646/zootaxa.4564.2.14
    Himes, J.G. (2020). Geographic distribution. Sceloporus scitulus (Striated Emerald Spiny Lizard). Herpetological Review, 51, 778–779.
    Ibarra-Contreras, C.A., García-Vázquez, U.O., García-Padilla, E., Mata-Silva, V., DeSantis, D.L., Johnson, J.D. & Wilson, L.D. (2016) Phrynosoma orbiculare (Linnaeus, 1758). Mesoamerican Herpetology, 3, 1046‒1047.
    Jadin, R.C., Blair, C., Orlofske, S.A., Jowers, M.J., Rivas, G.A., Vitt, L.J., Ray, J.M., Smith, E.N. & Murphy, J.C. (2020) Not withering on the evolutionary vine: systematic revision of the Brown Vine Snake (Reptilia: Squamata: Oxybelis) from its northern distribution. Organisms Diversity and Evolution, 20, 723–746.
    https://doi.org/10.1007/s13127-020-00461-0
    Jiménez-Arcos, V.H., Calzada-Arciniega, R.A., Alfao-Juantorena, L.A., Vázquez-Reyes, L.D., Blair, C. & Parra-Olea, G. (2019) A new species of Charadrahyla (Anura: Hylidae) from the cloud forest of western Oaxaca, Mexico. Zootaxa, 4554 (2), 371–385.
    https://doi.org/10.11646/zootaxa.4554.2.3
    Johnson, J.D., Mata-Silva, V. & Ramírez-Bautista, A. (2010) Geographic distribution and conservation of the herpetofauna of southeastern Mexico. In: Wilson, L.D., Townsend, J.H. & Johnson, J.D. (Eds.), Conservation of Mesomaerican Amphibians and Reptiles. Eagle Mountain Publishing, LC, Eagle Mountain, Utah, pp. 322–369.
    Johnson, J.D., Mata-Silva, V., García-Padilla, E. & Wilson, L.D. (2015) The herpetofauna of Chiapas, Mexico: composition, distribution, and composition. Mesoamerican Herpetology, 2, 271–329.
    Johnson, J.D., Wilson, L.D., Mata-Silva, V., García-Padilla, E. & DeSantis, D.L. (2017) The endemic herpetofauna of Mexico: organisms of global significance in severe peril. Mesoamerican Herpetology, 4, 543–620.
    Lynch, J.D. & Wake, D.B. (1989) Two new species of Pseudoeurycea (Amphibia: Caudata) from Oaxaca, Mexico. Contributions in Science, Natural History Museum of Los Angeles County, 411, 11–22.
    Mata-Silva, V., Johnson, J.D., Wilson, L.D. & García-Padilla, E. (2015) The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology, 2, 5–62.
    Mata-Silva, V., DeSantis, D.L., García-Padilla, E., Johnson, J.D. & Wilson, L.D. (2019) The endemic herpetofauna of Central America: a casualty of anthropocentrism. Amphibian & Reptile Conservation, 13, 1–64.
    Mata-Silva, V., García-Padilla, E., DeSantis, D.L., Rocha, A., Wilson, L.D., Simón-Salvador, P.R., Mayoral-Halla, C., Montiel-Altamirano, B.F. & Ramírez-Bautista, A. (2017) New herpetofaunal distribution records for the state of Oaxaca, Mexico. Mesoamerican Herpetology, 4, 679‒683.
    Mata-Silva, V., Rocha, A., Mata-González, S., Ramírez-Bautista, A., Berriozabal-Islas, C., García-Padilla, E., Johnson, J.D. & Wilson, L.D. (2019) First municipality records and additional information on the herpetofauna of Oaxaca, Mexico. Herpetological Review, 50, 533‒536.
    Mata-Silva, V., Rocha, A., Ramírez-Bautista, A., Berriozabal-Islas, C. & Wilson, L.D. (2019) A new species of forest snake of the genus Rhadinaea from tropical montane rainforest in the Sierra Madre del Sur of Oaxaca, Mexico (Squamata, Dipsadidae). ZooKeys, 813, 55–65.
    https://doi.org/10.3897/zookeys.813.29617
    McCranie, J.R., Matthews, A.J. & Hedges, B. (2020) A morphological and molecular revision of lizards of the genus Marisora Hedges and Conn (Squamata: Mabuyidae) from Central America and Mexico, with descriptions of four new species. Zootaxa, 4763 (3), 301–353. https://doi.org/10.11646/zootaxa.4763.3.1
    McCranie, J.R., Sunyer, J. & Martínez-Fonseca, J.G. (2019) Comments and updates to “Guía Ilustrada de Anfibios y Reptiles de Nicaragua along with taxonomic and related suggestions associated with the herpetofauna of Nicaragua. Revista Nicaraguense de Biodiversidad, 52, 1–44.
    Mendoza-Henao, A.M., Arias, E., Townsend, J.H. & Parra-Olea, G. (2020) Phylogeny-based species delimitation and integrative taxonomic revision of the Hyalinobatrachium fleischmanni species complex, with resurrection of H. viridissimum (Taylor, 1942). Systematics and Biodiversity, 18, 464–484.
    https://doi.org/10.1080/14772000.2020.1776781
    Meza-Lázaro, R.N. & Nieto-Montes de Oca, A. (2015) Long forsaken species diversity in the Middle American lizard Holcosus undulatus (Teiidae). Zoological Journal of the Linnean Society, 175, 189‒210.
    https://doi.org/10.1111/zoj.12264
    Nieto-Montes de Oca, A., Barley, A.J., Meza-Lázaro, R.N., García-Vázquez, U.O., Zamora-Abrego, J.G., Thomson, R.C. & Leach, A.D. (2016) Phylogenomics and species delimitation in the knob-scaled lizards of the genus Xenosaurus (Squamata: Xenosauridae) using ddRADseq data reveal a substantial underestimation of diversity. Molecular Phylogenetics and Evolution, 106, 241–253.
    https://doi.org/10.1016/j.ympev.2016.09.001
    O’Connell, K.A. & Smith, E.N. (2018) The effect of missing data on coalescent species delimitation and a taxonomic revision of whipsnakes (Colubridae: Masticophis). Molecular Phylogenetics and Evolution, 127, 356‒366.
    https://doi.org/10.1016/j.ympev.2018.03.018
    Palacios-Aguilar, R. & Flores-Vilella, O. (2020) Taxonomic revision and comments on two groups of the genus Coniophanes (Squamata: Dipsadidae). Vertebrate Zoology, 70, 111–124.
    Palacios-Aguilar, R. & García-Vázquez, U.O. (2020) A partial molecular phylogeny of Rhadinaea and related genera (Squamata, Dipsadidae) with comments on the generic assignment of Rhadinaea eduardoi. Zookeys, 943, 145–155.
    https://doi.org/10.3897/zookeys.943.50738
    Parham, J.F., Papenfuss, T.J., Buskirk, J.R., Parra-Olea, G., Chen, J.Y. & Simison, W.B. (2015) Trachemys ornata or not ornata: reassessment of a taxonomic revision for Mexican Trachemys. Proceedings of the California Academy of Sciences, 62, 359–367.
    Parra-Olea, G., García-París, M. & Wake, D.B. (1999) Status of some populations of Mexican salamanders (Amphibia: Plethodontidae). Revista de Biología Tropical, 47, 217–223.
    https://doi.org/10.15517/rbt.v47i1-2.19069
    Parra-Olea, G., Rovito, S.M., García-París, Maisano, J.A., Wake, D.B. & Hanken, M. (2016) Biology of tiny animals: three new species of minute salamanders (Plethodontidae: Thorius) from Oaxaca, Mexico. PeerJ, 4, e2694.
    https://doi.org/10.7717/peerj.2694
    Pavón-Vázquez, C.J., García-Vazquez, U.O., Bryson Jr., R.W., Feria-Ortiz, M., Manríquez-Morán, N.L. & Nieto-Montes de Oca, A. (2018) Integrative species delimitation in practice: revealing cryptic lineages within the short-nosed skink Plestiodon brevirostris (Squamata: Scincidae). Molecular Phylogenetics and Evolution, 129, 242–257. https://doi.org/10.1016/j.ympev.2018.08.020
    Pavón-Vázquez, C.J., Nieto-Montes de Oca, A., Mendoza-Hernández, A.A., Centenero-Alcal, E., Santa Cruz-Padilla, S.A. & Jiménez-Arcos, V.H. (2017) A new species of Plestiodon (Squamata: Scincidae) from the Balsas Basin, Mexico. Zootaxa, 4365 (2), 149‒172.
    https://doi.org/10.11646/zootaxa.4365.2.3
    Ramírez-Reyes, T., Flores-Villela, O., Piňero, D., Lathrop, A. & Murphy, R.W. (2021) Genomic assessment of the Phyllodactylus tuberculosus complex (Reptilia: Phyllodactylidae) in America. Zoologica Scripta, 2021, 1–14.
    https://doi.org/10.1111/zsc.12492
    Reyes-Velasco, J., Adams, R.H., Boissinot, S., Parkinson, C.L., Campbell, J.A., Castoe, T.A. & Smith, E.N. (2020) Genome-wide SNPs clarify lineage diversity confused by coloration in coralsnakes of the Micrurus diastema species complex (Serpentes: Elapidae). Molecular Phylogenetics and Evolution, 147, 106770.
    Rocha, A., Mata-Silva, V., García-Padilla, E., DeSantis, D.L. & Wilson, L.D. (2016) Third known specimen and first locality record in Oaxaca, Mexico, for Tantilla sertula Wilson and Campbell, 2000 (Squamata: Colubridae). Mesoamerican Herpetology, 3, 771‒774.
    Ron, S.R., Venegas, P.J., Ortega-Andrade, H.M., Gagliardi-Urrutia, G. & Salerno, P.E. (2016) Systematics of Ecnomiohyla tuberculosa with the description of a new species and comments on the taxonomy of Trachycephalus typhonius (Anura, Hylidae). Zookeys, 630, 115‒154.
    https://doi.org/10.3897/zookeys.630.9298
    Rovito, S.M., Parra-Olea, G., Recuero, E. & Wake, D.B. (2015) Diversification and biogeographical history of Neotropical plethodontid salamanders. Zoological Journal of the Linnean Society, 175, 167–188.
    https://doi.org/10.1111/zoj.12271
    Rossman, D.A., Ford, N.B. & Seigel, R.A. (1996) The Garter Snakes: Evolution and Ecology. University of Oklahoma Press, Norman, Oklahoma, 332 pp.
    Ruane, S., Bryson Jr., R.W., Pyron, R.A. & Burbrink, F.T. (2014) Coalescent species delimitation in milksnakes (genus Lampropeltis) and impacts on phylogenetic comparative analyses. Systematics Biology, 63, 231–250.
    https://doi.org/10.1093/sysbio/syt099
    Sánchez-García, F., García-Padilla, E., Wilson, L.D. & Mata-Silva, V. (2020) Geographic Distribution. Gerrhonotus ophiurus. Herpetological Review, 51, 776.
    Scarpetta, S., Gray, L., Nieto-Montes de Oca, A., Castañeda, M.R., Herrel, A., Losos, J.B., Luna-Reyes, R., Jiménez Lang, N. & Poe, S. (2015) Morphology and ecology of the Mexican cave anole Anolis alvarezdeltoroi. Mesoamerican Herpetology, 2, 261‒270.
    Schätti, B., Heimes, P., Tillack, F., Kucharzewski, C. & Torres-Pérez Coeto, J. (2020) Pituophis deppei (Duméril, Bibron & Duméril, 1854) or a reassessment of Mexican bullsnakes (Reptilia: Squamata: Colubridae). Vertebrate Zoology, 70, 483‒545.
    Siria-Hernández, C., García-Collazo, R., Chávez Tapia, C. & Mallory, F. (2005) Pseudoleptodeira uribei (Ramírez y Smith, 1992): Adición a la ofidiofauna de Oaxaca, México. Boletín Sociedad Herpetólogica Mexicana, 14, 25–26.
    Smith, H.M. (1959) New and noteworthy reptiles from Oaxaca, Mexico. Transactions of the Kansas Academy of Sciences, 62, 265–272.
    https://doi.org/10.2307/3626381
    Smith, H.M. & Pérez-Higareda, G. (1991) Clara evidencia de la coespecificidad de Geophis dubius y Geophis rostralis (Reptilia: Serpentes). Boletín de la Sociedad Herpetológica Mexicana, 3, 39–40.
    Smith, H.M. & Smith, R.B. (1977) Guide to Mexican Amphisbaenians and Crocodilians, Bibliographic Addendum II. In: Johnson, J. (Ed.), Synopsis of the Herpetofauna of Meìxico. Vol. V. North Bennington, Vermont, pp. 1–187.
    Valdez-Villavicencio, J., García-Padilla, E. & Mata-Silva, V. (2016) Anelytropsis papillosus Cope, 1855 (Squamata: Dibamidae), an overlooked species in the state of Oaxaca, Mexico. Mesoamerican Herpetology, 3, 178‒180.
    Vázquez-Vega, L.F., Canseco-Márquez, L., Acosta-Sánchez, M.M. & Flores-Villela, O.A. (2016) Geophis semidoliatus (Duméril, Bibron & Duméril, 1854). Mesoamerican Herpetology, 3, 514‒515.
    Wallach, V. (2016) Morphological review and taxonomic status of the Epictia phenops species group of Mesoamerica, with description of six new species and discussion of South American Epictia albifrons, E. goudotii, and E. tenella (Serpentes Leptotyphlopidae: Epictinae). Mesoamerican Herpetology, 3, 216‒374.
    Wallach, V. (2020) How to easily identify the flowerpot blindsnake, Indotyphlops braminus (Daudin, 1803), with proposal of a new genus (Serpentes: Typhlopidae). Pod@rcis, 11, 4–12.
    Wilson, L.D. & Townsend, J.H. (2007) A checklist and key to the snakes of the genus Geophis (Squamata: Colubridae: Dipsadinae), with commentary on distribution and conservation. Zootaxa, 1395 (1), 1–31.
    https://doi.org/10.11646/zootaxa.1395.1.1
    Wilson, L.D., Johnson, J.D. & Mata-Silva, V. (2013a) A conservation reassessment of the amphibians of Mexico based on the EVS measure. Amphibian & Reptile Conservation, 7, 97–127.
    Wilson, L.D., Mata-Silva, V. & Johnson, J.D. (2013b) A conservation reassessment of the reptiles of Mexico based on the EVS measure. Amphibian & Reptile Conservation, 7, 1–47.
    Wilson, L.D. & Lazcano, D. (2019) Biology and society: exposing the vital linkages. Biología y Sociedad, Febrero 2019, 29–54.

  2.