Abstract
We describe Petracola waka sp. nov. and re-describe the poorly known P. ventrimaculatus, both high-Andean gymnophthalmids from northern Peru. The new species is diagnosed by gracile habitus, smooth dorsal scales, an anterior superciliary that extends onto the dorsum, absence of loreal, three genials, 36–49 transverse dorsal scale rows, 23–28 transverse ventral scale rows, 0–7 femoral pores, absence of subocular-labial fusion, and a color pattern consisting of dark spots or fine lines or bands on a pale brown background. Petracola waka is known from the middle Río Marañon and CajamaraCajabamba basin (Río Crisnejos drainage) from small isolated syngamic systems at 2650–2900 m and is parapatrically distributed with respect to P. ventrimaculatus, which occurs at higher elevations.References
Boulenger, G.A. (1900) Descriptions of new batrachians and reptiles collected by Mr. P.O. Simons in Peru. Annals and Magazine of Natural History, 6, 181–186.
Castoe, T.A., Doan, T.M., & Parkinson, C.L. (2004) Data partitions and complex models in Bayesian analysis: The phylogeny of gymnophthalmid lizards. Systematic Biology, 53, 448–469.
Doan, T.M. & Castoe, T.A. (2003) Using morphological and molecular evidence to infer species boundaries within Proctoporus bolivianus Werner (Squamata: Gymnophthalmidae). Herpetologica, 59, 432–449.
Doan, T.M. & Castoe, T.A. (2005) Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143, 405–416.
Hillis, D.M. (1985) Evolutionary genetics of the Andean lizard genus Pholidobolus (Sauria: Gymnophthalmidae): Phylogeny, biogeography, and a comparison of tree construction techniques. Systematic Zoology, 34, 109–126.
Kizirian, D.A. (1996) A review of Ecuadorian Proctoporus (Squamata: Gymnophthalmidae) with descriptions of nine new species. Herpetological Monographs, 10, 85–155.
Köhler, G. & Lehr, E. (2004) Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with descriptions of three new species and a key to Peruvian species. Herpetologica, 60, 501–518.
Leviton, A.E., Gibbs Jr., R.H., Heal, E. & Dawson., C.E. (1985) Standards in herpetology and ichthyology. Part I. Standard symbolic codes for institutional resource collections in herpetology and ichthyology. Copeia, 1985, 802–832.
Montanucci, R.R. (1973) Systematics and evolution of the Andean lizard genus Pholidobolus (Sauria: Teiidae). University of Kansas Museum of Natural History Miscellaneous Publications, 59, 1–52.
Rowe, J.H. (1946) Inca culture at the time of the Spanish conquest. In: Steward, J.H. (ed.), The Andean Civilizations, Handbook of South American Indians, vol. 2,. Bureau of American Ethnology, Smithsonian Institution, Washington, D.C., pp.183–330.
Uzzell, T. (1970) Teiid lizards of the genus Proctoporus from Bolivia and Peru. Postilla, 142, 1–39.