Abstract
A compilation of halacarid and hydrachnid mite species found associated with sponges has been carried out based on published records. Altogether 78 halacarid mites species belonging to 15 genera viz. Agaue (9 species), Agauopsis (7 species), Arhodeoporus (1 species), Atelopsalis (1 species), Bradyagaue (1 species), Copidognathus (27 species), Halacarellus (7 species), Halacaropsis (2 species), Halacarus (5 species), Lohmannella (3 species), Maracarus (3 species), Rhombognathides (3 species), Rhombognathus (4 species), Spongihalacarus (1 species), and Thalassarachna (4 species) are reported. Twenty-four hydrachnid mite species belonging to three families viz. Hygrobatidae (2 species), Unionicolidae (20 species) and Pontarachnidae (2 species) are included. Some species of Unionicolidae are mentioned as possible sponge-mite species. Further studies, emphasizing developmental studies of unionicolid mites to get better ideas about associations with freshwater sponges are needed. Molecular sequencing will reveal more cryptic species and improve the quality of re-descriptions of currently recognized species in these sponge associated mites.
References
Abdo, D.A. (2007) Endofauna differences between two temperate marine sponges (Demospongiae; Haposclerida; Chalinidae) from southwest Australia. Marine Biology, 152, 845–854. https://doi.org/10.1007/s00227-007-0736-7
Abé, H. (1998) Rhombognathine Mites. Taxonomy, Phylogeny, and Biogeography. Hokkaido University Press, Sapporo, 219 pp.
Abé, H. (2010) Redescription of the First Japanese Halacarid Mite, Halacarus spongiphilus (Arachnida: Acari: Halacaridae). Species Diversity, 15, 109–117. https://doi.org/10.12782/specdiv.15.109
Abé, H. (2021) Taxonomic review of the subfamily Lohmannellinae (Acari: Trombidiformes: Halacaridae) with morphological comparisons among genera. Zootaxa, 4980 (2), 201–255. https://doi.org/10.11646/zootaxa.4980.2.1
André, M. (1928) Contribution a l’étude des Halacariens des environs de Monaco. Bulletin de l'Institut océanographique de Monaco, 521, 1–15.
André, M. (1929) In: Faune et Flore de la Méditerranée. Herausgag von der commission Internationale pour l’explor Sci. de la Méditerranée. Paris.
André, M. (1937) Description de trois Halacariens de Conchinchie. Bulletin du Muséum National d'Histoire Naturelle, Série 2, 9, 206–211.
André, M. (1946) Halacariens marins. Faune de France, 46, 1–152.
André, M. (1959) Acari I. Contribution a l'étude des halacariens de la Mer Rouge. In: Mission Robert Ph. Dollfus en Égypte. Vol. 26. Institut Français d’Archéologie Orientale, Le Caire, pp. 93–119.
Arndt, W. (1933) Die von Dr. Fritz Haas auf der Schomburgk-Afrika-Expedition 1931/32 gesammelten Süsswasserschwämme. Senckenbergiana, 15 (5–6), 302–309.
Arndt, W. (1936) Die von Dr. A. Monard in Angola gesammelten Süsswasserschwämme. Mit einem Überblick über die Spongillidenfauna Afrikas nach dem gegenwärtigen Stand unserer Kenntnisse. Arquivos do Museu Bocage, 7, 7–35.
Arndt, W. & Viets, K. (1938) Die biologischen (parasitologischen) Beziehingen zwischen Arachnoideen und Spongien. Zeitschrift für Parasitenkunde, Berlin, 10 (1), 67–93. https://doi.org/10.1007/BF02122224
Bartsch, I. (1973) Copidognathus raekor, eine neue Halacaride (Acari) von der Großen Meteorbank. Meteor Forschungs-Ergebnisse, Reihe D, 16, 65–68.
Bartsch, I. (1978) Halacaridae (Acari) aus der Tiefsee des atlantischen Ozeans. Cahiers de Biologie Marine, 19, 47–62.
Bartsch, I. (1979) Halacaridae (Acari) von der Atlantikküste Nordamerikas. Beschreibung der Arten. Mikrofauna des Meeresbodens, 79, 1–62.
Bartsch, I. (1984) Halacaridae (Acari) von den Westindischen Inseln. Bijdragen tot de Dierkunde, 54, 185–196. https://doi.org/10.1163/26660644-05402005
Bartsch, I. (1985) Halacaridae (Acari) from the Strangford Narrows and the Irish Sea. Proceedings of the Royal Irish Academy, Section B, 85, 21–35.
Bartsch, I. (1986) New species of Halacaridae (Acari) from New Zealand. New Zealand Journal of Zoology, 12, 547–560. https://doi.org/10.1080/03014223.1985.10428302
Bartsch, I. (1990) Antarctic Halacaroidea (Acari): Genera Agaue, Bradyagaue, and Halacarellus. Antarctic Research Series, 52, 185–217. https://doi.org/10.1029/AR052p0185
Bartsch, I. (1992a) Halacariden von den Inseln Moorea und Bora Bora, Gesellschaftsinseln (Arachnida: Acari). Senckenbergiana Biologica, 72, 465–488.
Bartsch, I. (1992b) Lohmannella dictyota n. sp. und L. africana n. sp. von der Südhalbkugel (Arachnida: Acari: Halacaridae). Senckenbergiana Biologica, 72, 457–464.
Bartsch, I. (1993a) Synopsis of the Antarctic Halacaroidea (Acari). Synopses of the Antarctic Benthos. Vol. 4. Koeltz, Koenigstein, 176 pp.
Bartsch, I. (1993b) Halacaridae (Acari) von Malaysia. Beschreibung von drei Arten der Gattung Copidognathus. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 11, 45–58.
Bartsch, I. (1993c) Halacarus (Halacaridae, Acari) from south-western Australia. In: Wells, F.E., Walker, D.I., Kirkman, H. & Lethbridge, R. (Eds.), The Marine Flora and Fauna of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 45–71.
Bartsch, I. (1993d) Rhombognathine mites (Halacaridae, Acari) from Rottnest Island. In: Wells, F.E., Walker, D.I., Kirkman, H. & Lethbridge, R. (Eds.), The Marine Flora and Fauna of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 19–43.
Bartsch, I. (1995) Halacarellus obsoletus n. sp. (Arachnida: Acari: Halacaridae), an associate with Antarctic hexactinellid sponges. Mitteilungen aus dem Zoologischen Museum in Berlin, 71, 121–128. https://doi.org/10.1002/mmnz.19950710113
Bartsch, I. (1996a) Agauopsis (Acari, Halacaridae) of the Sevastopol area; supplementary notes on taxonomy and ecology. Revue de Suisse Zoologie, 103, 697–712. https://doi.org/10.5962/bhl.part.79969
Bartsch, I. (1997a) A new species of the Copidognathus tricorneatus group (Acari: Halacaridae) from Western Australia with a review of this species-group. Species Diversity, 2, 155–166. https://doi.org/10.12782/specdiv.2.155
Bartsch, I. (1997b) Arhodeoporus (Acari: Halacaridae) from Rottnest Island, description of three new species. Acarologia, 38, 265–274.
Bartsch, I. (1998a) Halacarinae (Acari, Halacaroidea) from the northwestern Black Sea: A review. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 95, 143–178.
Bartsch, I. (1998b) Copidognathus dictyotellus nom. nov. replacement name for a homonym (Acari: Halacaridae). Species Diversity, 3, 317–318. https://doi.org/10.12782/specdiv.3.317
Bartsch, I. (1999) Halacaridae (Acari) from Western Australia. Four species of Copidognathus. In: Walker, D.I. & Wells, F.E. (Eds.), The Seagrass Flora and Fauna of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 315–331.
Bartsch, I. (2004) The Black Sea halacarid fauna (Halacaridae: Acari): faunal comparison with the Mediterranean, Eastern North Atlantic, North Sea, and Baltic and reflection on its origin. Mitteilungen aus dem Museum für Naturkunde in Berlin, Zoologische Reihe, 80, 143–158. https://doi.org/10.1002/mmnz.4850800202
Bartsch, I. (2005) The rhombognathine fauna of Australia (Rhombognathinae: Halacaridae: Acari) with notes on the fauna of Esperance, Western Australia. In: Wells, F.E., Walker, D.I. & Kendrick, G.A. (Eds.), The Marine Flora and Fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 375–397.
Bartsch, I. (2007) Halacarid mites (Acari: Halacaridae) from Esperance, Western Australia: Notes on taxonomy and faunal distribution of non-Copidognathinae. Records of the Western Australian Museum, 23, 359–392. https://doi.org/10.18195/issn.0312-3162.23(4).2007.359-392
Bartsch, I. (2009) Checklist of marine and freshwater halacarid mite genera and species (Halacaridae: Acari) with notes on synonyms, habitats, distribution and descriptions of the taxa. Zootaxa, 1998 (1), 1–170. https://doi.org/10.11646/zootaxa.1998.1.1
Bartsch, I. (2015) The Agauopsis brevipalpus group (Acari: Halacaridae), descriptions of tropical Indo-West Pacific species, a key to all species, their geographical distribution and reflections on dispersal routes. Acarologia, 55 (2), 147–169. https://doi.org/10.1051/acarologia/20152158
Bartsch, I. (2016a) Commented and illustrated key for identification of Agaue chevreuxi (Trouessart, 1889) and A. panopae (Lohmann, 1893) (Acari, Halacaridae). Ecologica Montenegrina, 8, 45–54. https://doi.org/10.37828/em.2016.8.6
Bartsch, I. (2016b) Antarctic Halacaridae (Acari), new records, these species characteristics and an updated list of species. Polish Polar Research, 37 (1), 131–154. https://doi.org/10.1515/popore-2016-0007
Bartsch, I. (2016c) Arhodeoporus, Camactognathus, Peladognathus, and Winlundia (Acari, Halacaridae) re-evaluation and geographical distribution. Acarologia, 56 (4), 553–571. https://doi.org/10.1051/acarologia/20164141
Bartsch, I. (2020) Lohmannella (Acari, Halacaridae) from a cold-water coral reef off Norway, description of two new and a list of North Atlantic species. Zootaxa, 4722 (3), 277–286. https://doi.org/10.11646/zootaxa.4722.3.5
Beaulieu, S.E. (2001a) Colonization of habitat islands in the deep sea: recruitment to glass sponge stalks. Deep-Sea Res. Part I Oceanogric Research Papers, 48, 1121–1137. https://doi.org/10.1016/S0967-0637(00)00055-8
Beaulieu, S.E. (2001b) Life on glass houses: sponge stalk communities in the deep sea. Marine Biology, 138, 803–817. https://doi.org/10.1007/s002270000500
Bilecenoðlu, M., Alfaya, J.E.F., Azzurro, E., Baldacconi, R., Boyaci, Y., Circosta, V., Compagno, L., Coppola, F., Deidun, A., Durgham, H., Durucan, F., Ergüden, D., Fernández-Álvarez, F., Gianguzza, P., Giglio, G., Gökoðlu, M., Gürlek, M., Ikhtiyar, S., Kabasakal, H., Karachle, V., Katsanevakis, S., Koutsogiannopoulos, D., Lanfranco, E., Micarelli, P., Özvarol, Y., Pena-Rivas, L., Poursanidis, D., Saliba, J., Sperone, E., Tibullo, D., Tiralongo, D., Tripepi, S., Turan, C., Vella, P., Yokeş, M.B. & Zava, B. (2013) New Mediterranean Marine Biodiversity Records (December, 2013). Mediterranean Marine Science, 14, 463–480. https://doi.org/10.12681/mms.676
Böttger, K. (1972) Vergleichend biologisch-ökologische Studien zum Entwicklungszyklus der Süβwassermilben (Hydrachnellae, Acari). II. Der Entwicklungszyklus von Limnesia maculata und Unionicola crassipes. Internationale Revue der gesamten Hydrobiologie, 57 (2), 263–319. https://doi.org/10.1002/iroh.19720570206
Caspers, H. (1951) Quantitative Untersuchungen über die Bodentierwelt des Schwarzen Neeres im Bulgarischen Küstenbereich. Archiv für Hydrobiologie, 45, 1–192.
Chatterjee, T. (2015) A checklist of halacarid and pontarachnid mites (Acari: Halacaridae and Pontarachnidae) associated with mangroves. Ecologica Montenegrina, 4, 334–341. https://doi.org/10.37828/em.2015.2.40
Chatterjee, T. (2020) Parasitic halacarid mites (Acari, Halacaridae): Survey on literature. Acta Biologica, 27, 5–10. https://doi.org/10.18276/ab.2020.27-01
Chatterjee, T. (2021a) A checklist of halacarid mites (Acari: Halacaridae) associated with decapod crustaceans (Arthropoda, Crustacea, Decapoda). Persian Journal of Acarology, 10 (2), 155–166. https://doi.org/10.22073/pja.v10i2.65624
Chatterjee, T. (2021b) A checklist of deep-sea halacarid mites (Acari, Halacaridae) found from more than 1000 m depth. Persian Journal of Acarology, 10 (3), 249–268. https://doi.org/10.22073/pja.v10i3.67563
Chatterjee, T. (2021c) A checklist of halacarid mites (Acari, Halacaridae) associated with echinoderms (Echinodermata). Persian Journal of Acarology, 10 (4), 359–369. https://doi.org/10.22073/pja.v10i4.70790
Chatterjee, T. (2022) A checklist of halacarid mites (Acari, Halacaridae) found among ascidians (Tunicata, Ascidiacea). Persian Journal of Acarology, 11 (in press).
Chatterjee, T. & Durucan, F. (2021) A checklist of halacarid mites (Acari, Halacaridae) found from more than 900 m asl altitudes. Persian Journal of Acarology, 10 (1), 21–40. https://doi.org/10.22073/pja.v10i1.63954
Chatterjee, T. & Durucan, F. (2022) A checklist of halacarid and pontarachnid mites (Acari, Halacaridae & Pontarachnidae) found in seagrass habitats. Persian Journal of Acarology, 11 (in press).
Chatterjee, T. & Sarma, A.L.N. (1991) Review of Indian Ocean Halacaridae and perspectives of future research. In: Gopal, V. & Asthana, V. (Eds.), Aqatic Sciences in India. Indian Association of Limnology and Oceanography, New Delhi, pp 189–198.
Chatterjee, T., Pfingstl, T. & Pešić, V. (2018) A checklist of marine littoral mites (Acari) associated with mangroves. Zootaxa, 4442 (2), 221–240. https://doi.org/10.11646/zootaxa.4442.2.2
Chatterjee, T., Schizas, N.V. & Pešić, V. (2019) A checklist of Pontarachnidae (Acari: Hydrachnidia) and notes on distributional patterns of the species. Zootaxa, 4619 (3), 527–544. https://doi.org/10.11646/zootaxa.4619.3.6
Chislenko, L.L. (1977) Garpaktitsidy (Copepoda, Harpacticoida) s gubok zemli Frantsa-Iosifa. [Harpacticids (Copepoda, Harpacticoida) from sponges of Franz Josef Land.] Biotsenosy shellfa remli Frantsa-Iosifa i fauna sopredel’nykh akuatorii.-Issledouan~a Faunj Morei, 14, 237–276. [in Russian]
Conroy, J.C. (1979) The taxonomy and ecology of Unionicola crassipes crassipes (Müller), a water mite parasitic on the freshwater sponge, Spongilla lacustris (Linné), in Marion Lake, British Columbia. Proceeding of the 4th International Congress of Acarology, 1974, 135–142.
Cook, D.R. (1974) Water Mite Genera and Subgenera. Memoirs of the American Entomological Institute, 21, 1–860.
Cook, D.R. (1980) Studies on Neotropical Water Mites. Memoirs of the American Entomological Institute, Ann Arbor, 31, 1–645.
Corredor, J.E., Wilkinson, C.R., Vicente, V.P., Morell, J.M. & Otero, E. (1987) Nitrate release by Caribbean reef sponges. Limnology Oceanography, 33, 114–120. https://doi.org/10.4319/lo.1988.33.1.0114
Crowell, R.M. (1990) Unionicola billieaehonore, new species, a sponge-associated Hydracarina (Acari: Unionicolidae) from New Zealand. New Zealand Journal of Zoology, 17 (2), 265–270. https://doi.org/10.1080/03014223.1990.10422601
Crowell, R.M. & Davids, C. (1979) Systematics of Unionicola laurentiana, n. sp., and U. nearctica, n. sp., sponge-associated Hydracarina (Parasitengona: Unionicolidae) from North America. The Ohio Journal of Science, 79 (4), 178–186.
Davids, C., Crowell, R.M. & de Groot, C.J. (1985) The developmental cycles of two co-occurring sponge mites Unionicola crassipes (Müller) and Unionicola minor (Soar) (Acari, Hydrachnellae). Hydrobiologia, 122, 199–205. https://doi.org/10.1007/BF00018279
Di Sabatino, A., Smit, H., Gerecke, R., Goldschmidt, T., Matsumoto, N. & Cicolani, B. (2008) Global diversity of water mites (Acari, Hydrachnidia; Arachnida) in freshwater, Hydrobiologia, 595, 303–315. https://doi.org/10.1007/s10750-007-9025-1
Duffy, J.E. (1996) Eusociality in a coral-reef shrimp. Nature, 381, 512–514. https://doi.org/10.1038/381512a0
Durucan, F. (2019) New records of Copidognathus (Acari: Halacaridae) from Antalya, Turkey. Persian Journal of Acarology, 8 (3), 189–210.
Durucan, F. (2020) Mediterranean and Black Sea marine halacarids (Halacaridae: Acari): A review. Journal of Black Sea/Mediterranean Environment, 26, 352–373.
Durucan, F. (2021a) Halacaridae (Prostigmata: Acari) of the Aegean Sea of Turkey (Çanakkale and Izmir). Persian Journal of Acarology, 10 (3), 341–346. https://doi.org/10.22073/pja.v10i3.67699
Durucan, F. (2021b) New records of marine water mites (Acari: Hydrachnidia, Pontarachnidae) from Turkey. Persian Journal of Acarology, 10 (2), 145–153. https://doi.org/10.22073/pja.v10i2.65665
Durucan, F. & Boyaci, Y.Ö. (2017) First record of Atelopsalis pacifica (Acari: Halacaridae) from Turkey. Natural and Engineering Sciences, 2 (3), 10–13. https://doi.org/10.28978/nesciences.348327
Durucan, F. & Boyaci, Y.Ö. (2018) Halacarid mites of the genus Agauopsis (Acari: Halacaridae) from Antalya Turkey. Ege Journal of Fisheries and Aquatic Sciences, 35 (1), 49–53. https://doi.org/10.12714/egejfas.2018.35.1.09
Edwards, D.D. & Vidrine, M.F. (2013) Mites of freshwater mollusks. M. F. Vidrine, Eunice, Louisiana, 336 pp.
Edwards, D.D., Ernsting, B.R. & Vidrine, M.F. (2010) Phylogenetic relationships among Unionicola (Acari: Unionicolidae) mussel-mites of North America based on mitochondrial cytochrome oxidase I sequences. Zootaxa, 2357 (1), 47–57. https://doi.org/10.11646/zootaxa.2537.1.4
Fiore, C.L. & Jutte, P.C. (2010) Characterization of macrofaunal assemblages associated with sponges and tunicates collected off the southeastern United States. Invertebrate Biology, 129, 105–120. https://doi.org/10.1111/j.1744-7410.2010.00184.x
Fishelson, L. (1966) Spirastrella inconstans Dendy (Porifera) as an ecological niche in the littoral zone of the Dahlak Archipelago (Eritrea). Bulletin of the Sea Fisheries Research Station of Israel, 41, 17–25.
Gaino, E., Lancioni, T., Porta , G.L. & Todini, B. (2004) The consortium of the sponge Ephydatia fluviatilis (L.) living on the common reed Phragmites australis in Lake Piediluco (central Italy). Hydrobiologia, 520, 165–178. https://doi.org/10.1023/B:HYDR.0000027735.11246.41
Garcia-Hernandez, J.E., Hammerman, N.M., Cruz-Motta, J.J. & Schizas, N.V. (2019) Associated organisms inhabiting the calcareous sponge Clathrina lutea in La Parguera, Puerto Rico. Caribbean Journal of Science, 49 (2–3), 239–254. https://doi.org/10.18475/cjos.v49i2.a12
Gimbel, O. (1919) Über einige neue Halacariden. Mitteilungen aus dem Zoologischen Museum Hamburg, 36, 105–130.
Gliwicz, Z. & Biesiadka, E. (1975) Pelagic water mites (Hydracarina) and their effect on the plankton community in a neotropical man-made lake. Archiv für Hydrobiologie, 76 (1), 65–88.
Gloeckner, V., Wehrl, M., Moitinho-Silva, L., Gernert, C., Schupp, P., Pawlik, J.R., Lindquist, N.L., Erpenbeck, D., Wörheide, G. & Hentschel, U. (2014) The HMA-LMA dichotomy revisited: an electron microscopical survey of 56 sponge species. Biological Bulletin, 227, 78–88. https://doi.org/10.1086/BBLv227n1p78
Green, J. & MacQuitty, M. (1987) Halacarid Mites. Synopses of the British Fauna, New Series, 36, 1–178.
Greene, A.K. (2008) Invertebrate endofauna associated with sponge and octocoral epifauna at Gary’s National Marine Sanctuary off the coast of Georgia. Ms. Thesis, The Graduate School of the College of Charleston, Charleston, South Carolina, 133 pp.
Gunner, I.E. (1770) Von versehledenen Seeschwämmen. Sehr Norwegian Gesellschaft Drontheim, 4, 65–70.
Hevers, J. (1975) Zur Systematik und Biologie der einheimeschen Unionicola-Arten (Hydrachnellae: Acari). Doctoral Dissertation, University of Kiel, Kiel, 354 pp.
Hevers, J. (1977) Revision der in Deutschland Vorkommenden Arten der Untergattung Unionicola s. str. (Hydrachnellae, Acari). Acarologia, 18 (4), 691–703.
Hevers, J. (1979) Morphologie und Systematik der Nymphen der Unionicola-Arten (Hydrachnellae, Acari) Deutschlands. Osnabrücker Naturwissenschaftliche Mitteilungen, 6, 73–92.
Hevers, J. (1980) Morphologie und Systematik der Larven der Unionicola-Arten (Hydrachnellae, Acari) Deutschlands. Acarologia, 21 (2), 249–266.
Hevers, J. (1984) Die Unionicola crassipes-Gruppe in Japan und in Iran. Deutsche Entomologische Zeitschrift, 31 (4-5), 265–280.
Hicks, G.R.F. (1986) Phylogenetic relationships within the harpacticoid copepod family Peltidiidae Sars, including the description of a new genus. Zoological Journal of the Linnean Society, 88, 349–362. https://doi.org/10.1111/j.1096-3642.1986.tb02252.x
Humes, A.G. (1985) Cnidarians and copepods: a success story. Transactions of the American Microscopical Society, 104, 313–320. https://doi.org/10.2307/3226484
Ho, J.S. (1984) Copepoda associated with sponges, cnidarians, and tunicates of the Sea of Japan. Reports of the Sato Marine Biological Station, Niigata University, 14, 23–61.
Huys, R. (1990) A new harpacticoid copepod family collected from Australian sponges and the status of the subfamily Rhynchothalestrinae Lang. Zoological Journal of the Linnean Society, 99, 51–115. https://doi.org/10.1111/j.1096-3642.1990.tb01222.x
Kersken, D., Göcke, C., Brandt, A., Lejzerowicz, F., Schwabe, E., Seefeldt, M. A., Veit-Köhler, S. & Janussen, D. (2014). The infauna of three widely distributed sponge species (Hexactinellida and Demospongiae) from the deep Ekström Shelf in the Weddell Sea, Antarctica. Deep-Sea Research II, 108, 101–112. https://doi.org/10.1016/j.dsr2.2014.06.005
Kishida, K. (1927) Ushio-dani, Ushio-dani-ka [Halacarus spongiphilus, Halacaridae]. In: Nippon Doubutsu Zukan [Illustrated Encyclopedia of Japanese Animals]. Hokuryukan, Tokyo, pp. 978. [in Japanese]
Klitgaard, A.B. (1991) Gnathia abyssorum (G.O. Sars, 1872) (Crustacea, Isopoda) associated with sponges. Sarsia, 76, 33–39. https://doi.org/10.1080/00364827.1991.10413464
Klitgaard, A.B. (1995) The fauna associated with outer shelf and upper slope sponges (Porifera, Demospongiae) at the Faroe Islands, northeastern Atlantic. Sarsia, 80, 1–22. https://doi.org/10.1080/00364827.1995.10413574
Koukouras, A., Voultsiadou-Koukoura, E., Chintiroglou, H. & Dounas, C. (1985) A comparison of the macrobenthic animal assemblages associated with seven sponge species. Cahiers de Biologie Marine, 26, 301–319.
Koukouras, A., Russo, A., Voultsiadou-Koukoura, E., Dounas, C. & Chintiroglou, C. (1992) Relationship of sponge macrofauna with the morphology of their hosts in the North Aegean Sea. Internationale Revue der gesamten Hydrobiologie, 4, 609–619. https://doi.org/10.1002/iroh.19920770406
Koukouras, A., Russo, A., Voultsiadou-Koukoura, E., Arvanitidis, C. & Stefanidou, D. (1996) Macrofauna associated with sponge species of different morphology. Marine Ecology, 17 (4), 569–582. https://doi.org/10.1111/j.1439-0485.1996.tb00418.x
Krantz, G.W. & Walter, D.E. (2009) A Manual of Acarology. 3rd Edition. Texas Tech, University Press, Lubbock, Texas, 807 pp.
Kunzmann, K. (1992) Die mit ausgewahlten Schwammen (Hexactinellida und Demospongia) aus dem Weddellmeer, Antarktis vergesellschaftete Fauna. Dissertation, Universitat Kiel, Kiel, 108 pp.
Kunzmann, K. (1996) Die mit ausgewählten Schwämmen (Hexactinellida und Demospongiae) aus dem Weddellmeer, Antarktis, vergesellschaftete Fauna (= Associated fauna of selected sponges (Hexactinellida and Demospongiae) from the Weddell Sea, Antarctica. Ber Palarforsch, 210, 1–93.
Leite1, F.P.P., Pavanil, L. & Tanaka, M.O. (2016) Temporal variation of epi- and endofaunal assemblages associated with the red sponge Tedania ignis on a rocky shore (São Sebastião Channel), SE Brazil. Iheringia Série Zoologia, 106. [published online] https://doi.org/10.1590/1678-4766e2016007
Lohmann, H. (1893) Die Halacarinen der Plankton-Expedition. Ergebnisse der Plankton-Expedition der Humboldt Stiftung, Bd. 2, 11–95.
Lohmann, H. (1907a) Über einige faunistische Ergebnisse der Deutschen Südpolar-Expedition unter besonderer Berücksichtigung der Meeresmilben. Schriften des Naturwissenchaftlichen Vereins für Schleswig-Holstein, 14, 1–14.
Lohmann, H. (1907b) Die Meeresmilben der Deutschen Südpolar-Expedition 1901–1903. Deutsche Südpolar-Expedition 1901–1903, 9. G. Fischer, Jena, pp. 361–413.
Long, E.R. (1968) The associates of four species of marine sponges of Oregon and Washington. Pacific Science, 22, 347–351.
Lundbald, O. (1935) An Investigation of some Hispaniolan Lakes. (Dr. R. M. BoNn's Expeditions.) Über einige Hydracarinen aus Haiti. Arkiv för zoologi, Stockholm, 28 A (13), l–30.
Magnino, G., Pronzato, R., Sara, A. & Gaino, E. (1999) Fauna associated with the horny sponge Anomoianthella lamella Pulitzer-Finali & Pronzato, 1999 (Ianthellidae, Demospongiae) from Papua-New Guinea. Italian Journal of Zoology, 66, 175–181. https://doi.org/10.1080/11250009909356253
Mari, M. & Morselli, I. (1990) Idracnelle ed Alacaridi (Acari) di una prateria di Posidonia oceanica (L.) delile dell’isola d’Ischia. Atti della Societa Toscana di Scienze Naturali, Memorie, Serie B, 96, 243–256.
Marliave, J.B., Conway, K.W., Gibbs, D.M., Lamb, A. & Gibbs, C. (2009) Biodiversity and rockfish recruitment in sponge gardens and bioherms of southern British Columbia, Canada. Marine Biology, 156, 2247–2254. https://doi.org/10.1007/s00227-009-1252-8
Morselli, I. & Mari, M. (1985) Ricerche sugli alacaridi delle coste Livornesi. IV.—Osservazioni su alcune specie raccolte su fondi sabbiosi della zona di Piombino. Atti della Societa Toscana di Scienze Naturali, Memorie, Serie B, 91, 201–220.
Morselli, I. & Mari, M. (1993) Alacaridi (Acari, Actinedida) di formazioni coralligene dell’infralitorale laziale (Italia centrale). Atti della Societa Toscana di Scienze Naturali, Memorie, Serie B, 99, 115–123.
Mwase, E.T. & Baker, A.S. (2006) An annotated checklist of (Arachnida: Acari) of Zambia. Zootaxa, 1106 (1), 1–24. https://doi.org/10.11646/zootaxa.1106.1.1
Newell, I.M. (1949) New genera and species of Halacaridae (Acari). American Museum Novitates, 1411, 1–22.
Newell, I.M. (1950) New species of Copidognathus (Acari, Halacaridae) from the Aleutians. American Museum Novitates, 1476, 1–9.
Newell, I.M. (1951a) New species of Agaue and Thalassarachna from the Aleutians (Acari, Halacaridae). American Museum Novitates, 1489, 1–19.
Newell, I.M. (1951b) Further studies on Alaskan Halacaridae (Acari). American Museum Novitates, 1536, 1–56.
Newell, I.M. (1951c) Copidognathus curtus Hall, 1912 and other species of Copidognathus from Western North America (Acari, Halacaridae). American Museum Novitates, 1499, 1–27.
Old, M. (1933) Contribution to the biology of the freshwater sponges (Spongillidae). Papers of the Michigan Academy of Science Arts and Letters, 17, 663–679.
Otero-Colina, G. (1988) Original data on aquatic acarians (Prostigmata, Parasitengona) of southeastern Mexico. Folia entomológica mexicana, 76, 195–224.
Otto, J.C. (1994) New species of Halacaridae from Australia (Acarina: Prostigmata). Acarologia, 35, 31–48.
Otto, J.C. (2000a) Halacaridae from the Great Barrier Reef and Coral Sea: The genera Lohmannella, Scaptognathides and Scaptoganthus (Acarina: Halacaridae: Lohmannellinae). Memoirs of the Queensland Museum, 45, 535–555.
Otto, J.C. (2000b) Spongihalacarus longiscutatus n. gen., n. sp., a marine mite (Acari: Prostigmata: Halacaridae) associated with an alga-sponge symbiosis from the Great Barrier Reef Lagoon in Australia. International Journal of Acarology, 26, 279–283. https://doi.org/10.1080/01647950008684200
Otto, J.C. (2001) Halacaridae of the Great Barrier Reef Lagoon and Coral Sea: The Copidognathus ornatus group (Acarina: Prostigmata: Halacaridae). Memoirs of the Queensland Museum, 46, 717–731.
Pansini, M. (1970) Inquilinismo in Spongia officinalis, Ircinia fasciculate, e Petrosia ficiformis della Riviera Ligure di Levante. Bolletino dei Musei et degli Istituti Biologici della Universita di Genova, 38, 5–17.
Pawlik, J.R. (1983) A sponge-eating worm from Bermuda: Branchiosyllis oculata (Polychaeta, Syllidae). Marine Ecology, 4, 65–79. https://doi.org/10.1111/j.1439-0485.1983.tb00288.x
Pearse, A.S. (1932) Animals in brackish water ponds and pools at Dry Tortugas. Publications of the Carnegie Institution of Washington, 435, 125–142.
Pearse, A.S. (1950) Notes on the inhabitants of certain sponges at Bimini. Ecology, 31, 149–151. https://doi.org/10.2307/1931369
Peattie, M. & Hoare, R. (1981) The sublittoral ecology of the Menai Strait. II. The sponge Halichondria panicea Pallas and its associated fauna. Estuarine Coastal Shelf Science, 13, 621–635. https://doi.org/10.1016/S0302-3524(81)80044-8
Pešić, V., Chatterjee, T., Ingole, B., Velip, D. & Pavićević, A. (2012) A new species of Litarachna Walter, 1925 (Acari, Hydrachnidia) from the West Indian Coast, with a discussion on the biodiversity of the family Pontarachnidae Koenike, 1910. Cahiers de Biologie Marine, 53, 541–546.
Pešić, V., Durucan, F. & Zawal, A. (2019) Marine mites (Acari: Hydrachnidia) of the Mediterranean Sea: Descriptions of two new species, key for identification and future prospects. Zootaxa, 4585 (3), 501–516. https://doi.org/10.11646/zootaxa.4585.3.6
Poore, A.G.B., Huggett, M.J., de Nys, R., Lowry, J.K. & Steinberg, P. (2000) Patterns of host use in alga- and sponge- associated amphipods. Marine Ecology Progress Series, 208, 183–196. https://doi.org/10.1139/z90-183
Proctor, H. & Pritchard, C. (1990) Variability in the life history of Unionicola crassipes, a sponge associated mite (Acari: Unionicollidae). Canadian Journal of Zoology, 68, 1227–1232. https://doi.org/10.1139/z90-183
Ribeiro, Z.M., Omena, E.P. & Muricy, G. (2003) Macrofauna associated to Mycale microsigmatosa (Porifera, Demospongiae) in Rio de Janeiro State, SE Brazil. Estuary, Coastal and Shelf Sciences, 57, 951–959. https://doi.org/10.1016/S0272-7714(02)00425-0
Riesgo, A., Pérez-Portela, R. & Arroyo, N.L. (2010) Halacarid mites (Acari: Halacaridae) associated with a North Atlantic subtidal population of the kelp Laminaria ochroleuca. Journal of Natural History, 44 (11), 651–667. https://doi.org/10.1080/00222930903528222
Rützler, K. (1976) Ecology of Tunisian commercial sponges. Tethys, 7, 249–264.
Santucci, E. (1922) La Geodia cydonium come centro di associazione biologica. Memorie comitato Talassografico Italiano, 103, 1–20.
Sarma, A.L.N. & Chatterjee, T. (1993) Record of Atelopsalis pacifica Bartsch, 1985 (Halacaridae: Acari) from eastern Indian Ocean. Journal of the Bombay Natural History Society, 90, 117–119.
Schejter, L., Chiesa, I.L., Doti, B. & Bremec, C. (2012) Mycale (Aegogropila) magellanica (Porifera: Demospongiae) in the southwestern Atlantic Ocean: endobiotic fauna and new distributional information. Scientia Marina, 76, 753–761. https://doi.org/10.3989/scimar.03490.21A
Schönberg, C.H.L., Hosie, A.M., Fromont, J., Marsh, L. & O’Hara, T. (2016) Apartment-style living on a kebab sponge. Marine Biodiversity, 46 (2), 331–332. https://doi.org/10.1007/s12526-015-0393-x
Schröder, K. (1935) Spongillidenstudien VII Susswasserschwamme von Neuseeland, Borneo und Madagaskar. Zoologischer Anzeiger, 109 (5–6), 97–106.
Schuchert, P. & Reiswig, H.M. (2006) Brinckmannia hexactinellidophila, n. gen., n. sp.: a hydroid living in tissues of glass sponges of the reefs, fjords, and seamounts of Pacific Canada and Alaska. Canadian Journal of Zoology, 84 (4), 564–572. https://doi.org/10.1139/z06-031
Sivadas, S.K., Redij, A. Sagare, P. Thakur, N. & Ingole, B.S. (2014) Temporal variation in macroinvertebrates associated with intertidal sponge Ircinia fusca (Carter 1880) from Ratnagiri, West coast, India. Indian Journal of Marine Sciences, 43, 921–926.
Skilleter, G.A., Russell, B.D., Degnan, B.M. & Garson, M.J. (2005) Living in a potentially toxic environment: comparisons of endofauna in two congeneric sponges from the Great Barrier Reef. Marine Ecology Progress Series, 304, 67–75. https://doi.org/10.3354/meps304067
Smit, H. 2020. Water Mites of the World with keys to the Families, Subfamilies, Genera and Subgenera (Acari: Hydrachnidia). Monografieen van de NedelandseEntomologische Vereniging, 12, 1–774.
Smit, H., van Maanen, B., van den Hoek, T., Wiggers, R. & Knol, B. (2003) New records of rare water mites from the Netherlands (Acari, Hydrachnidia). Nederlandse Faunistische Mededelingen, 18, 123–136.
Soar, C.D. (1900) British freshwater mites. Science-Gossip, London, New Series, 6 (72), 363–365.
Stadler, H. (1935) Pflanzen und Tierwelf der Flußshole des Mains. Verhandlungen des Internationalen Verein Limnologie, 7, 493. https://doi.org/10.1080/03680770.1935.11898577
Stalstedt, J., Bergsten, J. & Ronquist, F. (2013) “Forms” of water mites (Acari: Hydrachnidia): intraspecific variation or valid species? Ecology and Evolution, 3 (10), 3415–3435. https://doi.org/10.1002/ece3.704
Stoll, O. (1887) Arachnida Acaridea. In: Guate-Godman, F.D. & Salvin, O., Centrali-Americana, London, Zoologica, part LIX, pp. 1–55.
Sube, J. (1970) Etude des associes a quelques spongiaires de la region setoise. Reactions tissulaires de hotes vis-à-vis de leurs commensaux et parasites. PhD Thesis, Universite de Montpellier, Montpellier, 90 pp.
Trouessart, E. (1889a) Diagnoses d’espèces et genres nouveaux d’acariens marins (Halacaridae) des côtes de France. Le Naturaliste, 11, 161.
Trouessart, E. (1889b) Sur les acariens marins des côtes de France. Comptes Rendus de l’Académie des Sciences, Paris, 108, 1178–1181. https://doi.org/10.5962/bhl.part.29830
Trouessart, E. (1889c) D’acariens marins (Halacaridae) des côtes de France. Diagnoses d’espèces et genres nouveaux. Le Naturaliste, 11, 181.
Trouessart, E. (1894) Note sur les acariens marins (Halacaridae) dragués par M.P. Hallez dans le Pas-de-Calais. Revue Biologique du Nord de la France, 6, 154–184.esur les
Trouessart, E. (1896) Description d'espèces nouvelles d'acariens marins. Bulletin de la Société Entomologique de France, 1896, 250–253.
Trouessart, E. (1914) Acariens. In: Deuxième Expédition Antarctique Française (1908–1910), commandée par le Dr. Jean Charcot. Sciences naturelles, Documents Scientifiques. Masson, Paris, pp. 1–16.
Vervoort, W. (1964) Freeliving Copepoda from Ifaluk Atoll in the Caroline Islands with notes on related species. Bulletin uf the United States National Museum, 236, 1–431. https://doi.org/10.5479/si.03629236.236.1
Vidrine, M.F. (1984) A new species of mite of the genus Unionicola (Acari: Unionicolidae) from the sponge Trochospongilla leidyi (Bowerbank) in Louisiana. Proceedings of the Louisiana Academy of Sciences, 47, 23–26.
Vidrine, M.F. (1992) Revision of the subgenus Unionicola (Acari: Unionicolidae). The Texas Journal of Science, 44 (4), 459–467.
Vidrine, M.F. (1996) North American Najadicola and Unionicola: diagnoses and distributions. Gail Q. Vidrine Collectibles, Eunice, Louisiana, 356 pp.
Vidrine, M.F. (2019) Louisiana Freshwater Mussels. M. F. Vidrine, Eunice, Louisiana, 261 pp.
Vidrine, M.F. & Edwards, D.D. (2021) Amblemine gill mites. M. F. Vidrine, Eunice, Louisiana, 290 pp.
Vidrine, M.F., Bouchon, R.J. & Poirrier, M.A. (1986) New North American records of water mites (Acari: Unionicolidae: Unionicola: Unionicola and Pentatax). International Journal of Acarology, 12 (4), 229–232. https://doi.org/10.1080/01647958608683469
Vidrine, M.F., Quillman-Vidrine, G.J., Vidrine, M. F.II, Vidrine, D.J. & Vidrine, C.E. (2004) Freshwater Mussels (Bivalvia: Unionidae) in the Cajun Prairie Ecosystem in Southwestern Louisiana. Proceedings of the North American Prairie Conferences, 2004, 77. [https://digitalcommons.unl.edu/napcproceedings/77]
Vidrine, M.F., Borsari, B., Fuller, S.L.H. & Bereza, D.J. (2010) Mites in mollusks in the upper Mississippi river and their community structure. Proceedings of the North American Prairie Conference, 21, 171–178.
Viets, K. (1927a) Die Halacaridae der Nordsee. Zeitschrift für Wissenschaftliche Zoologie, 130, 83–173.
Viets, K. (1927b) Attractides spongicolus n. sp., eine hydracarine aus einem SuBwasserschwamm von Celebes. Zoologischer Anzeiger, 71, 109–112.
Viets, K. (1928) Die Halacariden des Schärgaards bei Bergen (Norwegen). Bergens Museums Årbok 1927, Naturvidenskapelig Rekke, 6, 1–14.
Viets, K. (1935) Die Wassermilben von Sumatra, Java und Bali nach den Ergebnissen der Deutschen Limnologischen Sunda-Expedition. Archiv für Hydrobiologie, Supplement 13, 5 (4), 595–738.
Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv für Naturgeschichte, Neue Folge, 9, 1–135.
Viets, K. (1950) Meeresmilben (Halacaridae, Acari) der Fauna Antarctica. Further Zoological Results of the Swedish Antarctic Expedition 1901–1903, 4 (3), 1–44.
Viets, K. (1953) Die aus Afrika bekannten Wassermilben (Hydrachnellae, Acari). Hydrobiologia, 5, 1–178. https://doi.org/10.1007/BF00023587
Viets, K. (1956) Die Milben des Süßwassers und des Meeres. Katalog der Halacaridae, Meeresmilben. II. Abschnitt. In: Viets, K. (Ed.), Die Milben des Süßwassers und des Meeres. Hydrachnellae et Halacaridae. Fischer Verlag, Jena, pp. 641–870.
Viets, K.H. & Plate, H. (1954) Die Oekologischen (Parasitologischen) Beziehungen zwischen Wassermilben (Hydrachnellae, Acari) und Susswasser-Mollusken. Zeitschrift für angewandte Entomologie, 35, 459–494. https://doi.org/10.1111/j.1439-0418.1954.tb00736.x
Viets, K.O. (1975) Wassermilben (Hydrachnellae, Acari) aus stillgewassen in Guatemala. Studies on Neotropical Fauna and Environment, 10, 57–76. https://doi.org/10.1080/01650527509360482
Voultsiadou-Koukoura, H.E., Koukouras, A. & Eleftheriou, A. (1987) Macrofauna associated with the sponge Verongia aerophoba in the North Aegean Sea. Estuarine, Coastal and Shelf Science, 24, 265–278. https://doi.org/10.1016/0272-7714(87)90069-2
Wendt, P.H., Van Dolah, R.F. & O’Rourke, C.B. (1985) A comparative study of the invertebrate macrofauna associated with seven sponge and coral species collected from the South Atlantic bight. The Journal of the Elisha Mitchell Scientific Society, 3, 187–203.
Wolcott, R.H. (1899) On the North American species of the genus Atax (Fabr.) Bruz. Transactions of the American Microscopical Society, 20, 193–259. https://doi.org/10.2307/3221258
Wulff, J.L. (2006) Ecological interactions of marine sponges. Canadian Journal of Zoology, 84, 145–166. https://doi.org/10.1139/z06-019