Abstract
This checklist contains taxonomic information and distributions for 720 drosophilid species, including all species descended from the most recent common ancestor of the Hawaiian Drosophilidae and all adventive species present in the Hawaiian Islands. The ancestor of the Hawaiian Drosophilidae colonized the archipelago roughly 25 million years ago and now includes 689 described taxa. This includes species placed in the genus Scaptomyza (273 spp), only some of which are Hawaiian endemics (148 spp), and all endemic Hawaiian Drosophila (416 spp). There are also 33 adventive species that have been introduced to Hawaii in the past ~200 years. Taxonomic placement, to the level of species subgroup, and all references related to replacement names and synonyms are included. This is the first comprehensive list to be published in over a decade and includes many recent changes and additions to the fauna, including 130 new species names. This checklist will serve as the basis for future revisionary work on the endemic Hawaiian Drosophilidae, particularly the genus Scaptomyza.
References
Ahearn, J.N. & Baimai, V. (1987) Cytogenetic study of three closely related species of Hawaiian Drosophila. Genome, 29, 47–57. https://doi.org/10.1139/g87-008
Asquith, A. & Messing, R.H. (1992) Attraction of Hawaiian Ground Litter Invertebrates to Protein Hydrolysate Bait. Environmental Entomology, 21, 1022–1028. https://doi.org/10.1093/ee/21.5.1022
Bächli, G. (1986) Die Drosophiliden-Arten (Diptera) in der Sammlung des Naturhistorischen Museums Wien. Annalen des Naturhistorischen Museums in Wien. Serie B für Botanik und Zoologie, 90, 131–148.
Bächli, G. (1987) List of type specimens of Drosophilidae (Diptera) in the collections of the Zoological Museum, Amsterdam. Bulletin. Zoologisch Museum Universiteit van Amsterdam, 11, 81–93.
Bächli, G. (2021) TaxoDros: the database on taxonomy of Drosophilidae. Available from: https://www.taxodros.uzh.ch (accessed 5 December 2021)
Bächli, G. & Vilela, C.R. (1991) On the identity of Drosophila immatura Walker and Drosophila remota Walker (Diptera, Drosophilidae). Mitteilungen der Schweizerischen Entomologischen Gesellshaft, 64, 27–31.
Bächli, G., Vilela, C.R., Escher, A.S. & Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. Leiden, Brill, 362 pp. https://doi.org/10.1163/9789047414681
Basden, E.B. (1976) Oswald Duda and Scaptomyza, 1921 (Dipt., Drosophilidae). Entomologist’s Monthly Magazine, 111, 185.
Becker, T. (1908) Dipteren der Kanarischen Inseln. Mitteilungen aus dem Zoologischen Museum in Berlin, 4, 1–180.
Becker, T. (1919) Diptèrés. Brachycères. In: Mission du Service geographique de l’Armée. Mésure d’un Arc de Méridien Equatorial en Amérique du Sud, 1899–1906. Gauthier-Villard, Paris, pp. 163–215, pls. 14–17. https://doi.org/10.5962/bhl.title.980
Bezzi, M. (1928) Diptera brachycera and athericera of the Fiji Islands based on material in the British Museum (Natural History). British Museum (Natural History), London, 220 pp.
Bock, I.R. (1986) The Drosophilidae (Insecta: Diptera) of Norfolk Island. Australian Journal of Zoology, 34, 305–313. https://doi.org/10.1071/ZO9860305
Bonacum, J. (2001) Molecular Systematics of the Hawaiian Drosophilidae. PhD Thesis, Yale University, New Haven, Connecticut, 168 pp.
Bonacum, J., O’Grady, P.M., Kambysellis, M. & DeSalle, R. (2005) Phylogeny and age of diversification of the planitibia species group of the Hawaiian Drosophila. Molecular Phylogenetics and Evolution, 37, 73–82. https://doi.org/10.1016/j.ympev.2005.03.008
Brake, I. & Bächlli, G. (2008) Drosophilidae (Diptera). World Catalogue of Insects, 9, 1–412. https://doi.org/10.1163/9789004261037
Brauer, F. (1883) Die Zweiflugler des Kaiserlichen Museums zu Wien. III. Systematische Studien auf Grundlage der Dipteren-Larven nebst einer Zusammenstellung von Beispielen aus der Literatur uber dieselben und Beschreibung neuer Formen. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse, 47, 1–100.
Brischke, C.G.A. (1881) Die Blattminirer in Danzig’s Umgebung. Schriften der Naturforschenden Gesellschaft in Danzig, 5, 233–290.
Brncic, D. (1955) The Chilean species of Scaptomyza Hardy (Diptera, Drosophilidae). Revista Chilena de Entomologia, 4, 237–250.
Brncic, D. (1983) A review of the genus Scaptomyza Hardy (Diptera, Drosophilidae) in Chile with the description of a new species. Revista Chilena de Historia Natural, 56, 71–76.
Bryan, E.H. (1934) A Review of the Hawaiian Diptera, with Descriptions of New Species. Proceedings of the Hawaiian Entomological Society, 8, 399–468.
Bryan, E.H. (1938) Key to the Hawaiian Drosophilidae and descriptions of new species. Proceedings of the Hawaiian Entomological Society, 10, 25–42.
Burla, H. (1954a) Distinction between four species of the “melanogaster” group, “Drosophila seguyi”, “D. montium”, “D. kikkawai” sp. n. and “D. auraria” (Drosophilidae, Diptera). Revista Brasileira de Biologia, 14, 41–54.
Burla, H. (1954b) Zur Kenntnis der Drosophiliden der Elfenbeinkuste (Franzosisch West-Afrika). Revue Suisse de zoologie, 61, 1–218. https://doi.org/10.5962/bhl.part.75413
Burla, H. (1957) Ostafrikanische drosophiliden (Dipt.). Jahreshefte des Vereins für Vaterlandische Naturkunde in Wurttemberg, 112, 36–49.
Carson, H.L. (1965) Chromosomal morphism in geographically widespread species of Drosophila. In: Baker, H.G. & Stebbins, G.L. (Eds.), The Genetics of Coloninizg Species. Academic Press, New York, New York, pp. 503–531.
Carson, H.L. (1966) Chromosomal races of Drosophila crucigera from the islands of Oahu and Kauai, State of Hawaii. University of Texas Publications, 6615, 405–412.
Carson, H.L., Hardy, D.E., Spieth, H.T. & Stone, W.S. (1970) The Evolutionary Biology of the Hawaiian Drosophilidae. In: Hecht, M.K. & Steere, W.C. (Eds.), Essays in Evolution and Genetics in Honor of Theodosius Dobzhansky. Appleton-Century-Crofts, New York, New York, pp. 437–543. https://doi.org/10.1007/978-1-4615-9585-4_15
Carson, H.L. & Kaneshiro, K.Y. (1976) Drosophila of Hawaii: systematics and ecological genetics. Annual Review of Ecology and Systematics, 7, 311–345. https://doi.org/10.1146/annurev.es.07.110176.001523
Carson, H.L. & Stalker, H.D. (1968a) Polytene Chromosome Relationships in Hawaiian Species of Drosophila. I. The D. grimshawi subgroup. The University of Texas Publication, 6818, 335–354.
Carson, H.L. & Stalker, H.D. (1968b) Polytene Chromosome Relationships in Hawaiian Species of Drosophila. II. The D. planitibia subgroup. The University of Texas Publication, 6818, 355–365.
Carson, H.L. & Stalker, H.D. (1968c) Polytene Chromosome Relationships in Hawaiian Species of Drosophila. III. The D. adiastola and D. punalua subgroups. The University of Texas Publication, 6818, 367–380.
Carson, H.L., Val, F.C. & Wheeler, M.R. (1983) Drosophilidae of the Galapagos Islands, with descriptions of two new species. International Journal of Entomology, 25, 239–248.
Chen, H.Z. (1994) Five new species and a new record of drosophilid flies (Diptera, Drosophilidae) from Sichuan province, China. Journal of Fudan University (Natural Science), 33, 335–347.
Collart, A. (1937) Les Zaprionus du Congo belge (Diptera: Drosophilidae). Bulletin du Musee Royale d’Histoire Naturelle de Belgique, 13, 1–15.
Collin, J.E. (1953) On the British species of Scaptomyza Hardy and Parascaptomyza Duda (Diptera, Drosophilidae). The Entomologist, 86, 148–151.
Coquillett, D.W. (1901) Three New Species of Diptera. Entomological News, 12, 16–18.
Craddock, E.M. & Johnson, W.E. (1979) Genetic Variation in Hawaiian Drosophila. V. Chromosomal and Allozymic Diversity in Drosophila silvestris and its Homosequential Species. Evolution, 33, 137–155. https://doi.org/10.2307/2407372
Craddock, E.M. & Johnson, W.E. (1981) Genetic Variability in a montane rain forest. In: Mueller-Dombois, D., Bridges, K.W. & Carson, H.L. (Eds.), Island ecosystems: biological organization in selected Hawaiian communities. Hutchinson Ross, Stroudsburg, pp. 445–451.
Craddock, E.M., Kambysellis, M.P., Franchi, L., Francisco, P., Grey, M., Hutchinson, A., Nanhoo, S. & Antar, S. (2018) Ultrastructural variation and adaptive evolution of the ovipositor in the endemic Hawaiian Drosophilidae. Journal of Morphology, 279, 1725–1752. https://doi.org/10.1002/jmor.20884
De, A. & Gupta, J.P. (1996) Records of drosophilid species from West Bengal with description of one new and two previously unrecorded species from India (Insecta: Diptera: Drosophilidae). Senckenbergiana Biologica, 76, 129–133.
Dobzhansky, T. & Pavan, C. (1943) Studies on Brazilian species of Drosophila. Boletim. Faculdade de Filosofia, Ciencias e Letras, Universidade de Sao Paulo, 36, 7–72.
Doleschall, C.L. (1858) Derde bijdrage tot de kennis der Dipteren fauna van Nederlandsch Indie. Natuurkundig tijdschrift voor Nederlandsch Indie (Natuurkundige Vereeniging in Nederlandsch Indie), 17, 73–128.
Duda, O. (1921) Kritische Bemerkungen zur Gattung Scaptomyza Hardy (Dipteren). Jahresheft des Vereins fur schlesische Insektenkunde zu Breslau, 13, 57–69.
Duda, O. (1923) Die orientalischen und australischen Drosophiliden-Arten (Dipteren) des ungarischen National-Museums zu Budapest. Annales Historico-Naturales Musei Nationalis Hungarici [Termeszettudomanyi Muzeum evkonyve], 20, 24–59.
Duda, O. (1924) Beitrag zur Systematik der Drosophiliden unter besonderer Berucksichtigung der palaarktischen u. orientalischen Arten (Dipteren). Archiv fur Naturgeschichte, 90 (A), 3, 172–234.
Duda, O. (1927) Die südamerikanischen Drosophiliden (Dipteren) unter Berücksichtigung auch der anderen neotropischen sowie der nearktischen Arten. Archiv für Naturgeschichte, 91 (A)11/1, 1–228.
Duda, O. (1935) Drosophilidae. In: Lindner, E. (Ed.), Die Fliegen der Palearktischen Region. E. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, pp. 65–118.
Duda, O. (1939) Revision der afrikanischen Drosophiliden (Diptera). I. Annales Historico-Naturales Musei Nationalis Hungarici [Termeszettudomanyi Muzeum evkonyve], 32, 1–57.
Escher, S., Ekenstedt, J., Elberg, K. & Saura, A. (2006) The Drosophilidae (Diptera) of Estonia. Entomologica Fennica, 17, 13–20. https://doi.org/10.33338/ef.84283
Escher, S.A., Ekenstedt, J., Karpa, A. & Saura, A. (2002) The Drosophilidae (Diptera) of Latvia. Latvijas Entomologs, 39, 68–76.
Escher, S.A., Ekenstedt, J., Pakalniđkis, S., Rimđaitë, J. & Saura, A. (2004) The Drosophilidae (Diptera) of Lithuania. Acta Zoologica Lituanica, 14, 48–55. https://doi.org/10.1080/13921657.2004.10512579
Fallén, C.F. (1823) Diptera Sveciae. Geomyzides. Berling, Lundae, 8 pp.
Fartyal, R.S. & Singh, B.K. (2001) List of Drosophilid Species so far described and recorded from Kumaon region, India. Drosophila Information Service, 83, 71–73.
Frey, R. (1945) Tiergeographische Studien über die Dipterenfauna der Azoren. I. Verzeichnis der bisher von den Azoren bekannten Dipteren. Societas Scientiarum Fennica. Commentationes Biologicae VIII, 10, 1–114.
Frey, R. (1954) Diptera Brachycera und Sciaridae von Tristan da Cunha. Results of the Norwegian Expedition to Tristan da Cunha, 26, 1–55.
Goñi, B. & Vilela, C.R. (2016) Two new Neotropical species of Drosophilinae (Diptera: Drosophilidae) from Uruguay. Zoologia, 33, 1–13. https://doi.org/10.1590/s1984-4689zool-20160142
Gornostaev, N.G. (2018) An Annotated list of the drosophilid flies (Diptera: Drosophilidae) of Chukotka. Far Eastern Entomologist, 368, 16–19. https://doi.org/10.25221/fee.368.3
Gornostaev, N.G., Kulikov, A.M., Sorokina, S.Y. & Tyukmaeva, V.I. (2018) Review of the drosophilid flies (Diptera: Drosophilidae) of Kamchatka. Far Eastern Entomologist, 359, 16–20. https://doi.org/10.25221/fee.359.4
Grimaldi, D.A. (1987) Amber Fossil Drosophilidae (Diptera), with Particular Reference to the Hispaniolan Taxa. American Museum Navitates, 2880, 1–23.
Grimaldi, D.A. (1990) New Distributional Records of Scaptomyza (Bunostoma) australis from South Pacific Islands and Biogeographic Implications. Journal of the New York Entomological Society, 98, 484–488.
Grimshaw, P.H. (1901) Part I. Diptera. In: Sharpe, D. (Ed.), Fauna Hawaiiensis. Vol. 3 (1). University Press, Cambridge, pp. 1–77.
Grimshaw, P.H. (1902) Diptera (Suppl.). Fam. Drosophilidae. In: Sharpe, D. (Ed.), Fauna Hawaiiensis. Vol. 3 (2). University Press, Cambridge, pp. 79–86.
Gruner, D.S. (2004) Arthropods From Ohi’a Lehuna (Myrtaceae: Metrosideros polymorpha), With New Records for the Hawaiian Islands. Bishop Museum Occasional Papers, 78, 33–51.
Gupta, J.P. (1970) Description of a new species of Phorticella Zaprionus (Drosophilidae) from India. Proceedings of the Indian National Science Academy, 36, 62–70.
Hackman, W. (1955) On the genera Scaptomyza Hardy and Parascaptomyza Duda (Dipt. Drosophilidae). Notulae Entomologicae, 35, 74–91.
Hackman, W. (1959) On the genus Scaptomyza Hardy (Dipt., Drosophilidae) with descriptions of new species from various parts of the world. Acta Zoologica Fennica, 97, 3–73.
Hackman, W. (1962) On Hawaiian Scaptomyza species (Dipt., Drosophilidae). Notulae Entomologicae, 42, 33–42.
Hardy, D.E. (1952) Additions and corrections to Bryan’s check list of the Hawaiian Diptera. Proceedings of the Hawaiian Entomological Society, 14, 443–484.
Hardy, D.E. (1956) Dettopsomyia nigrovittata (Malloch). Proceedings of the Hawaiian Entomological Society, 16 (1), 17.
Hardy, D.E. (1965) Diptera: Cyclorrhapha II, Series Schizophora, Section Acalypteratae I. Family Drosophilidae. Insects of Hawaii. University of Hawaii Press, Honolulu, 814 pp.
Hardy, D.E. (1966) VIII. Descriptions and notes on Hawaiian Drosophilidae (Diptera). The University of Texas Publications, 6615, 195–244.
Hardy, D.E. (1969) Notes on Hawaiian “Idiomyia” (Drosophila). The University of Texas Publication, 6918, 71–77.
Hardy, D.E. (1977) Review of Hawaiian Drosophila (Antopocerus) Hardy. Proceedings of the Entomological Society of Washington, 79, 82–95. https://doi.org/10.2307/1850070
Hardy, D.E. (1978a) A New Symmorphic Sibling Species of Drosophila (Diptera) from the Island of Maui, Hawaii. American Midland Naturalist, 99, 350–351. https://doi.org/10.2307/2424811
Hardy, D.E. (1978b) Replacement name for Drosophila nigricolor Hardy (Diptera: Drosophilidae). Proceedings of the Entomological Society of Washington, 80, 102.
Hardy, D.E., Delfinado, M.D. & Fujii, D. (1981) Litter-inhabiting Diptera. In: Mueller-Dombois, D., Bridges, K.W. & Carson, H.L. (Eds.), Island ecosystems: biological organization in selected Hawaiian communities. Hutchinson Ross, Stroudsburg, pp. 147–156.
Hardy, D.E. & Kaneshiro, K.Y. (1968) New Picture-Winged Drosophila from Hawaii. The University of Texas Publication, 6818, 171–262.
Hardy, D.E. & Kaneshiro, K.Y. (1969) Descriptions of New Hawaiian Drosophila. The University of Texas Publication, 6918, 39–54.
Hardy, D.E. & Kaneshiro, K.Y. (1971) New Picture-Winged Drosophila From Hawaii, Part II. (Drosophilidae, Diptera). The University of Texas Publication, 7103, 151–170.
Hardy, D.E. & Kaneshiro, K.Y. (1972) New Picture-Winged Drosophila From Hawaii, Part III. (Drosophilidae, Diptera). The University of Texas Publication, 7213, 155–161.
Hardy, D.E. & Kaneshiro, K.Y. (1975a) Studies in Hawaiian Drosophila, Miscellaneous New Species, no. I: Mitchelli Subgroup. Proceedings of the Hawaiian Entomological Society, 22, 57–64.
Hardy, D.E. & Kaneshiro, K.Y. (1975b) Studies on Hawaiian Drosophila, modified-mouthparts species. Proceedings of the Hawaiian Entomological Society, 22, 51–55.
Hardy, D.E. & Kaneshiro, K.Y. (1979) A review of the modified tarsus species group of Hawaiian Drosophila (Drosophilidae; Diptera). Proceedings of the Hawaiian Entomological Society, 23, 71–90.
Hardy, D.E., Kaneshiro, K.Y., Val, F.C. & O’Grady, P.M. (2001) Review of the haleakalae species group of Hawaiian Drosophila (Diptera: Drosophilidae). Bishop Museum Bulletin in Entomology, 9, 1–88.
Hardy, J. (1849) Note on remedies for the turnip-fly amongst the ancients, and on the turnip-fly of New Holland, with notice of a new genus and species of Diptera. History of the Berwickshire Naturalist’s Club, 2, 359–362.
Harrison, R.A. (1959) Acalypterate Diptera. N.Z. Department of Scientific and Industrial Research Bulletin, 128, 1–381.
Heed, W.B. (1968) Ecology of the Hawaiian Drosophila. The University of Texas Publications, 6818, 387–419.
Heeger, E. (1852) Beitrage zur Naturgeschichte der Insecten. Sitzungsberichte der Kaiserliche Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse, 9, 774–781.
Hellqvist, S., Ericson, N. & Struwe, I. (2017) Intressanta fynd av daggflugor, med en uppdaterad provinskatalog över de svenska arterna (Diptera: Drosophilidae). Entomologisk Tidskrift, 138, 189–202.
Hunt, J.A. & Carson, H.L. (1983) Evolutionary relationships of four species of Hawaiian Drosophila as measured by DNA reassociation. Genetics, 104, 353–364. https://doi.org/10.1093/genetics/104.2.353
Hutton, F.W. (1901) Synopsis of the Diptera brachycera of New Zealand. Transactions and Proceedings of the New Zealand Institute, 33, 1–95.
Johnson, C.W. (1895) Diptera of Florida, with Additional Descriptions of New Genera and Species, by D. W. Coquillett. Proceedings of the Academy of Natural Sciences of Philadelphia, 47, 303–340.
Johnson, C.W. (1913) The distribution of some species of Drosophila. Psyche (Journal of entomology), 20, 202–204. https://doi.org/10.1155/1913/41505
Kahanpää, J. (2014) Checklist of the Braulidae, Camillidae, Diastatidae and Drosophilidae of Finland (Diptera, Ephydroidea). ZooKeys, 338, 333–338. https://doi.org/10.3897/zookeys.441.7057
Kambysellis, M.P. (1993) Ultrastructural diversity in the egg chorion of Hawaiian Drosophila and Scaptomyza: ecological and phylogenetic considerations. International Journal of Insect Morphology & Embryology, 22, 417–446. https://doi.org/10.1016/0020-7322(93)90022-S
Kaneko, A. & Shima, T. (1962) Drosophila Survey of Hokkaido, XVIII.: Drosophilidae From Four Localities of Hokkaido (With 3 Text-figures and 1 Table). 北海道大學理學部紀要, 15, 74–79.
Kaneshiro, K.Y. (1969) The Drosophila crassifemur group of species in a new subgenus. University of Texas Publications, 6918, 79–83.
Kaneshiro, K.Y. (1976) A revision of the generic concepts in the biosystematics of Hawaiian Drosophilidae. Proceedings of the Hawaiian Entomological Society, 22, 255–278.
Kaneshiro, K.Y. (1983) Drosophila (Sophophora) suzukii (Matsumura). Proceedings of the Hawaiian Entomological Society, 24, 179.
Kaneshiro, K.Y., Carson, H.L., Clayton, F.E. & Heed, W.B. (1973) Niche separation in a pair of homosequential Drosophila species from the island of Hawaii. American Naturalist, 107, 766–774. https://doi.org/10.1086/282874
Kaneshiro, K.Y. & Kambysellis, M.P. (1999) Description of a New Allopatric Sibling Species of Hawaiian Picture-Winged Drosophila. Pacific Science, 53, 208–213.
Kang, Y.S., Lee, T.J. & Bahng, K.W. (1965) Three New Species and Two Unrecorded Species of Drosophilidae from Korea. The Korean Journal of Zoology, 8, 51–54.
Karpa, A. (2008) Catalogue of Latvian Flies (Diptera: Brachycera). Latvijas entomologs, 46, 4–43.
Katoh, T., Izumitani, H.F., Yamashita, S. & Watada, M. (2017) Multiple origins of Hawaiian drosophilids: Phylogeography of Scaptomyza hardy (Diptera: Drosophilidae). Entomological Science, 20, 33–44. https://doi.org/10.1111/ens.12222
Kikkawa, H. & Peng, F.T. (1938) Drosophila Species of Japan and Adjacent Localities. Japanese Journal of Zoology [Nihon dobutsugaku shuho], 7, 507–552.
Knab, F. (1914) Drosophilidae with parasitic larvae. Insecutor Inscitiae Menstruus, 2, 165–169.
Kozhevnikov, B.T. (1936) Experimentally produced karyotypical isolation. (in Russian, with English summary). Biologicheskii Zhurnal [Journal de Biologie; Zeitschrift fur Biologie], 5, 727–752.
Kumar, A. & Gupta, J.P. (1992) Two new and two unrecorded species of Drosophilidae from Sikkim, India (Insecta: Diptera). Senckenbergiana Biologica, 72, 45–51.
Lamb, C.B. (1914) Reports of the Percy Sladen Trust Expedition. No. XV. Diptera: Drosophilidae. The Transactions of the Linnean Society of London, 16, 325–351. https://doi.org/10.1111/j.1096-3642.1913.tb00152.x
Lapoint, R.T., Gidaya, A. & O’Grady, P.M. (2011) Phylogenetic relationships in the spoon tarsus subgroup of Hawaiian Drosophila: Conflict and concordance between gene trees. Molecular Phylogenetics and Evolution, 58 (2011), 492–501. https://doi.org/10.1016/j.ympev.2010.12.015
Lapoint, R.T., Magnacca, K.N. & O’Grady, P.M. (2009) Review of the spoon tarsus subgroup of Hawaiian Drosophila (Drosophilidae: Diptera), with a description of one new species. Zootaxa, 2003 (1), 53–68. https://doi.org/10.11646/zootaxa.2003.1.3
Lapoint, R.T., Magnacca, K.N. & O’Grady, P.M. (2014) Phylogenetics of the antopocerus-modified tarsus clade of Hawaiian Drosophila: Diversification across the Hawaiian Islands. PLoS ONE, 9 (11), e113227. https://doi.org/10.1371/journal.pone.0113227
Lapoint, R.T., O’Grady, P.M. & Whiteman, N.K. (2013) Diversification and dispersal of the Hawaiian Drosophilidae: the evolution of Scaptomyza. Molecular Phylogenetics and Evolution, 69, 95–108. https://doi.org/10.1016/j.ympev.2013.04.032
Leblanc, L., O’Grady, P.M., Rubinoff, D. & Montgomery, S.L. (2009) New Immigrant Drosophilidae in Hawaii, and a Checklist of the Established Immigrant Species. Proceedings of the Hawaiian Entomological Society, 41, 121–127.
Leblanc, L., Rubinoff, D. & Wright, M.G. (2013) Conservation Implications of Changes in Endemic Hawaiian Drosophilidae Diversity across Land Use Gradients. PLoS ONE, 8, 1–15. https://doi.org/10.1371/journal.pone.0062464
Lin, F.J. & Ting, J.I. (1971) Several additions to the fauna of Taiwan Drosophilidae (Diptera). Bulletin of the Institute of Zoology,Academica Sinica, Taiwan, 10, 17–35.
Lin, F.J. & Tseng, H.C. (1971) A new species of polychaeta species group of subgenus Drosophila from Taiwan (Drosophila, Drosophilidae, Diptera). Bulletin of the Institute of Zoology, Academia Sinica [Tung wu yen chiu so chi k’an], 10, 69–72.
Lin, F.J., Tseng, H.C. & Lee, W.Y. (1977) A catalogue of the family Drosophilidae in Taiwan (Diptera). Quarterly Journal of the Taiwan Museum, 30, 345–372.
Loew, H. (1862) Diptera Americae septentrionalis indigena. Centuria secunda. Berliner entomologische Zeitschrift, 6, 185–232.
Loew, H. (1863) Diptera Americae Septentrionalis Indigena. Centuria Tertia. Berliner Entomologische Zeitschrift, 7, 1–55. https://doi.org/10.1002/mmnd.18630070104
Macquart, J. (1843) Dipteres exotiques nouveaux ou peu connus. Tome 2, Part 3. Memoires de la Societe [Royale] des Sciences, de l’Agriculture et des Arts de Lille, 1842, 162–460.
Magnacca, K.N. (2015) Notes on native and alien Hymenoptera and Diptera (Insecta) from the Hawaiian Islands. Bishop Museum Occasional Papers, 116, 19–22.
Magnacca, K.N., Foote, D. & O’Grady, P.M. (2008) A review of the endemic Hawaiian Drosophilidae and their host plants. Zootaxa, 1728 (1), 1–58. https://doi.org/10.11646/zootaxa.1728.1.1
Magnacca, K.N. & O’Grady, P.M. (2006) Subgroup structure for the modified mouthparts species group of Hawaiian Drosophila. Proceedings of the Hawaiian Entomological Society, 38, 87–101.
Magnacca, K.N. & O’Grady, P.M. (2008a) New combinations in Hawaiian Drosophila and Scaptomyza (Diptera: Drosophilidae). Zootaxa, 1926 (1), 53–60. https://doi.org/10.11646/zootaxa.1926.1.5
Magnacca, K.N. & O’Grady, P.M. (2008b) Revision of the “nudidrosophila” and “ateledrosophila” species groups of Hawaiian Drosophila (Diptera: Drosophilidae), with descriptions of twenty-two new species. Systematic Entomology, 33 (2), 395–428. https://doi.org/10.1111/j.1365-3113.2007.00400.x
Magnacca, K.N. & O’Grady, P.M. (2009) Revision of the Modified Mouthparts Species Group of Hawaiian Drosophila (Diptera: Drosophilidae): The Ceratostoma, Freycinetiae, Semifuscata, and Setiger Subgroups, and Unplaced Species. UC Publication in Entomology, 130, 1–94.
Magnacca, K.N. & Price, D.K. (2012) New species of Hawaiian picture wing Drosophila (Diptera: Drosophilidae) with a key to species. Zootaxa, 3188 (1), 1–30. https://doi.org/10.11646/zootaxa.3188.1.1
Magnacca, K.N. & Price, D.K. (2015) Rapid adaptive radiation and host plant conservation in the Hawaiian picture wing Drosophila (Diptera: Drosophilidae). Molecular Phylogenetics and Evolution, 92, 226–242. https://doi.org/10.1016/j.ympev.2015.06.014
Mainland, G.B. (1949) New Drosophilids. Proceedings of the Hawaiian Entomological Society, 13, 326–327.
Malloch, J.R. (1923) Notes on Australian Diptera with Desscriptions. Proceedings of the Linnean Society of New South Wales, 48, 601–622.
Malloch, J.R. (1924a) Descriptions of Neotropical two-winged flies of the family Drosophilidae. Proceedings of the United States National Museum, 66, 1–11. https://doi.org/10.5479/si.00963801.2540
Malloch, J.R. (1924b) Notes on Australian Diptera. No. IV. Proceedings of the Linnean Society of New South Wales, 49, 348–359.
Malloch, J.R. (1932) New species and other records of Otitidae (Ortalidae), Piophilidae, Clusiidae, Chloropidae, and Drosophilidae from the Marquesas. Bulletin of the Bernice P. Bishop Museum, 98, 205–223.
Malloch, J.R. (1933) Some acalyptrate Diptera from the Marquesas Islands. Bernice P. Bishop Museum Bulletin, 114, 3–31.
Malloch, J.R. (1934a) Additional New Species and Other Records of Acalyptrate Diptera (Sapromyzidae, Asteiidae, Drosophilidae, Ephydridae and Trypetidae) From the Marquesas Islands. Bulletin of the Bernice P. Bishop Museum, 114, 179–200.
Malloch, J.R. (1934b) Diptera of Patagonia and South Chile. Part VI. British Museum (Natural History), 5, 435–455.
Malloch, J.R. (1934c) Diptera, Drosophilidae. In: Insects of Samoa and other Samoan terrestrial arthropoda. Part VI. British Museum Natural History, London, pp. 267–328.
Malloch, J.R. (1938) Two Genera of Hawaiian Drosophilidae (Diptera). Proceedings of the Hawaiian Entomological Society, 10, 4–6.
Mangan, R.L. (1978) Competitive interactions among host plant specific Drosophila species. Ph.D. Thesis. University of Arizona, Tucson, Arizona, 204 pp.
Markow, T.A. & O’Grady, P. (2006) Drosophila: a Guide to Species Identification and Use. Academic Press (Elsevier), London, 272 pp. https://doi.org/10.1016/B978-012473052-6/50003-2
Matsumura, S. (1931) 6000 illustrated insects of Japan-empire (in Japanese). Toko Shoin, Tokyo, 1694 pp.
McEvey, S.F. (1990) New Species of Scaptomyza From Madagascar and Mauritius (Diptera : Drosophilidae). Annales de la Societe Entomologique de France, 26, 51–64.
Meigen, J.W. (1830) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Sechster Theil. Schulz, Hamm, 401 pp.
Meigen, J.W. (1838) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Siebenter Theil oder Supplementband. Schulz, Hamm, xii + 434 + [1] pp., 8 pls.
De Meijere, J.C.H. (1911) Studien uber sudostasiatische Dipteren. VI. Plates 18-22. Tijdschrift voor entomologie, 54, 258–432.
De Meijere, J.C.H. (1914) Studien über südostasiatische Dipteren IX. Tijdschrift voor Entomologie, 57, 137–275.
De Meijere, J.C.H. (1946) Zevende Supplement op de Nieuwe Naamlijst van Nederlandsche Diptera van 1898. (Eerste Supplement op mijne Naamlijst van 1939). Tijdschrift voor entomologie, 87, 1–25.
Mik, J. (1886) Eine neue Drosophila aus Nieder-Oesterreich und den Aschanti-Landern. (Ein dipterologischer Beitrag). Wiener entomologische Zeitung, 5, 328–331. https://doi.org/10.5962/bhl.part.20609
Momma, E. (1968) Domestic Drosophila in Hawaii. Drosophila Information Service, 43, 168.
Montgomery, S.L. (1975) Comparative Breeding Site Ecology and the Adaptive Radiation of Picture-Winged Drosophila (Diptera: Drosophilidae) in Hawaii. Proceedings of the Hawaiian Entomological Society, 22, 65–103.
Nartshuk, E.P. (2014) Fruit flies (Diptera: Drosophilidae) of the Russian Arctic. Zoosystematica Rossica, 23, 256–263. https://doi.org/10.31610/zsr/2014.23.2.256
Nishida, G.M. (2002) Hawaiian terrestrial arthropod checklist. 4th Edition. Hawaii Biological Survey, Bishop Museum Press, Honolulu, 313 pp.
Nishiharu, S. (1979) Three New Species of Drosophilidae from Japan (Diptera). Kontyû, Tokyo, 47, 38–43.
O’Grady, P.M., Beardsley, J.W. & Perreira, W.D. (2002) New records for introduced Drosophilidae (Diptera) in Hawai’i. Bishop Museum Occasional Papers, 69, 34–35.
O’Grady, P.M., Bonacum, J., DeSalle, R. & Do Val, F.C. (2003a) The placement of Engiscaptomyza, Grimshawomyia, and Titanochaeta, three clades of endemic Hawaiian Drosophilidae (Diptera). Zootaxa, 159 (1), 1–16. https://doi.org/10.11646/zootaxa.159.1.1
O’Grady, P.M. & DeSalle, R. (2008) Out of Hawaii: the origin and biogeography of the genus Scaptomyza (Diptera: Drosophilidae). Biology Letters, 4, 195–199. https://doi.org/10.1098/rsbl.2007.0575
O’Grady, P.M. & DeSalle, R. (2018) Phylogeny of the genus Drosophila. Genetics, 209, 1–25. https://doi.org/10.1534/genetics.117.300583
O’Grady, P.M., Kam, M.W.Y., Val, F.C. & Perreira, W.D. (2003b) Revision of the Drosophila mimica Subgroup, with Descriptions of Ten New Species. Annals of the Entomological Society of America, 96, 12–38. https://doi.org/10.1603/0013-8746(2003)096[0012:ROTDMS]2.0.CO;2
O’Grady, P.M., Lapoint, R.T., Bonacum, J., Lasola, J., Owen, E., Wu, Y. & DeSalle, R. (2011) Phylogenetic and ecological relationships of the Hawaiian Drosophila inferred by mitochondrial DNA analysis. Molecular Phylogenetics and Evolution, 58 (2), 344–256. https://doi.org/10.1016/j.ympev.2010.11.022
O’Grady, P.M., Val, F.C., Hardy, D.E. & Kaneshiro, K.Y. (2001) The rustica species group of Hawaiian Drosophila (Diptera: Drosophilidae). Pan-Pacific Entomologist, 77, 254–260.
O’Grady, P.M. & Zilversmit, M. (2004) Phylogenetic relationships within the Drosophila halekalae group inferred by molecular and morphological characters (Diptera, Drosophilidae). Bishop Museum Bulletin in Entomology, 12, 117–134.
Okada, T. (1956) Systematic study of Drosophilidae and Allied Families of Japan. Gihodo Co. Ltd., Tokyo, 183 pp.
Okada, T. (1964a) Drosophilidae (Diptera) of Southeast Asia collected by the Thai-Japanese Biological Expedition 1961-62. Nature and Life in Southeast Asia, 3, 439–466.
Okada, T. (1964b) New and unrecorded species of Drosophilidae in the Amami Islands, Japan. Kontyu, 32, 105–115.
Okada, T. (1965) Drosophilidae of the Okinawa islands. Kontyû, 33, 327–350.
Okada, T. (1966) Diptera from Nepal: Cryptochaetidae, Diastatidae and Drosophilidae. Bulletin of the British Museum (Natural History), 6, 1–129. https://doi.org/10.5962/p.78200
Okada, T. (1973a) Descriptions of four new species of drosophilidae of the Bonins, with taxometrical analyses of the Scaptomyza species (Diptera). Kontyu, 41, 83–90.
Okada, T. (1973b) Drosophilidae and Diastatidae from Mongolia (Diptera). Annales Historico-Naturales Musei Nationalis Hungarici, 65, 271–279.
Okada, T. (1973c) Four new species of Drosophilidae from Japan (Diptera). Kontyu, 41, 434–439.
Okada, T. (1974) Drosophilidae (Diptera) from Korea. Annales historico-naturales Musei nationalis hungarici, 66, 269–275.
Okada, T. (1988) Family Drosophilidae (Diptera) from the Lund University Ceylon expedition in 1962 and Borneo collections in 1978-1979. Entomologica scandinavica. Supplementum 30, 111–151.
Oldenberg, L. (1914) Beitrag zur Kenntnis der europaischen Drosophiliden (Dipt.). Archiv fur Naturgeschichte, 80 (A), 1–42.
Patterson, J.T. & Wheeler, M.R. (1942) Description of new species of the subgenera Hirtodrosophila and Drosophila. The University of Texas Publications, 4213, 67–109.
Peñafiel, A.D. & Rafael, V. (2018) Dos especies nuevas del género Drosophila y el registro de Parascaptomyza clavifera en la provincia del Carchi, Ecuador. Revista Ecuatoriana de Medicina y Ciencias Biológicas, 39 (1), 51–61. https://doi.org/10.26807/remcb.v39i1.571
Perkins, R.C.L. (1910) Supplement (to Diptera). Fauna Hawaiiensis, 2, 697–700.
Perreira, W.D. & Kaneshiro, K.Y. (1990) Three New Species of Picture-Winged Drosophila from the Hawaiian Islands. Proceedings of the Hawaiian Entomological Society, 30, 79–84.
Pipkin, S.B. (1964) New flower breeding species of Drosophila (Diptera: Drosophilidae). Proceedings of the Entomological Society of Washington, 66, 217–245.
Plotnikov, I.S., Sidorenko, V.S. & Krivokhatsky, V.A. (2013) Zoogeographic analysis of the Palaearctic fauna of Drosophilid flies (Diptera, Drosophilidae) with description of clusterization software for provincial faunas. Entomological Review, 93, 831–843. https://doi.org/10.1134/S0013873813070051
Rakshit, A., Chatterjee, P. & Chetterjee, R.N. (2012) Scaptomyza jadavpuri sp. Nov. is a new member of picture wing Drosophilidae of the Scaptomyza complex. Drosophila Information Service, 95, 150–153.
Ray-Chaudhuri, S.P. & Mukherjee, D.P. (1941) Genetic and systematic studies on Indian Drosophila. I. Description of two new species of Drosophila: life-history and preliminary studies on the genetic constitution of Drosophila emulata, sp. nov. Indian Journal of Entomology, 3, 215–224.
Rockwood, E.S., Kanapi, C.G., Wheeler, M.R. & Stone, W.S. (1971) Allozyme changes during the evolution of Hawaiian Drosophila. The University of Texas Publication, 7103, 193–212.
Rondani, C. (1875) Notizie sul moscherino dell’uva. Bollettino del Comizio Agrario Parmense, 8, 145–149.
von Roser, C. (1840) Erster Nachtrag zu dem im Jahre 1834 bekannt gemachten Verzeichnisse in Wurttemberg vorkommender zweiflugliger Insekten. Correspondenzblatt des Koniglich Wurttembergishen Landwirthschaftlichen Vereins, 1840, 49–64.
Sarswat, M., Dewan, S. & Fartyal, R.S. (2016) Mitochondrial DNA sequence variation in Drosophilid species (Diptera: Drosophilidae) along altitudinal gradient from Central Himalayan region of India. Journal of Genetics, 95, 357–367. https://doi.org/10.1007/s12041-016-0646-x
Sati, P.C., Fartyal, R.S., Sarswat, M., Dewan, S., Kandpal, M., Kanita, Patel, M.K. & Pradhan, S. (2013) Check-list of drosophilid species so far described and recorded from Uttarkhand state, India. Drosophila Information Service, 96, 79–83.
Scopoli, J.A. (1763) Entomologia carniolica exhibens insecta carnioliae indigena et distributa in ordines, genera, species, varietates. Methodo linnaeana. Trattner, Vienna, 424 pp. https://doi.org/10.5962/bhl.title.119976
Séguy, E. (1934) Étude sur quelques muscides de L’amerique Latine. Revista de la Sociedade Entomologica de Argentina, 6, 9–16.
Séguy, E. (1936) Voyage de MM. L. Chopard et A. Mequignon aux Acores (aout-septembre 1930). x. Dipteres. Annales de la Societe Entomologique de France, 105, 11–26.
Séguy, E. (1938) Mission scientifique de l ‘Omo. Diptera. I. Nematocera et Brachycera. Mémoires du Muséum national d’histoire naturelle, Paris, 8, 319–380.
Sidorenko, V.S. (1993a) On two species of Drosophilidae (Diptera) described by O. DUDA from Russian Far East. Metteilungen der Schweizerischen Entomologischen Gesellschaft, 66, 75–78.
Sidorenko, V.S. (1993b) Tribe Drosophilini of the Asian part of the USSR. Zeitschrift fur Entomologie, 14, 253–268.
Sidorenko, V.S. (1995) New data on Asian drosophilid flies (Diptera, Drosophilidae). Part 2. Far Eastern Entomologist, 21, 1–4.
Sidorenko, V.S. (1998) New data on Asian Drosophilid flies (Diptera, Drosophilidae). Part 3. Far Eastern Entomologist, 56, 1–8.
Singh, B.K. & Bhati, M. (1988) A Preliminary Report on the Drosophilidae of Kumaun Region with Description of Two New Species and Three New Records. Oriental Insects, 22, 147–161. https://doi.org/10.1080/00305316.1988.11835486
Singh, B.K. & Dash, S. (1993) Drosophilidae of Uttarakhand region with the description of one new species (Insecta: Diptera). Proceedings of the Zoological Society, Calcutta, 46, 131–140.
Smith, C.A. (1958) Drosophilidae Found in Tomato Fields in Indiana. Proceedings of the Indiana Academy of Science, 67, 138–144.
Spencer, W.P. (1940) Subspecies, hybrids and speciation in Drosophila hydei and Drosophila virilis. American Naturalist, 74, 157–179. https://doi.org/10.1086/280882
Spieth, H.T. (1966) Courtship behavior of endemic Hawaiian Drosophila. University of Texas Publications, 6615, 245–313.
Spieth, H.T. (1982) Behavioral Biology and Evolution of the Hawaiian Picture-Winged Species Group of Drosophila. Evolutionary Biology, 14, 351–438.
Spieth, H.T. (1984) Courtship behaviors of the Hawaiian picture-winged Drosophila. University of California Publications. Entomology, 103, 1–92.
Stark, J.B. & O’Grady, P.M. (2010) Morphological variation in the forelegs of the Hawaiian Drosophilidae. I. The AMC clade. Journal of Morphology, 271, 86–103. https://doi.org/10.1002/jmor.10783
Sturtevant, A.H. (1916) Notes on North American Drosophilidae with descriptions of twenty-three new species. Annals of the Entomological Society of America, 9, 323–343. https://doi.org/10.1093/aesa/9.4.323
Sturtevant, A.H. (1919) A new species closely resembling Drosophila melanogaster. Psyche, 26, 153–155. https://doi.org/10.1155/1919/97402
Sturtevant, A.H. (1921) The North American species of Drosophila. Carnegie Institution of Washington Publication, 301, 1–150. https://doi.org/10.5962/bhl.title.33564
Sturtevant, A.H. (1942) The classification of the genus Drosophila, with descriptions of nine new species. The University of Texas Publication, 4213, 5–51.
Swezey, O.H. (1934) Records of immigrant insects for 1933. Proceedings of the Hawaiian Entomological Society, 8, 544.
Swezey, O.H. (1952) Note: Insects from decaying blossoms. Proceedings of the Hawaiian Entomological Society, 14, 357.
Swezey, O.H. & Davis, C.J. (1947) New insect records. Proceedings of the Hawaiian Entomological Society, 13, 14.
Takada, H. (1958) Drosophila Survey of Hokkaido, X.: Drosophilidae from several localities of Hokkaido (With 3 Text-figures and 5 Tables). 北海道大學理學部紀要, 14, 120–127.
Takada, H. (1959) Drosophila survey of Hokkaido IX. On Drosophila okadai sp. nov. with supplementary notes on the female of Scaptomyza polygonia Okada. Annotationes Zoologicae Japoneses, 32, 152–155.
Takada, H. (1966) Male Genitalia of some Hawaiian Drosophilidae. University of Texas Publications, 6615, 315–333.
Takada, H. (1970) Scaptomyza (Parascaptomyza) pallida (Zetterstedt) and two related new species, S. (P.) elmoi n. sp. and S. (P.) himalayana n. sp. (Diptera: Drosophilidae). Annotationes Zoologicae Japoneses, 43, 142–147.
Takada, H., Beppu, K. & Toda, M.J. (1979) Drosophila survey of Hokkaido, XXXVI. New and unrecorded species of Drosophilidae (Diptera). Journal of the Faculty of General Education, Sapporo University, 14, 105–129.
Takada, H. & Momma, E. (1975) Distribution and Population Constitution of Drosophila in South East Asia and Oceania: II. Drosophilidae in the Suburbs of Kuala Lumpur, West Malaysia. Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology, 20, 9–48.
Takada, H. & Yoon, J.S. (1989) Three new Drosophila species (Diptera: Drosophilidae) from British Columbia, Hawaii, and the Canary Islands. Entomological News, 100, 111–121.
Thomas, R.H. & Hunt, J.A. (1991) The molecular evolution of the alcohol dehydrogenase locus and the phylogeny of Hawaiian Drosophila. Molecular Biology and Evolution, 8, 687–702. https://doi.org/10.1093/oxfordjournals.molbev.a040678
Thomson, C. (1869) Diptera species novas descripsit. In: Kongliga svenska fregatten Eugenies resa omkring jorden under befal of C. A. Virgen, Aren 1851–1853. 2 (Zoologi) 1, Insecta P. A. Norstedt & Soner, Stockholm, pp. 443–614.
Throckmorton, L.H. (1966) The relationships of the endemic Hawaiian Drosophilidae. University of Texas Publications, 6615, 335–396.
Toda, M., Riihimaa, A. & Fuyama, Y. (1987) Additional notes on drosophilid flies (Diptera, Drosophilidae) in the Bonin Islands, with descriptions of two new species. Kontyû, Tokyo, 55, 240–258.
Toda, M.J. (1979) A preliminary note on winter Drosophilid flies in southern Japan, with special reference to reproductive conditions. Low temperature science, Series B, Biological sciences, 37, 39–45.
Toda, M.J. (2018) DrosWLD-Species: taxonomic information database for world species of Drosophilidae. Available from: https://bioinfo.museum.hokudai.ac.jp/db/modules/st (accessed 5 December 2021)
Toda, M.J. & Kwon, O.K. (1983) Collection records of Drosophilid flies from the Quelpart Island, Korea. Drosophila Information Service, 59, 130–132.
Toda, M.J., Sidorenko, V.S., Watabe, H.A., Kholin, S.K. & Vinokurov, N.N. (1996) A Revision of the Drosophilidae (Diptera) in East Siberia and Russian Far East: Taxonomy and Biogeography. Zoological Science, 13, 455–477.
Tsacas, L. (1972) The “Genus” Euscaptomyza Seguy (Diptera: Drosophilidae), with the Description of Two New African Species. University of Texas Publications, 7213, 345–354.
Tsacas, L. (1980) Family Drosophilidae. In: Crosskey, R.W. (Ed.), Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, pp. 673–685.
Tsacas, L. (1990) Drosophilidae de l’Afrique australe (Diptera). Annals of the Natal Museum, 31, 103–161.
Tsacas, L. & Cogan, B.H. (1976) La faune terrestre de l’île de Sainte-Hélène. Troisième partie. II. Insectes, 13. Diptera, 19. Fam. Drosophilidae. Annales de la Muséum Royald’ Afrique Centrale, 215, 82–95.
Villeneuve, J. (1911) Description de deux nouveaux Dipteres. Wiener entomologische Zeitung, 30, 81–84.
Villeneuve, J. (1923) Description d’une nouvelle espece du genre Drosophila Fallen (Dipt. Muscidae). Bulletin de la Societe Entomologique de France, 1923, 28. https://doi.org/10.3406/bsef.1923.27103
Walker, F. (1849) Printed by Order of the Trustees. List of the specimens of dipterous insects in the collection of the British Museum, 1849 (Part IV), 689–1172.
Walker, F. (1853) Insecta Britannica: Diptera. Vol. II. Reeve and Benham, London, 370 pp.
Walker, F. (1859) Catalogue of the Dipterous Insects collected in the Aru Islands by Mr. A. R. Wallace, with Descriptions of New Species. Journal of the Proceedings of the Linnean Society, 3, 111–131. https://doi.org/10.1111/j.1096-3642.1859.tb00076.x
Waltl, J. (1837) Neue Gattungen von Mucken bey Passau. Isis, 4, 279–283.
Watabe, H., Toda, M.J. & Vinokurov, N.N. (1994) A preliminary report on the drosophilid fauna of Sakha SSR, the East Siberia. Drosophila Information Service, 75, 145–147.
Watabe, H.A., Toda, M.J., Li, G., Duan, C., Imitty, R., Entomack, B. & Muhtar, A. (1993) Drosophilid fauna (Diptera, Drosophilidae) of Chinese Central Asia. Japanese Journal of Entomology, 61, 525–545.
Wheeler, M.R. (1949) Taxonomic studies on the Drosophilidae. University of Texas Publications, 4920, 157–206.
Wheeler, M.R. (1952) The Drosophilidae of the Nearctic Region, exclusive of the genus Drosophila. University of Texas Publications, 5204, 162–218.
Wheeler, M.R. (1987) Drosophilidae. In: McAlpine, J.F. (Ed.), Manual of Nearctic Diptera. Research Branch Agriculture Canada, Ottawa, pp. 1011–1018.
Wheeler, M.R. & Kaneshiro, K.Y. (1974) Notes and exhibitions. Proceedings of the Hawaiian Entomological Society, 21, 319–320.
Wheeler, M.R. & Takada, H. (1966) The Nearctic and Neotropical species of Scaptomyza Hardy (Diptera: Drosophilidae). University of Texas Publications, 6615, 37–78.
Willbrand, B., Pfeiffer, D., Leblanc, L. & Yassin, A. (2018) First report of African fig fly, Zaprionus indianus Gupta (Diptera: Drosophilidae), on the island of Maui, Hawaii, USA, in 2017 and potential impacts to the Hawaiian entomofauna. Proceedings of the Hawaiian Entomological Society, 50, 55–65.
Williston, S.W. (1896) On the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London, 1896, 253–446. https://doi.org/10.1111/j.1365-2311.1896.tb00965.x
Wirth, W.W. (1952) Two new spider egg predators from the Hawaiian Islands (Diptera: Drosophilidae). Proceedings of the Hawaiian Entomological Society, 14, 415–417.
Wollaston, T.V. (1858) Brief Diagnostic Characters of undescribed Madeiran Insects (concluded). Annals and Magazine of Natural History, 1 (3), 113–125. https://doi.org/10.1080/00222935808696882
van der Wulp, F.M. (1892) Eenige uitlandsche Diptera. Tijdschrift voor entomologie, 34, 193–218.
van der Wulp, F.M. (1897) Zur Dipteren-Fauna von Ceylon. Termeszettudomanyi Fuzetek, 20, 136–144.
Yoon, J.S., Resch, K., Wheeler, M.R. & Richardson, R.H. (1975) Evolution in Hawaiian Drosophilidae: Chromosomal phylogeny of the Drosophila crassifemur complex. Evolution, 29, 249–256. https://doi.org/10.2307/2407213
Zetterstedt, J.W. (1838) Dipterologis Scandinaviae. Sectio tertia. Diptera. In: Insecta Lapponica. L. Voss, Lipsiae [Leipzig], pp. 477–868
Zetterstedt, J.W. (1847) Diptera Scandinaviae disposita et descripta. Tomus sextus. Officina Lundbergiana, Lundae, pp. 2163–2580.
Zhang, W., Chen, H., Peng, T. & Lin, F. (1996) Drosophilidae. In: Xue, W. & Chao, C. (Eds.), Flies of China. Liaoning Science and Technology Press, Shenyang, pp. 280–414.
Zimmerman, E.C. (1938) Two insect immigrants new to the Hawaiian fauna. Proceedings of the Hawaiian Entomological Society, 10, 131–132.