Abstract
The genera and species of worldwide wormlions (Diptera: Rhagionidae) are listed, with annotated references to nomenclature, synonymies and generic combinations, type localities, primary type depositories, distribution, and citations for the most recent revisions. The most diverse genera of the family are Vermileo Macquart, Vermipardus Stuckenberg and Lampromyia Macquart, with 13 described species each. The bulk of Vermileonidae diversity, with nearly half of the valid species, remains in the Afrotropical region.
References
Adar, S., Dor, R. & Scharf, I. (2016a) Habitat choice and complex decision making in a trap-building predator. Behavioral Ecology, 27 (5), 1491–1498. https://doi.org/10.1093/beheco/arw071
Adar, S., Scharf, I. & Dor, R. (2016b) The effect of previous experience on trap construction and movement distance in a pit-building predator. The Science of Nature, 103 (9–10), 83. https://doi.org/10.1007/s00114-016-1405-9
Bar-Ziv, M.A., Bega, D., Subach, A. & Scharf, I. (2019) Wormlions prefer both fine and deep sand but only deep sand leads to better performance. Current Zoology, 65 (4), 393–400. https://doi.org/10.1093/cz/zoy065
Bar-Ziv, M.A. & Scharf, I. (2018) Thermal acclimation is not induced by habitat-of-origin, maintenance temperature, or acute exposure to low or high temperatures in a pit-building wormlion (Vermileo sp.). Journal of Thermal Biology, 74, 181–186. https://doi.org/10.1016/j.jtherbio.2018.03.024
Bar-Ziv, M.A., Subach, A., Hirsch-Ionescu, A., Belmaker, J., Zweifler, A. & Scharf, I. (2018) Comparison of wormlions and their immediate habitat under man-made and natural shelters: suggesting factors making wormlions successful in cities. Zoology, 130, 38–46. https://doi.org/10.1016/j.zool.2018.08.004
Bequaert, J. (1922) The Predaceous Enemies of Ants. The Auk, 40 (1), 162. https://doi.org/10.2307/4073749
Beuk, P. & Pape, T. (2013) Fauna Europaea: Vermileonidae. Fauna Europaea. Version 2.6.2. Available from: http://www.faunaeur.org (accessed 24 October 2020)
Bezzi, M. (1898) Contribuzioni alla fauna ditterologica italiana del Dott. Mario Bezzi. II. Ditteri delle Marche degli Abruzzi. Bolletino della Società Entomologica Italiana, 30, 28.
Bezzi, M. (1900) Zur Synonymie und Verbreitung des Psammorcyter vermileo Deg. Wiener Entomologische Zeitung, 19, 56–57.
Bezzi, M. (1926) South African Rhagionidae (Diptera) in the South African Museum. Annals of the South African Museum, 23, 297–324.
Bigot, J.M.F. (1885) [Note on Lampromyia Macquart.]. Bulletin entomologique de la Société Entomologique de France, 6 (5), lxviii.
Blanchard, E. (1840) Histoire naturelle des insectes. Orthoptères, névroptères, hémiptères, hyménoptères, lépidoptères et diptères. P. Dumenil Editeur, Paris, 672 pp. https://doi.org/10.5962/bhl.title.59226
Brauer, F. (1883a) Die Zweifugler des Kaiserlichen Museums zu Wien III, Pars I. Denkschriften der Kaiserlichen Akademie der Wissenschafter, 47, 43–44.
Brauer, F. (1883b) Vermileo Degeeri Macq. Wiener Entomologische Zeitung, 2, 114.
Brauns, A. (1954) Puppen terricoler Dipterenlarven: eine Einführung in die Kenntnis der Ruhestadien bodenlebender Zweiflüglerlarven der Waldbiozönose auf systematischer Grundlage. Musterschmidt Wissenschaftlicher Verlag, Göttingen, 156 pp.
Brunetti, E. (1927) Notes on Malayan Diptera, with descriptions of new species. Journal of the Federated Malay States Museums, 13, 281–309.
Buchner, P. (1940) Uber den Wurmlowen (Vermileo vermileo). Natur und Volk, 70, 116–131.
Carles-Tolrá, M. (2001) Nuevos datos sobre dípteros iberobaleares (Diptera: Orthorrhapha y Cyclorrhapha). Boletín de la Asociación Española de Entomología, 25, 53–95.
Carles-Tolrá, M. (2010) Algunos Dípteros de la Comunidad de Madrid (España) (Insecta: Diptera). Heteropterus Revista de Entomología, 10, 171–176.
Carles-Tolrá, M. (2016) Nemestrinidae, Vermileonidae, Xylophagidae, Borboropsidae, Cryptochetidae y Xenasteiidae: Seis familias de dípteros nuevas para Portugal (Insecta: Diptera). Boletín de la Sociedad Entomológica Aragonesa, 59, 157–158.
Carles-Tolrá, M. & Baéz, M. (2002) Vermileonidae. In: Carles-Tolrá, M. (Ed.), Catálogo de los Diptera de España, Portugal y Andorra (Insecta). Monografías de la Sociedad Entomológica Aragonesa, Zaragoza, pp. 118.
Carles-Tolrá, M. & Cuesta-Segura, A.D. (2020) Vermileo immaculatus sp. n.: a new vermileonid species from Malta. Arquivos Entomolóxicos, 22, 265–277.
Chassagnard, M.T. & Tsacas, L. (1974) Morphologie de la tête et des pièces buccales imaginales de Chrysopilus auratus F. et de Vermileo vermileo de G. (Diptera: Rhagionidae). International Journal of Insect Morphology and Embryology, 3 (1), 13–32. https://doi.org/10.1016/S0020-7322(74)81003-2
Coquillett, D.W. (1904) Reports on Californian and Nevadan Diptera, I. Invertebrata Pacifica, 1, 17–40.
Costa, O.G. (1844a) Descrizione di dodici specie nuove dell’ordine de’Ditteri ed illustrazione di altre quattordici meno ovvie raccolte nella state del 1834. Atti della Reale Accademia delle Scienze, Sezione della Società Reale Borbonica, 5 (2), 81–107.
Costa, O.G. (1844b) Descrizione d’una novella specie del genere Ceroplatus en enumeratione de’Ditteri raccolti ne’viaggi del 1834 e 35. Atti della Reale Accademia delle Scienze, Sezione della Società Reale Borbonica, 5 (2), 109–120.
Coucke, L. (1893) Matériaux pour une étude des leptides de Belgique, y compris ler genres Xylophagus et Coenomyia. Annales de la Societé Entomologique de Belgique, 37, 130.
Courtiller, M.A. (1867) Leptis vermileo (Fabricius). Annales de la Société Linneenne de Maine-et-Loire, 9, 172.
Cuvier, F.G.B. (1817) Le règne animal distribué d’après son organisation, pour servir de base a l’histoire naturelle des animaux et d’introduction a l’anatomie comparée. Tomé III. Chez Déterville, Paris, 618 pp.
De Geer, C. (1752) Rön om Mask-Lejonet. Kungliga Svenska Vetenskapsakademiens Handlingar, 13, 180–265.
De Geer, C. (1776) Memoires pour servir à l’histoire des insectes. Tome Sixième. P. Hesselberg, Stockholm, 608 pp.
DeLeon, D. (1938) Notes on the distribution of Vermileo in the United States and Mexico with a description of a new species (Diptera: Rhagionidae). Entomological News, 49, 167–171.
De Romand, B.E. (1833) Note sur la larva du Leptis vermileo. Annales de la Société Entomologique de France, 2, 498–499.
Devetak, D. (2008a) Wormlion Vermileo vermileo (L.) (Diptera: Vermileonidae) in Slovenia and Croatia. Annales Series Historia Naturalis, 18, 283–286.
Devetak, D. (2008b) Substrate particle size-preference of wormlion Vermileo vermileo (Diptera: Vermileonidae) larvae and their interaction with antlions. European Journal of Entomology, 105 (4), 631–635. https://doi.org/10.14411/eje.2008.085
Devetak, D. & Arnett, A.E. (2015) Preference of antlion and wormlion larvae (Neuroptera: Myrmeleontidae; Diptera: Vermileonidae) for substrates according to substrate particle sizes. European Journal of Entomology, 112 (3), 500–509. https://doi.org/10.14411/eje.2015.052
Doczkal, D. (2017) Kapitel 7.14: Remarkable records of insects (Diptera, Hymenoptera, Lepidoptera) from the Grenzach project. In: Ssymank, A. & Doczkal, D. (Eds.), Biodiversität des südwestlichen Dinkelbergrandes und des Rheintals bei Grenzach-Wyhlen. Naturkundemuseum Mauritianum Altenburg, Altenburg, pp. 821–835.
Dor, R., Rosenstein, S. & Scharf, I. (2014) Foraging behaviour of a neglected pit-building predator: the wormlion. Animal Behaviour, 93, 69–76. https://doi.org/10.1016/j.anbehav.2014.04.020
Dufour, L. (1850) Description et iconographie de quelques diptères de l’Espagne. Annales de la Société Entomologique de France, 2 (8), 131–155.
Ebejer, M.J. (1995) Notes and new records of the larger Brachycera (Diptera) of Malta. The Central Mediterranean Naturalist, 2 (3), 86–96.
Ebejer, M.J., & Gatt, P. (2021) A checklist of the Diptera (Insecta) of the Maltese Islands. Zootaxa, 5018 (1), 1–75. https://doi.org/10.11646/zootaxa.5018.1.1
Ebejer, M.J., Ketanni, K. & Gatt, P. (2019) First records of families and species of Diptera (Insecta) from Morocco. Boletín de la Sociedad Entomológica Aragonesa, 64, 143–153.
Edwards, F.W. (1932) A note on the genus Vermitigris, (Diptera, Rhagionidae). Proceedings of the Royal Entomological Society of London, Series B, Taxonomy, 1 (10), 217–220. https://doi.org/10.1111/j.1365-3113.1932.tb01333.x
Edwards, F.W. (1935) “Worm-lions” from Portugal and the Sudan (Diptera, Rhagionidae), with a description of a new species. Proceedings of the Royal Entomological Society of London, 10 (2), 54–57.
Engel, E.O. (1929) A new species of Lampromyia (Dipt.) from South Africa. Annals of the Transvaal Museum, 13 (3), 172–174.
Engel, E.O. & Cuthbertson, A. (1937) On the biology of some Rhodesian Diptera together with descriptions of three species of Asilidae new to science. Transactions of the Rhodesia Scientific Association, 35, 147–162.
Engel, E.O. & Gill, E.L. (1929) Notes on two larvae of south African Diptera belonging to the families Leptidae and Asilidae. Transactions of the Royal Society of South Africa, 18 (2), 147–162.
Evenhuis, N.L. (1994) Catalogue of the fossil flies of the world (Insecta: Diptera). Backhuys Publishers, Leiden, 310 pp.
Evenhuis, N.L. (2021) The insect and spider collections of the world website. Available from: http://hbs.bishopmuseum.org/codens/ (accessed 20 November 2020)
Fabricius, J.C. (1775) Systema entomologiae: sistens insectorvm classes, ordines, genera, species adiectis synonymis, locis, desriptionibus, observationibus. Kortii, Flensbvrgi et Lipsiae [Flensburg & Leipzig], 832 pp. https://doi.org/10.5962/bhl.title.36510
Fabricius, J.C. (1781) Species insectorvm exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin adjectis observationibus, descriptionibus. Tome. I C.E. Bohnii, Hambvrgi et Kilonii [Hamburg & Kiel], 552 pp. https://doi.org/10.5962/bhl.title.36509
Fabricius, J.C. (1787) Mantissa insectorvm, sistens eorum species nuper detectas, adjectis characteribus genericis, differentiis specificis, emendationibus observationibus. Tom. II. Christ. Gottl. Proft, Hafniae [Copenhagen], 382 pp. https://doi.org/10.5962/bhl.title.36471
Fabricius, J.C. (1794) Entomologica systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tome. IV. Christ. Gottl. Proft, Hafniae [Copenhagen], 472 pp. https://doi.org/10.5962/bhl.title.125869
Fabricius, J.C. (1805) Systema antliatorum: secundum ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Carolus Reichard, Brunsvigae [Brunswick], xiv + 392 pp. https://doi.org/10.5962/bhl.title.15806
Fernández, J.M. (1952) La Lampromyia canariensis Macq. Graellsia, 10, 47–55.
Frederiksen, J. & Hemmingsen, A.M. (1972) Ethological and autecological studies on canarian Vermileonidae (Diptera Brachycera, Rhagionidae: Leptidae). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, 135, 37–59.
GBIF (2020) The Global Biodiversity Information Facility. Available from https://www.gbif.org/what-is-gbif (accessed 18 November 2020)
Greene, C.T. (1926) Descriptions of larvae and pupae of two-winged flies belonging to the family Leptidae. Proceedings of the United States National Museum, 70 (2651), 1–20. https://doi.org/10.5479/si.00963801.70-2651.1
Griffini, A. (1895) Sopra alcuni Leptidi e Tabanidi del Piemonte. Bolletino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino, 10 (218), 1–19.
Griffiths, G.C.D. (1994) Relationships among the major subgroups of Brachycera (Diptera): A critical review. The Canadian Entomologist, 126 (3), 861–880. https://doi.org/10.4039/Ent126861-3
Hafez, M. & El-Moursy, A.A. (1956a) Studies on desert insects in Egypt. I. Field and laboratory investigations on the worm-lion, Vermileo vermileo L. (Diptera: Rhagionidae). Bulletin de la Société Entomologique d’Égypte, 15, 279–299.
Hafez, M. & El-Moursy, A.A. (1956b) Studies on desert insects in Egypt. II. On the general biology of Vermileo vermileo L. (Diptera: Rhagionidae). Bulletin de la Société Entomologique d’Égypte, 15, 333–348.
Hafez, M. & El-Moursy, A.A. (1963) Comparative notes on the habits of the pit-buiding worm-lions and ant-lions. Bulletin de la Société Entomologique d’Égypte, 47, 125–132.
Hemmingsen, A.M. (1963) The Ant-lion-like Sand Trap of the larva of Lampromyia canariensis Macq. (Diptera, Leptidae = Rhagionidae, Vermileoninae). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 125, 237–267.
Hemmingsen, A.M. (1968) A review of the instinctive behavior in the worm-lions Vermileo vermileo L. and Lampromyia pallida Macquart (Diptera, Brachycera, Rhagionidae, Vermileoninae). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, 131, 289–302.
Hemmingsen, A.M. & Nielsen, B.R. (1971) Species differences in ovipository instincts within the Vermileoninae (Diptera Brachycera, Rhagionidae = Leptidae). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, 134, 149–203.
Hennig, W. (1967) Die sogenannten ‘niederen Brachycera’ im baltischen Bernstein (Diptera: Fam. Xylophagidae, Xylomyidae, Rhagionidae, Tabanidae). Stuttgarter Beiträge zur Naturkunde, 174, 1–51.
Hubenov, Z. (2019) The Dipterans (Insecta: Diptera) of the Vrachanska Planina Mountains. In: Bechev, D., Gorgiev, D. (Eds.), Faunistic diversity of Vrachanski Balkan Nature Park. Part 2. Supplement 7. Plovdiv University Press, Plovdiv, pp. 107–166.
James, M.T. (1965) Family Rhagionidae (Leptidae). In: Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, H., Coulson, J.R. (Eds.), A catalog of the Diptera of America north of Mexico. Agriculture Handbook Nº 276. Agricultural Research Service, United States Department of Agriculture, Washington, D.C., pp. 342–348.
Johnson, C.W. & Brues, C.T. (1919) A revised list of the Diptera of Jamaica. Bulletin of the American Museum of Natural History, 41, 441–449.
Katz, N. & Scharf, I. (2018) Habitat geometry and limited perceptual range affect habitat choice of a trap-building predator. Behavioral Ecology, 29 (4), 958–964. https://doi.org/10.1093/beheco/ary046
Katz, N., Shavit, R., Pruitt, J.N. & Scharf, I. (2017) Group dynamics and relocation decisions of a trap-building predator are differentially affected by biotic and abiotic factors. Current Zoology, 63 (6), 647–655. https://doi.org/10.1093/cz/zow120
Katz, N., Subach, A., Pruitt, J.N. & Scharf, I. (2016) Habitat preference of wormlions and their behavioural repeatability under illumination/shade conditions. Ecological Entomology, 41 (6), 716–726. https://doi.org/10.1111/een.12347
Kehlmaier, C. (2014) A new Lampromyia Macquart from Europe (Diptera: Vermileonidae). Zootaxa, 3887 (4), 481–493. https://doi.org/10.11646/zootaxa.3887.4.6
Kehlmaier, C. (2021) DNA barcoding reveals an unexpected diversity in Old World Vermileonidae (Insecta: Diptera). Bonn Zoological Bulletin, 70 (2), 339–349. https://doi.org/10.20363/BZB-2021.70.2.339
Kerr, P.H. (2010) Phylogeny and classification of Rhagionidae, with implications for Tabanomorpha (Diptera: Brachycera). Zootaxa, 2592 (1), 1–133. https://doi.org/10.11646/zootaxa.2592.1.1
Kertész, K. (1908) Catalogus dipterorum Hucusque Descriptorum. Vol. III. G. Engelmann, Budapestini, 367 pp. https://doi.org/10.5962/bhl.title.5141
Kirk-Spriggs, A.H. & McGregor, G. (2009) Disjunctions in the Diptera (Insecta) fauna of the Mediterranean Province and southern Africa and a discussion of biogeographical considerations: ecological overview article. Transactions of the Royal Society of South Africa, 64 (1), 32–52. https://doi.org/10.1080/00359190909519236
Lamarck, J.B.P.A.M. (1816) Histoire naturelle des animaux sans vertèbres: présentant les caractères généraux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales espèces qui s’y rapportent: précédée d’une introduction offrant la détermination des caractères essentiels de l’animal, sa distinction du végétal et des autres corps naturels, enfin, l’exposition des principes fondamentaux de la zoologie. Verdière, Paris, 423 pp. https://doi.org/10.5962/bhl.title.12712
Le Faucheux, M. (1961) Contribution a l’étude du cycle biologique de Vermileo Degeeri Macquart (Diptère: Rhagionidae): Ponte et éclosion des jeunes larves. Bulletin de la Société Scientifique de Bretagne, 36, 133–141.
Leonard, M.D. (1930) A revision of the Dipterous family Rhagionidae (Leptidae) in the United States and Canada. Memoirs of the American Entomological Society, 7, 1–181.
Lindner, E. (1925) Neue exotische Dipteren (Rhagionidae et Tabanidae). Konowia, 4, 20–24.
Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurenti Salvii, Holmiae [Stockholm], 824 pp.
Ludwig, P., Melzer, R.R. & Ehrhardt, V. (2001) Larval morphology and classification of wormlions (Diptera: Vermileonidae). Mitteilungen der Deutschen Gesellschaft für Allgemeine und Angewandte Entomologie, 13, 89–94.
Ludwig, P., Smols, M. & Melzer, R.R. (1996) Die Mundwerkzeuge des Wurmlowen Vermileo vermileo L. und ihre Funktion (Diptera, Vermileonidae). Nachrichtenblatt der Bayerischen Entomologen, 45 (1), 9–14.
Macquart, P.J.M. (1834) Histoire naturelle des insectes. Diptères. Tome premiere. Roret, Paris, 578 pp. https://doi.org/10.5962/bhl.title.14274
Macquart, P.J.M. (1835) Description d’un nouveau genre d’Insectes Diptères de la famille des Notacanthes. Mémoires de la Société des Sciences, de l’Agriculture et des Arts de Lille, 1834, 504–509.
Macquart, P.J.M. (1838) Diptères exotiques nouveaux ou peu connus. Roret, Paris, 323 pp.
Majer, J. (1988) Family Vermileonidae. In: Soós, A., Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Vol. 5. Academiai Kiadó, Budapest, pp. 34–35.
Marchal, L. (1897) Notes d’entomologie biologique sur une excursion en Algérie et en Tunisie. Memoires de la Société Zoologique de France, 10, 5–25.
Marchesi, P., Cotty, A., Burri, A., Fournier, J. & Sierro, A. (2005) Inventaire faunistique du site protégé de Montorge (Sion, Valais). Bulletin de la Murithienne, 123, 61–82.
Meigen, J.W. (1820) Systematische Beschreibung der bekannten Europäischen zweiflügeligen Insekten. Zweiter Theil mit zehn Kupfertafeln. Vol. 2. Friedrich Wilhelm Forstmann und Beaufort Sohn, Aachen, 363 pp. https://doi.org/10.5962/bhl.title.12464
Mik, J. (1887) Ueber Apogon Dufourii Perr. Wiener Entomologische Zeitung, 6, 311–313.
Mik, J. (1894) Dipterologische miscellen. Wiener Entomologische Zeitung, 13, 165.
Miler, K., Yahya, B.E. & Czarnoleski, M. (2019) Substrate moisture, particle size and temperature preferences of trap-building larvae of sympatric antlions and wormlions from the rainforest of Borneo. Ecological Entomology, 44 (4), 488–493. https://doi.org/10.1111/een.12725
Nagatomi, A. (1977) Classification of lower Brachycera (Diptera). Journal of Natural History, 11 (3), 321–335. https://doi.org/10.1080/00222937700770231
Nagatomi, A., Yang, C. & Yang, D. (1999) The Chinese species and the world genera of Vermileonidae (Diptera). Japan Society of Tropical Ecology, 1, 1–155.
Oldroyd, H. (1947) A new Vermitigris (Diptera: Rhagionidae) From Borneo. Proceedings of the Royal Entomological Society of London, Series B, Taxonomy, 16 (1–2), 21–22. https://doi.org/10.1111/j.1365-3113.1947.tb00842.x
Osten Sacken, C.R. (1883) Synonymica concerning exotic dipterology. No. II. Berliner Entomologische Zeitschrift, 27, 295–298. https://doi.org/10.1002/mmnd.18830270219
Ovtshinnikova, O.G. (2000) Male genital musculature of the Vermileonidae (Diptera, Brachycera) and systematic position of the family. Ëntomologicheskoe Obozrenie, 79 (3), 513–521.
Ovtshinnikova, O.G., Wang, Z., Ovchinnikov, A.N., Wang, X. & Krivokhatsky, V.A. (2014) Description of the larva and biological notes on Vermiophis taihangensis Yang & Chen, 1993 (Diptera: Vermileonidae) from China. Zootaxa, 3790 (3), 487–494. https://doi.org/10.11646/zootaxa.3790.3.7
Palm, J. (1869) Beitrag zur Dipterenfauna Tirols. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 30, 395–454.
Papp, Z. & Soltész, Z. (2019) First record of wormlion Vermileo vermileo (Diptera: Vermileonidae) from Greece. Entomologia Hellenica, 28 (1), 5–10.
Pechuman, L.L. (1938) A synopsis of the new world species of Vermileo (Diptera-Rhagionidae). Bulletin of the Brooklyn Entomological Society, 33, 84–89.
Pérez, A.J., Morales, E., Oromí, P. & López, H. (2003) Fauna de artrópodos de Montaña Clara (Islas Canarias) II: Hexápodos (no coleópteros). Vieraea: Folia Scientarum Biologicarum Canariensium, 31, 237–251.
Perris, É. (1852) Seconde excursion dans les Grandes Landes. Lettre adressée à M. Mulsant. Publications de la Société Linnéenne de Lyon, 8 (1), 145–216.
Petersen, W.H. & Baker, C.W. (2006) First record of wormlions (Diptera: Vermileonidae) in Idaho. Journal of the Idaho Academy of Science, 42 (1), 1–6.
Réaumur, R.A.F. de (1753) Histoire du ver-lion mouche. Histoire de l’Académie Royale des Sciences. Avec les Mémoires de Mathématique & de Physique, Pour la Même Année, 1753, 402–419.
Röder, V. (1892) Ueber Apogon Dufourii Perr. nebst einer Berichtigung zu Schiner’s Fauna (die Fliegen). Entomologische Nachrichten, 18, 248–249.
Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol. 1. Genera Italica ordinis dipterorum, ordinatim disposita et distincta et in familias et stirpes aggregata. Ex Tipographia A. Stoccih, Parmae, 226 pp.
Samocha, Y. & Scharf, I. (2020) Comparison of wormlion behavior under man-made and natural shelters: urban wormlions more strongly prefer shaded, fine-sand microhabitats, construct larger pits and respond faster to prey. Current Zoology, 66 (1), 91–98. https://doi.org/10.1093/cz/zoz006
Scharf, I., Daniel, A., MacMillan, H.A. & Katz, N. (2017) The effect of fasting and body reserves on cold tolerance in 2 pit-building insect predators. Current Zoology, 63 (3), 287–294. https://doi.org/10.1093/cz/zow049
Scharf, I. & Dor, R. (2015) The effects of starvation and repeated disturbance on mass loss, pit construction, and spatial pattern in a trap-building predator. Ecological Entomology, 40 (4), 381–389. https://doi.org/10.1111/een.12196
Scharf, I., Gilad, T., Bar-Ziv, M.A., Katz, N., Gregorian, E., Pruitt, J.N. & Subach, A. (2018) The contribution of shelter from rain to the success of pit-building predators in urban habitats. Animal Behaviour, 142, 139–145. https://doi.org/10.1016/j.anbehav.2018.06.010
Scharf, I., Silberklang, A., Avidov, B. & Subach, A. (2020) Do pit-building predators prefer or avoid barriers? Wormlions’ preference for walls depends on light conditions. Scientific Reports, 10 (1), 1–12. https://doi.org/10.1038/s41598-020-67979-3
Schiner, J.R. (1862) Fauna Austriaca. Die Fliegen (Diptera). Nach der analytischen Methode bearbeitet, mit der Characteristik sämmtlicher europäischer Gattungen, der Beschreibung aller in Deutschland vorkommenden Arten und dem Aufzählung aller bisher beschriebenen europäischen Arten. C. Gerold’s Sohn, Wien, 674 pp. https://doi.org/10.5962/bhl.title.8525
Séguy, E. (1926) Faune de France: Diptères (Brachycères) (Stratiomyiidae, Erinnidae, Cœnomyiidae, Rhagionidae, Tabanidae, Codidae, Nemestrinidae, Mydaidae, Bombyliidae, Therevidae, Omphralidae). Paul Lechevalier, Paris, 308 pp.
Séguy, E. (1928) Diptères nouveaux de l’Afrique mineure. Bulletin de la Société Entomologique de France, 33 (3), 45–46.
Séguy, E. (1930) Contributions a l’etude des dipteres du Maroc. Mémoires de la Société des Sciences Naturelles du Maroc, 24, 1–206.
Shelford, R.H. & Shelford, R.S. (1900) Notes from the Sarawak Museum. Journal of the Straits Branch of the Royal Asiatic Society, 33, 256–261.
Strobl, G. (1906) Spanische Dipteren. II. Beitrag. Memorias de la Real Sociedad Española de Historia Natural, 3, 271–422.
Stuckenberg, B.R. (1955) New and little-known South African Rhagionidae (Diptera). Journal of the Entomological Society of Southern Africa, 18 (2), 255–265.
Stuckenberg, B.R. (1960) Diptera (Brachycera): Rhagionidae. South African Animal Life Series, 7, 216–308.
Stuckenberg, B.R. (1961) Records and descriptions of Blepharoceridae, Erinnidae and Rhagionidae from South Africa (Diptera). Annals of the Natal Museum, 15 (11), 109–124.
Stuckenberg, B.R. (1965a) Notes on the Palaearctic species of Vermileo, with the description of a new species from Crete (Diptera: Rhagionidae). Annals and Magazine of Natural History, 8 (92), 495–500. https://doi.org/10.1080/00222936508651603
Stuckenberg, B.R. (1965b) A new species of Lampromyia Macquart (Diptera: Rhagionidae) from the Richtersveld. Journal of the Entomological Society of Southern Africa, 28 (1), 107–109.
Stuckenberg, B.R. (1971) A taxonomic study on the species of Lampromyia Macq. in the Canary Islands (Diptera: Rhagionidae). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, 134, 77–88.
Stuckenberg, B.R. (1995a) A taxonomic revision of Vermipardus Stuckenberg, 1960, with description of new species and notes on the biology and biogeography of the genus (Diptera: Vermileonidae). Annals of the Natal Museum, 36 (1), 215–253.
Stuckenberg, B.R. (1995b) Vermilynx, a new genus for the wormlion fly Lampromyia vanson; Stuckenberg of the Richtersveld, southern Africa (Diptera: Vermileonidae). African Entomology, 3 (1), 29–34.
Stuckenberg, B.R. (1996a) A revised generic classification of the wormlion flies of Southern Africa previously placed in Lampromyia Macquart, with reinstatement of Leptynoma Westwood 1876. Annals of the Natal Museum, 37 (1), 239–266.
Stuckenberg, B.R. (1996b) A new species of the wormlion genus Vermilynx stuckenberg (Diptera: Vermileonidae) from Namaqualand, South Africa. African Entomology, 4 (2), 197–202.
Stuckenberg, B.R. (1997) Three new species of Vermipardus Stuckenberg, and a revised cladogram for the genus with novel biogeographical implications (Diptera: Vermileonidae). Annals of the Natal Museum, 38 (1), 173–196.
Stuckenberg, B.R. (1998a) A revision of the Palaearctic speices of Lampromyia Macquart (Diptera, Vermileonidae), with the description of a new Iberian species and a cladogram for the genus. Bonner Zoologische Beitraege, 48, 67–96.
Stuckenberg, B.R. (1998b) A new Namibian wormlion species, with an account of the biogeography of Leptynoma Westwood s. str. and its association with anthophily in the Fynbos and Succulent Karoo Biomes (Diptera, Vermileonidae). Annals of the Natal Museum, 39 (1), 165–183.
Stuckenberg, B.R. (2000a) Namamyia, a new genus of Vermileonidae (Diptera) from Namaqualand, with a conspectus of the southern African genera and an account of the adaptations for anthophily. Annals of the Natal Museum, 41, 181–202.
Stuckenberg, B.R. (2000b) The Vermileonidae (Diptera) of Namibia, with the description of a new species of Perianthomyia Stuckenberg from the Brandberg Massif. Cimbebasia, 16, 47–54.
Stuckenberg, B.R. (2001) Pruning the tree: a critical review of classifications of the Homeodactyla (Diptera, Brachycera), with new perspectives and an alternative classification. Studia Dipterologica, 8, 3–41.
Stuckenberg, B.R. (2002a) Namaquamyia, a replacement name for a South African genus of Vermileonidae (Diptera). African Invertebrates, 43, 123.
Stuckenberg, B.R. (2002b) A new genus and species of Vermileonidae (Diptera: Brachycera) from Madagascar. Tijdschrift voor Entomologie, 145 (1), 1–8. https://doi.org/10.1163/22119434-900000093
Stuckenberg, B.R. (2003) A redescription of Isalomyia irwini Stuckenberg, the wormlion fly of Madagascar, related to an Arabian species for which the new genus Alhajarmyia is erected (Diptera: Vermileonidae). African Invertebrates, 44 (2), 191–201.
Stuckenberg, B.R. & Fisher, M. (1999) A new species of Lampromyia Macquart, from Oman: the first record of Vermileonidae (Diptera) from the Arabian Peninsula. Annals of the Natal Museum, 40 (1), 127–136.
Swart, V.R., Kirk-Spriggs, A.H. & Copeland, R.S. (2015) A new species of Alhajarmyia Stuckenberg (Diptera: Vermileonidae), the first wormlion fly described from East Africa and its biogeographical implications. Zootaxa, 4044 (4), 556–566. https://doi.org/10.11646/zootaxa.4044.4.5
Teskey, G.E. (1981) Vermileonidae. In: McAlpine, J.E., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R., Wood, D.M. (Eds.), Manual of Nearctic Diptera. Vol. 1. Research Branch, Agriculture Canada, Ottawa, pp. 529–532.
Thompson, F.C. & Pont, A.C. (1993) Systematic Database of Musca names. Theses Zoologica, 20, 1–219.
Tsacas, L. (1962) Recherches sur la structure et le fonctionnement de la tête et des pièces buccales larvaires des Rhagionidae (Diptères). Mémoires du Muséum national d’histoire naturelle. Serie A, Zoology, 27, 147–235.
Verrall, G.H. (1909) British flies vol. V: Stratiomyidae and succeeding families of the Diptera Brachycera of Great Britain. Gurney & Jackson, London, 780 pp. https://doi.org/10.5962/bhl.title.8944
Von Siebold, C.T.E. (1861) Ueber die Larve von Leptis vermileo. Amtlicher Bericht über die Versammlung Deutscher Naturforscher und Aerzte, 35, 105–107.
Walker, F. (1852) Diptera. Part II. In: Saunders, W.W. (Ed.), Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Saunders. Vol. 1. Van Voorst, London, pp. 77–156.
Wang, L., Ding, S., Cameron, S.L., Li, X., Liu, Y., Yao, G. & Yang, D. (2021) Middle Jurassic origin in India: a new look at evolution of Vermileonidae and time-scaled relationships of lower brachyceran flies. Zoological Journal of the Linnean Society, 20, 1–22. https://doi.org/10.1093/zoolinnean/zlab042
Westwood, J.O. (1876) Description of new genera and species of the family Acroceridae. Transactions of the Entomological Society of London, 1876, 505–518.
Wheeler, W.M (1918) Vermileo comstocki, sp. nov., an interesting leptid fly from California. Proceeding of the New England Zoölogical Club, 6, 83–84.
Wheeler, W.M. (1930) Demons of the dust. W.W. Norton & Company, London, 378 pp.
Wheeler, W.M. (1931) A Cuban Vermileo. Psyche, 38 (4), 165–169. https://doi.org/10.1155/1931/21245
Wheeler, W.M. (1934) A specimen of the Jamaican Vermileo. Psyche, 41 (4), 236–237. https://doi.org/10.1155/1934/42124
Wiedemann, C.R.W. (1830) Aussereuropäische zweiflügelige Insekten. Schulzischen Buchhandlung, Hamm, 684 pp. https://doi.org/10.5962/bhl.title.14603
Wiegmann, B.M., Trautwein, M.D., Winkler, I.S., Barr, N.B., Kim, J.W., Lambkin, C., Bertone, M.A., Cassel, B.K., Bayless, K.M., Heimberg, A.M., Wheeler, B.M., Peterson, K.J., Pape, T., Sinclair, B.J., Skevington, J.H., Blagoderov, V., Caravas, J., Kutty, S.N., Schmidt-Ott, U., Kampmeier, G.E., Thompson, F.C., Grimaldi, D.A., Beckenbach, A.T., Courtney, G.W., Friedrich, M., Meier, R. & Yeates, D.K. (2011) Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the United States of America, 108 (14), 5690–5695. https://doi.org/10.1073/pnas.1012675108
Williston, S.W. (1886) On two interesting new genera of Leptidae. Entomologica Americana, 2, 107.
Williston, S.W. (1895) Two remarkable genera of Diptera. The Kansas University quarterly, 4, 107–108.
Williston, S.W. (1900) Supplement. In: Godman, F.D., Salvin, O. (Eds.), Biologia Centrali-Americana. Zoologia-Insecta-Diptera. R.H. Porter, London, pp. 217–248.
Woodley, N.E. (2009) Vermileonidae (worm-lions). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E., Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, pp. 481–483.
Woodley, N.E. & Swart, V.R. (2017) Vermileonidae. In: Kirk-Spriggs, A.H., Sinclair, B.J. (Eds.), Manual of Afrotropical Diptera. Vol. 1. South African Nacional Biodiversity Institute, Pretoria, pp. 865–875.
Yang, C. (1979a) A new genus and species of wormlion from China (Diptera: Rhagionidae). Entomotaxonomia, 1 (2), 83–89.
Yang, C. (1979b) Biological observations on Vermiophis ganquanensis Yang. Entomotaxonomia, 1 (2), 131–140.
Yang, C. (1988) A primary note of Chinese Vermitigris and a new species of Guangxi (Diptera: Vermileonidae). Entomotaxonomia, 10 (3–4), 178–182.
Yang, C. & Chen, H. (1986) A new species of the genus Vermiophis from the Wudang Mountains, China (Diptera: Vermileonidae). Journal of Huazhong Agricultural University, 5 (4), 321–325.
Yang, C. & Chen, H. (1987) Diptera: Vermileonidae. In: Zhang, S. (Ed.), Agricultural insects, spiders, plant diseases and weeds of Xizang. Vol. 2. Xizang People’s Press, Lasa, pp. 157–160.
Yang, C. & Chen, H. (1993) Notes on Vermileonidae and four new species of Vermiophis (Diptera: Brachycera) from China. Entomotaxonomia, 15 (2), 127–136.
Yeates, D.K. (2002) Relationships of extant lower Brachycera (Diptera): a quantitative synthesis of morphological characters. Zoologica Scripta, 31 (1), 105–121. https://doi.org/10.1046/j.0300-3256.2001.00077.x
Zammit-Maempel, G. (1985) Biology and ecology of Ghar Dalam cave, Malta. Atti della Società Toscana di Scienze Naturali, Serie A, 92, 351–374.