Abstract
The following species of the genus Lycoriella Frey sensu Vilkamaa & Menzel from the Holarctic region are newly described and illustrated: Lycoriella ampla sp. n. (Canada: Ontario), L. barkalovi sp. n. (Russia: Krasnoyarsk region), L. canningsi sp. n. (Canada: British Columbia), L. eurystylata sp. n. (Canada: Quebec), L. excavata sp. n. (USA: Colorado), L. jakovlevi sp. n. (Russia: Karelia), L. kinbasketi sp. n. (Canada: British Columbia), L. longa sp. n. (USA: Colorado), L. nivicola sp. n. (Canada: Nunavut, NWT; Greenland), L. pearyi sp. n. (Greenland), L. taimyrensis sp. n. (Russia: Krasnoyarsk region) and L. tundrae sp. n. (Russia: Krasnoyarsk region). Lycoriella parva (Holmgren, 1869) is redescribed and illustrated and new faunistic records are given for some previously described species of Lycoriella. The newly described species raise the number of known species of Lycoriella from 38 to 50.
References
Bertram, G.C.L. & Lack, D. (1938) Notes on the animal ecology of Bear Island. Journal of Animal Ecology, 7 (1), 27–52. https://doi.org/10.2307/1113
BOLD Systems (2021) Barcode of Life Data System. Version 4. Available from: http://v4.boldsystems.org/ (accessed 12 August 2021)
Broadley, A., Kauschke, E. & Mohrig, W. (2018) Black fungus gnats (Diptera: Sciaridae) found in association with cultivated plants and mushrooms in Australia, with notes on cosmopolitan pest species and biosecurity interceptions. Zootaxa, 4415 (2), 201–242. https://doi.org/10.11646/zootaxa.4415.2.1
Coulson, S.J. (2008) The terrestrial and freshwater invertebrate fauna of Svalbard: a check and reference list. Report online 17 October 2008. Self-publishing, Tunebreen, 197 pp. Available from: http://svalbardinsects.net/assets/files/pdf/Full%20report.pdf (accessed 25 October 2020)
Coulson, S.J. (2013) The terrestrial and freshwater invertebrate fauna of Svalbard: a checklist. Report online 6 June 2013. Self-publishing, Tunebreen, 185 pp. + 21 pp. (unpaginated). Available from: https://www.yumpu.com/en/document/view/16680115/checklist-of-the-terrestrial-and-freshwater-invertebrate-fauna-unis (accessed 25 October 2020)
Coulson, S.J. & Refseth, D. (2004) Chapter 3. The terrestrial and freshwater invertebrate fauna of Svalbard (and Jan Mayen). In: Prestrud, P., Strøm, H. & Goldman, H.V. (Eds.), A catalogue of the terrestrial and marine animals of Svalbard. Skrifter, 201. Norwegian Polar Institute, Tromsø, pp. 57–122.
Dobat, K. (1975) Die Höhlenfauna der Schwäbischen Alb mit Einschluß des Dinkelberges, des Schwarzwaldes und des Wutachgebietes. Abhandlungen zur Karst- und Höhlenkunde, Reihe D (Paläontologie, Zoologie), 2, 259–381. [also as Jahreshefte der Gesellschaft für Naturkunde in Württemberg, 130, 259–381]
Edwards, F.W. (1935) LI.—Diptera from Bear Island. Annals and Magazine of Natural History, Series 10, 15, 531–543. https://doi.org/10.1080/00222933508654996
El Ouazzani, N.E.H., Heller, K. & Kettani, K. (2019) The first checklist of black fungus gnats (Diptera: Sciaridae) of Morocco. Annales de la Société entomologique de France, Nouvelle Série, 55 (3), 274–290. https://doi.org/10.1080/00379271.2019.1570826
Freeman, P. (1983) Sciarid flies. Diptera, Sciaridae. Handbooks for the Identification of British Insects, 9 (6), 1–68.
Frey, R. (1948) Entwurf einer neuen Klassifikation der Mückenfamilie Sciaridae (Lycoriidae). II. Die nordeuropäischen Arten. Notulae Entomologicae, 27 (2–4), 33–112.
Gerbachevskaja, A.A. (1969) 25. Sem. Sciaridae (Lycoriidae). In: Bei-Bienko, G.Y. (Ed.), Opredelitel’ nasekomych evropeiskoi chasti SSSR. Tom 5. Chast 1. Dvukrylye, blokhi. Nauka, Leningrad, pp. 320–356. [in Russian]
Gerbachevskaja, A.A. (1988) 25. Family Sciaridae (Lycoriidae). In: Bei-Bienko, G.Y. (Ed.), Keys to the insects of the European part of the USSR. Vol. 5. Diptera and Siphonaptera. Part 1. Amerind Publishing Co. Pvt. Ldt., New Delhi, Bombay, Calcutta and New York, pp. 488–538.
Gerbachevskaja-Pavluchenko, A.A. (1986) Family Sciaridae. In: Soós, Á. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Vol. 4. Sciaridae—Anisopodidae. Akadémiai Kiadó, Budapest, pp. 9, 11–72.
Hackman, W. (1963) Studies on the dipterous fauna in burrows of voles (Microtus, Clethrionomys) in Finland. Acta Zoologica Fennica, 102, 1–64.
Heidari Latibari, M., Moravvej, G., Heller, K., Rulik, B. & Sadeghi Namaghi, H. (2016) New records of Black Fungus Gnats (Diptera: Sciaridae) from Iran, including the reinstatement of Bradysia cellarum Frey. Studia dipterologica, 22 (1), 39–44. [2015]
Heller, K. & Menzel, F. (2017) Fauna Europaea: Sciaridae. In: Beuk, P.L.T. & Pape, T. (Eds.), Fauna Europaea: Diptera, Nematocera. Fauna Europaea, Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin. Database Version 2017.06. Available from: https://fauna-eu.org/cdm_dataportal/taxon/fbaa51a2-6eb4-46e6-ae1a-fbfd243a7950 (accessed 12 August 2021)
Heller, K., Vilkamaa, P. & Hippa, H. (2009) An annotated check list of Swedish black fungus gnats (Diptera, Sciaridae). Sahlbergia, 15 (1), 23–51.
Hippa, H. & Vilkamaa, P. (1991) The genus Prosciara Frey (Diptera, Sciaridae). Entomologica Fennica, 2 (3), 113‒155. https://doi.org/10.33338/ef.83544
Hippa, H., Vilkamaa, P. & Heller, K. (2010) Review of the Holarctic Corynoptera Winnertz, 1867, s. str. (Diptera, Sciaridae). Zootaxa, 2659 (1), 1–197. https://doi.org/10.11646/zootaxa.2695.1.1
Holmgren, A.E. (1869) Bidrag till Kännedomen om Beeren Eilands och Spetsbergens insekt-fauna. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 8 (5), 2–55.
Holmgren, A.E. (1883) Insecta a viris doctissimis Nordenskiöld illum ducem sequentibus in insulis Waigatsch et Novaja Semlia anno 1875 collecta. Hymenoptera et Diptera. Entomologisk Tidskrift, 4 (3‒4), 141‒190.
Jacobson, G. (1898) IV. Zoologicheskiya issledovaniya na Novoy Zemle v 1896 g. Nasekomya Novoy Zemli [Zoological studies in Novaya Zemlya in 1896. Insects of Novaya Zemlya]. Mémoires de l’Académie impériale des sciences de St.-Pétersbourg, Classe physico-mathématique, Series 8, 8 (1), 171‒244. [in Russian]
Jakovlev, E.B. (1994) Dvukrylye palearktiki, svazannye s gribami i myksomycetami [Palaearctic Diptera associated with fungi and myxomycetes]. Karelian Research Center, Russian Academy of Sciences, Forest Research Institute, Petrosavodsk, 127 pp. [in Russian]
Johannsen, O.A. (1912) The fungus gnats of North America, Part IV. Bulletin of the Maine Agricultural Experimental Station, 200, 57‒146. https://doi.org/10.5962/bhl.title.86614
Kolcsár, L.-P. & Heller, K. (2019) First DNA barcodes and records of Sciaridae (Insecta, Diptera) from Romania. Turkish Journal of Zoology, 43 (3), 290–296. https://doi.org/10.3906/zoo-1801-6
Krivosheina, N.P. & Mohrig, W. (1986) Dvukrylye sem. Sciaridae (Diptera) evropeiskoy chasti SSSR [Sciaridae (Diptera) of the European part of the USSR]. Entomologicheskoe Obozrenie, 65 (1), 153‒163. [in Russian]
Lengersdorf, F. (1928–1930) 7. Lycoriidae (Sciaridae). In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. Band 2.1. E. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, pp. 1–71. [1926–1930]
Lengersdorf, F. (1930) Bemerkungen zu den Zetterstedtschen, Staegerschen und Holmgrenschen Sciara-Typen. (Dipt. Sciar.). Deutsche Entomologische Zeitschrift, 1930 (1), 49–56. https://doi.org/10.1002/mmnd.193019300107
Lengersdorf, F. (1940) Beitrag zur Kenntnis der Sciariden (Lycoriiden) aus Finnland. Zoologischer Anzeiger, 131 (1–2), 23–29.
Lindroth, C.H., Andersson, H., Bödvarsson, H. & Richter, S.H. (1973) Surtsey, Iceland. The development of a new fauna, 1963–1970. Terrestrial Invertebrates. Entomologica Scandinavica, Supplementum 5. Munksgaard, Copenhagen, 280 pp.
Lundbeck, W. (1898) Diptera groenlandica. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjøbenhavn, 5 (10), 236–314, 2 pls.
Märkel, F. (1844) IV. Beiträge zur Kenntnis der unter Ameisen lebenden Insekten. Zweites Stück. Zeitschrift für die Entomologie, 5 (1–2), 193–271.
McAlpine, J.F. (1964) Arthropods of the bleakest barren lands: composition and distribution of the arthropod fauna of the northwestern Queen Elizabeth Islands. The Canadian Entomologist, 96 (1–2), 127–129. https://doi.org/10.4039/Ent96127-1
Menzel, F. (2018) Neue Funde von Trauermücken (Insecta: Diptera: Sciaridae) in Deutschland nebst einigen Korrekturen zur Nomenklatur und Systematik. VERNATE, 37, 401–433.
Menzel, F., Gammelmo, Ø., Olsen, K.M. & Köhler, A. (2020), The Black Fungus Gnats (Diptera: Sciaridae) of Norway--Part I: species records published until December 2019, with an updated checklist. ZooKeys, 957, 17–104. https://doi.org/10.3897/zookeys.957.46528
Menzel, F. & Heller, K. (2005) Sechs neue Arten aus den Gattungen Bradysia, Camptochaeta und Corynoptera (Diptera: Sciaridae) nebst einigen Bemerkungen zur Nomenklatur europäischer Trauermücken. Studia dipterologica, 11 (2), 335–357. [2004]
Menzel, F., & Heller, K. (2007) Bemerkungen zur Nomenklatur der Sciariden (Diptera, Bibionomorpha: Sciaridae). Studia dipterologica, 13 (2), 209–229. [2006]
Menzel, F. & Mohrig, W. (1991) Beiträge zur Faunistik und Ökologie des Naturschutzgebietes „Apfelstädter Ried“, Kreis Erfurt-Land. Teil VI—Diptera: Sciaridae. Veröffentlichungen des Naturkundemuseums Erfurt, 10, 27–45.
Menzel, F. & Mohrig, W. (2000) Revision der paläarktischen Trauermücken (Diptera: Sciaridae). Studia dipterologica Supplement, Volume 6 (1999). Ampyx-Verlag, Halle (Saale), 761 pp.
Menzel, F., Mohrig, W. & Barták, M. (2000) Sciaridae. In: Barták, M. & Vaňhara, J. (Eds.), Diptera in an industrially affected region (North-Western Bohemia, Bílina and Duchcov environs). I. Folia Facultatis Scientiarium Naturalium Universitatis Masarykianae Brunensis (Biologia). Vol. 104. Masaryk University, Brno, pp. 73–81.
Menzel, F., Mohrig, W. & Groth, I. (1990) Beiträge zur Insektenfauna der DDR: Diptera – Sciaridae. Beiträge zur Entomologie, 40 (2), 301–400. https://doi.org/10.21248/contrib.entomol.40.2.301-400
Menzel, F., Smith, J.E. & Chandler, P.J. (2006) The sciarid fauna of the British Isles (Diptera: Sciaridae), including descriptions of six new species. Zoological Journal of the Linnean Society, 146 (1), 1–147. https://doi.org/10.1111/j.1096-3642.2006.00190.x
Menzel, F., Vilkamaa, P. & Smith, J.E. (2013) Overview of the Black Fungus Gnats from the Tristan da Cunha archipelago, including a redescription of Hyperlasion viridiventris (Frey) (Diptera, Sciaroidea: Sciaridae). Contributions to Entomology, 63 (2), 283–296. https://doi.org/10.21248/contrib.entomol.63.2.283-296
Mohrig, W. (2003) Black fungus gnats of Central America. Part I. (Diptera, Sciaridae). Beiträge zur Entomologie, 53 (1), 1–69. https://doi.org/10.21248/contrib.entomol.53.1.1-69
Mohrig, W. (2016) Die Trauermücken (Diptera: Sciaridae) von Papua-Neuguinea. Teil IV—Gattungen Bradysia und Chiasmata gen. nov. Studia dipterologica, 22 (1), 3–38. [2015]
Mohrig, W., Heller, K., Hippa, H., Vilkamaa, P. & Menzel, F. (2013) Revision of Black Fungus Gnats (Diptera: Sciaridae) of North America. Studia dipterologica, 19 (1–2), 141–286. [2012]
Mohrig, W., Mamaev, B. & Krivosheina, N. (1983) Beiträge zur Kenntnis der Trauermücken der Sowjetunion (Diptera, Sciaridae). Teil V. Sciariden aus Mittelasien. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 110 (2), 141–155.
Mohrig, W., Krivosheina, N. & Mamaev, B. (1985) Beiträge zur Kenntnis der Trauermücken (Diptera, Sciaridae) der Sowjetunion. Teil VIII: Neue Arten aus europäischen Gebieten. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 112 (3), 299–310.
Mohrig, W., Krivosheina, N. & Mamaev, B. (1990) Beiträge zur Kenntnis der Trauermücken (Diptera, Sciaridae) der Sowjetunion. Teil XIV: Gattungen Plastosciara, Lycoriella und Scatopsciara. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 117 (1), 11–21.
Mukkala, V.-M., Haarto, A., Koponen, S., Mukkala, L., Rinne, V. & Salmela, J. (2005) Ilmajoen Kivistönmäen hyönteisistä, hämähäkkieläimistä ja muista selkärangattomista. W-album. Turun Eläin- ja Kasvitieteellinen Seura ry Hyönteiskerho, 2, 1–34.
Pakalniđkis, S., Bernotienë, R., Lutovinovas, E., Petrađiűnas, A, Podënas, S., Rimđaitë, J., Sżther, O.A. & Spungis, V. (2006) Checklist of Lithuanian Diptera. New and rare for Lithuania insect species, records and descriptions [Naujos ir retos Lietuvos vabzdţiř rűđys, pranešimai ir aprašymai], 18, 16–154.
Pakalniškis, S., Rimđaitë, J., Sprangauskaitë-Bernotienë, R., Butautaitë, R. & Podënas, S. (2000) Checklist of Lithuanian Diptera. Acta Zoologica Lituanica, 10 (1), 3–58. https://doi.org/10.1080/13921657.2000.10512316
Rübsaamen, E.H. (1898) VIII. Grönländische Mycetophiliden, Sciariden, Cecidomyiden, Psylliden, Aphiden und Gallen. Bibliotheca Zoologica, 20 (8), 103–119.
Rudzinski, H.-G. (1989) Der Einfluß von Schadstoffbelastung (Bodenversauerung) und Düngungsmaßnahmen auf die Abundanzdynamik der Trauermücken in Fichtenbeständen des Fichtelgebirges (Diptera, Nematocera: Sciaridae). Nachrichtenblatt der bayerischen Entomologen, 38 (3), 71–78.
Rudzinski, H.-G. (1994) Neue Mitteilungen zur Trauermückenfauna Österreichs (Diptera Nematocera: Sciaridae). Entomofauna, 15 (24), 281–292.
Rudzinski, H.-G. (2003) Die Trauermücken (Insecta: Diptera: Sciaridae) des Naturschutzgebietes Bommecketal in Plettenberg (Sauerland). In: Erbeling, J. & Grundmann, B. (Eds.), Das Bommecketal in Plettenberg (Sauerland). Naturkundliche Monografie eines Naturschutzgebietes. Der Sauerländische Naturbeobachter Nr. 28. Naturwissenschaftliche Vereinigung Lüdenscheid e.V. & Naturschutzzentrum Märkischer Kreis e.V., Lüdenscheid, pp. 190–197.
Salmela, J., Siivonen, S., Dominiak, P., Haarto, A., Heller, K., Kanervo, J., Martikainen, P., Mäkilä, M., Paasivirta, L., Rinne, A., Salokannel, J., Söderman, G. & Vilkamaa, P. (2015) Malaise-pyynti Lapin suojelualueilla 2012–2014 [Malaise-trapping of insects in conservation areas in Lapland within 2012–2014]. Metsähallituksen luonnonsuojelujulkaisuja, Sarja A, 221, 1–141.
Salmela, J. & Vilkamaa, P. (2005) Sciaridae (Diptera) from central Finland: faunistics and taxonomy. Entomologica Fennica, 16 (4), 287–300. https://doi.org/10.33338/ef.84273
Santini, L. & Lucchi, A. (1994) I ditteri Sciaridi nelle colture protette. Informatore Fitopatologico, 9, 15–24.
Schmitz, H. (1919) V.—Zur Kenntnis der Gattung Bradysia Winnertz (Sciaridae, Dipt.). Zoologische Mededeelingen, 5, 25–32.
Schmitz, H. (1920) Eine neue, vielleicht myrmecophile, Sciaride aus den Niederlanden. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere, 43 (1–4), 361–364.
Shin, S., Jung, S., Menzel, F., Heller, K., Lee, H. & Lee, S. (2013) Molecular phylogeny of black fungus gnats (Diptera: Sciaroidea: Sciaridae) and the evolution of larval habitats. Molecular Phylogenetics and Evolution, 66 (3), 833–846. https://doi.org/10.1016/j.ympev.2012.11.008
Steffan, W.A. (1966) A generic revision of the family Sciaridae (Diptera) of America north of Mexico. University of California Publications in Entomology, 44, 1–77.
Stone, A. & Laffoon, J.L. (1965) Family Sciaridae (Lycoriidae). In: Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, R.H. & Coulson, J.R. (Eds.), A catalog of the Diptera of America north of Mexico. Agriculture Handbook 276. United States Department of Agriculture, Washington, D.C., pp. 229–236.
Tuomikoski, R. (1960) Zur Kenntnis der Sciariden (Dipt.) Finnlands. Annales Zoologici Societatis Zoologicae Botanicae Fennicae „Vanamo“, 21 (4), 1–164.
Tuomikoski, R. (1967) Notes on the arthropod fauna of Spitsbergen I. 6. Mycetophilidae and Sciaridae from Spitsbergen, collected by Dr. J. Kaisila in 1965. Annales Entomologici Fennici, 33 (1), 43–51.
Vilkamaa, P. & Hippa, H. (1999) The genus Pnyxiopalpus gen. n. (Diptera: Sciaridae). Systematic Entomology, 24 (3), 209–241. https://doi.org/10.1046/j.1365-3113.1999.00074.x
Vilkamaa, P. & Hippa, H. (2011) Phylogeny of Pnyxiopalpus Vilkamaa & Hippa, with the description of P. persimplex sp. n. (Diptera, Sciaridae). Zootaxa, 2743 (1), 56–62. https://doi.org/10.11646/zootaxa.2743.1.5
Vilkamaa, P., Hippa, H. & Heller, K. (2013) Taxonomy of the Sciaridae (Diptera) of Northern Europe: description of eight new species. Studia dipterologica, 20 (1), 47–58.
Vilkamaa, P. & Menzel, F. (2019) Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa, 4665 (1), 1–67. https://doi.org/10.11646/zootaxa.4665.1.1
Vilkamaa, P., Salmela, J. & Hippa, H. (2007) Black fungus-gnats in deciduous forest habitat in northern Europe, with the description of Bradysia arcula sp. n. (Diptera: Sciaridae). Entomologica Fennica, 18 (4), 226–231. https://doi.org/10.33338/ef.84403
Walker, F. (1848) List of the specimens of dipterous insects in the collection of the British Museum. Part 1. Trustees of the British Museum, London, 229 pp.
Wirta, H., Várkonyi, G., Rasmussen, C., Kaartinen, R., Schmidt, N.M., Hebert, P.D.N., Barták, M., Blagoev, G., Disney, H., Ertl, S., Gjelstrup, P., Gwiazdowicz, D.J., Huldén, L., Ilmonen, J., Jakovlev, J., Jaschhof, M., Kahanpää, J., Kankaanpää, T., Krogh, P.H., Labbee, R., Lettner, C., Michelsen, V., Nielsen, S.A., Nielsen, T.R., Paasivirta, L., Pedersen, S., Pohjoismäki, J., Salmela, J., Vilkamaa, P., Väre, H., von Tschirnhaus, M. & Roslin, T. (2016) Establishing a community-wide DNA barcode library as a new tool for arctic research. Molecular Ecology Resources, 16 (3), 809–822 + 60 unpaginated pp. supported information. https://doi.org/10.1111/1755-0998.12489
Zetterstedt, J.W. (1851) s.n. In: Diptera Scandinaviae disposita et descripta. Vol. 10. ex officina Lundbergiana, sumtibus regiis, Lundae [Lund], pp. 3711–4090.