Skip to main content Skip to main navigation menu Skip to site footer
Type: Monograph
Published: 2022-03-09
Page range: 1-211
Abstract views: 634
PDF downloaded: 48

The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “Third Pole” (Coleoptera, Staphylinidae, Pselaphinae)

Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China.
Coleoptera beetles diversity Staphylinidae Pselaphinae Batrisitae taxonomy new genus new species new synonym identification key species distribution map Qinghai-Tibet Plateau China

Abstract

The taxonomic knowledge of the pselaphine tribe Batrisini (Coleoptera: Staphylinidae) from the Tibet Autonomous Region is summarized, with 80 species placed in 13 genera. Two genera and 68 species are described as new, aided with illustrations of habitus and diagnostic characters: Arthromelodes alpitorus sp. nov., A. angulatus sp. nov., A. aniqiao sp. nov., A. bicolor sp. nov., A. complexus sp. nov., A. cona sp. nov., A. crassicornis sp. nov., A. criniger sp. nov., A. cylindricus sp. nov., A. flosculus sp. nov., A. gyamda sp. nov., A. intricatus sp. nov., A. lage sp. nov., A. langjicuo sp. nov., A. latithorax sp. nov., A. lebus sp. nov., A. lotus sp. nov., A. markam sp. nov., A. monba sp. nov., A. nepaeformis sp. nov., A. nomurai sp. nov., A. songxiaobini sp. nov., A. speciosus sp. nov., A. torus sp. nov., A. zhangmu sp. nov., A. zhentangensis sp. nov., Batriscenaulax campestris sp. nov., Batrisiella acclivis sp. nov., Batrisiella beibeng sp. nov., Batrisiella bomea sp. nov., Batrisiella concisa sp. nov., Batrisiella curvitibialis sp. nov., Batrisiella ganma sp. nov., Batrisiella guttata sp. nov., Batrisiella zhujianqingi sp. nov., Batrisodes guoguotang sp. nov., Batrisodes hanmi sp. nov., Coryphomimus levigatus gen. et sp. nov., Coryphomodes budda sp. nov., Coryphomodes cephalicus sp. nov., Coryphomodes chenzhilini sp. nov., Gyrongita uniformis gen. et sp. nov., Sathytes aequalis sp. nov., S. asura sp. nov., S. caichenyangi sp. nov., S. dawai sp. nov., S. duojii sp. nov., S. laticornis sp. nov., S. pengzhongi sp. nov., S. proclivis sp. nov., S. pseudograndis sp. nov., S. sympatricus sp. nov., S. virupaksa sp. nov., S. yigongensis sp. nov., Tribasodites bari sp. nov., T. cellulanus sp. nov., T. constrictus sp. nov., T. dilophus sp. nov., T. elongatus sp. nov., T. grandiceps sp. nov., T. gyirong sp. nov., T. kiypu sp. nov., T. mirabilis sp. nov., T. prolixicornis sp. nov., T. pugiunculus sp. nov., T. vertexalis sp. nov., T. yatung sp. nov., and Trisinus quadrilobus sp. nov. The genus Plocamarthrus Jeannel, 1960 is proposed as a subjective junior synonym of Arthromelodes Jeannel 1954, syn. nov., resulting in Arthromelodes championi (Jeannel, 1960) comb. nov. (type species of Plocamarthrus), A. cameroni (Jeannel, 1960) comb. nov., A. planiceps (Jeannel, 1960) comb. nov., and A. abdominalis (Jeannel, 1960) comb. nov. Arthromelodes championi and Sathytes simplex Löbl, 1979 are reported to occur in Tibet, which are also new country records for China. Additional collecting data are provided for Hingstoniella lata Jeannel, 1960, Sathytes panzhaohuii Yin & Shen, 2020, Sathytes transversus Yin & Shen, 2020, and Sathytes xizangensis Yin & Shen, 2020. Keys for identification of the genera and species are presented. Maps showing the distribution of each species are provided, supplemented with habitat photos. Lists of Tibetan species of Batrisini and of other pselaphine groups are given, adding up to 95 species.

 

References

  1. Aubé, C. (1833) Note sur la famille des Psélaphiens. Annales de la Société Entomologique de France, 2, 502–511.
    Chandler, D.S. (1997) A catalog of the Coleoptera of America north of Mexico. Family: Pselaphidae. United States Department of Agriculture, Agriculture Handbook 529–31. United States Department of Agriculture, Washington, D.C., ix + 118 pp.
    Chandler, D.S. (2001) Biology, morphology, and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International, 15, 1–560.
    Besuchet, C.(1981) Contribution à l'étude des Batrisodes paléarctiques (Coleoptera: Pselaphidae). Revue Suisse de Zoologie, 88 (1), 275–296. https://doi.org/10.5962/bhl.part.82372
    Hampe, C. (1863) Ein kleiner Beitrag zur gross-österreichischen Käferfauna. Wiener Entomologische Monatschrift, 7 (8), 285–290.
    Hortal, J., de Bello, F., Diniz-Filho, A.J.F., Lewinsohn, T.M., Lobo, J.M. & Ladle, R.J. (2015) Seven shortfalls that beset large-scale knowledge of biodiversity. Annual Review of Ecology, Evolution, and Systematics, 46, 523–549. https://doi.org/10.1146/annurev-ecolsys-112414-054400
    Huang, M.-C., Li, L.-Z. & Yin, Z.-W. (2018) Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species. Acta Entomologica Musei Nationalis Pragae, 58 (2), 457–478. https://doi.org/10.2478/aemnp-2018-0035
    Huang, M.-C. & Yin, Z.-W. (2019) The Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) of Nepal. Revue suisse de Zoologie, 126 (2), 165–196. https://doi.org/10.5281/zenodo.3463445
    Huang, M.-C. & Yin, Z.-W. (2020) Validation of five Pselaphodes species names in Huang & Yin (2019): The Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) of Nepal. Revue suisse de Zoologie, 127 (1), 241–242. https://doi.org/10.35929/RSZ.0016
    Inoue, S. & Maruyama, M. (2020) A new termitophilous species of Tribasodites Jeannel from Taiwan and southern China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4786 (3), 417–424. https://doi.org/10.11646/zootaxa.4786.3.7
    Jeannel, R. (1952) Psélaphides de Saigon. Revue Française d'Entomologie, 19 (2), 69–113.
    Jeannel, R. (1954a) Psélaphides recueillis par N. Leleup au Congo Belge sous les auspices de l'Institut pour la Recherche Scientifique en Afrique Centrale (I.R.S.A.C.). VI–XI. Annales du Musée Royal du Congo Belge, Tervuren, Série 8º: Sciences Zoologiques, 33, 1–175.
    Jeannel, R. (1954b) Les Psélaphides de Madagascar. Mémoires de l'Institut Scientifique de Madagascar, E: Entomologie, 4, 139–344.
    Jeannel, R. (1957) Sur quelques Psélaphides du Tonkin recueillis par le Père A. de Cooman. Revue Française d'Entomologie, 24 (1), 5–32.
    Jeannel, R. (1958) Révision des Psélaphides du Japon. Mémoires du Muséum National d'Histoire Naturelle, A: Zoologie, 18 (1), 1–138.
    Jeannel, R. (1959) Révision des Psélaphides de l'Afrique intertropicale. Annales du Musée Royal du Congo Belge, Tervuren, Série 8º: Sciences Zoologiques, 75, 1–742.
    Jeannel, R. (1960) Sur les Psélaphides (Coleoptera) de l'Inde septentrionale. Bulletin of the British Museum (Natural History), Entomology, 9, 403–456.
    Jeannel, R. (1961) Sur les Psélaphides de Ceylan. Bulletin of the British Museum (Natural History), Entomology, 10 (10), 423–456.
    Jiang, R.-X. & Yin, Z.-W. (2016) Two new species of Batrisodes Reitter (Coleoptera: Staphylinidae: Pselaphinae) from China. Zootaxa, 4205 (2), 194–200. https://doi.org/10.11646/zootaxa.4205.2.9
    Jiang, R.-X. & Yin, Z.-W. (2017) Eight new species and two new records of Batriscenellus Jeannel (Coleoptera: Staphylinidae: Pselaphinae) from China and India. Zootaxa, 4318 (3), 561–575. https://doi.org/10.11646/zootaxa.4318.3.8
    Jiang, Y., Shi, B., Su, G., Lu, Y., Li, Q., Meng, J., Ding, Y., Song, S. & Dai, L. (2021) Spatiotemporal analysis of ecological vulnerability in the Tibet Autonomous Region based on a pressure-state-response-management framework. Ecological Indicators, 130 (3), 108054. https://doi.org/10.1016/j.ecolind.2021.108054
    Kurbatov, S.A. (2007) Revision of the genus Intestinarius gen. n. from Southeast Asia, with notes on a probable autapomorphy of Batrisitae (Coleoptera: Staphylinidae: Pselaphinae). Russian Entomological Journal, 16 (3), 281–295.
    Leleup, N. (1970) Contributions à l'étude des Coléoptères Psélaphides de l'Afrique. 7. Une nouvelle sous-tribu de Batrisini de l'Afrique intertropicale. Bulletin et Annales de la Société Royale d'Entomologie de Belgique, 105, 305–341. [1969]
    Leleup, N. (1971) Contributions à l'étude des Coléoptères Psélaphides de l'Afrique. 9. Un genre inédit d'Ambicocerina et nouvelle diagnose de la sous-tribu. Bulletin et Annales de la Société Royale d'Entomologie de Belgique, 106, 331–339. [1970]
    Leleup, N. (1973) Contributions à l'étude des Coléoptères Psélaphides de l'Afrique. 13. Description de trois genres inédits et de quatorze espèces nouvelles des régions interpropicales, nouvelle définition des Ambicocerina et création de la tribu des Barrosellini. Bulletin et Annales de la Société Royale Belge d'Entomologie, 109 (1–3), 43–91.
    Li, H. & Song W. (2021) Spatiotemporal Distribution and Influencing Factors of Ecosystem Vulnerability on Qinghai-Tibet Plateau. International Journal of Environmental Research and Public Health, 18 (12), 6508. https://doi.org/10.3390/ijerph18126508
    Li, L.-Z. (Chief Ed.), Hu, J.-Y., Peng, Z., Tang, L., Yin, Z.-W. & Zhao, M.-J. (Eds.) (2019) Catalogue of Chinese Coleoptera. Vol. 3. Staphylinidae. Science Press, Beijing, xix + 720 pp.
    Löbl I. (1979) Révision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285–307. https://doi.org/10.5962/bhl.part.82294
    Löbl, I. & Kodada, J. (2021) On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Hartmann, M., Barclay, M.V.L. & Weipert, J. (Eds.), Biodiversität und Naturausstattung im Himalaya VII. Naturkundemuseum, Erfurt, pp. 349–368. https://doi.org/10.5281/zenodo.5793818
    Löbl, I. & Kurbatov, S.A. (2001) The Batrisini of Sri Lanka (Coleoptera: Staphylinidae: Pselaphinae). Revue Suisse de Zoologie, 108 (3), 559–697. https://doi.org/10.5962/bhl.part.80163
    Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Archiv für Naturgeschichte, 84 (A), i–xxxi + 1–696 pp. [1918]
    Myers, N., Mittermeier, R.A., Mittermeier, C.G., da Fonseca, G.A.B. & Kent, J. (2000) Biodiversity hotspots for conservation priorities. Nature, 403, 853–858. https://doi.org/10.1038/35002501
    Newton, A.F. (2018) Staphyliniformia world catalog database. In: Bánki, O., Roskov, Y., Döring, M., Ower, G., Vandepitte, L., Hobern, D., Remsen, D., Schalk, P., DeWalt, R.E., Keping, M., Miller, J., Orrell, T., Aalbu, R., Adlard, R., Adriaenssens, E., Aedo, C., Aescht, E., Akkari, N. & Alonso-Zarazaga, M.A. (Eds.), Catalogue of Life Checklist. November 2018. Available from: https://www.catalogueoflife.org/data/dataset/1204 (accessed 12 February 2020) https://doi.org/10.48580/d4tn-3gk
    Newton, A.F. Jr. & Chandler, D.S.(1989) World catalog of the genera of Pselaphidae (Coleoptera). Fieldiana, New Series, Zoology, 53, 1–93. https://doi.org/10.5962/bhl.title.3209
    Nomura, S. (1986) Descriptions of two new myrmecophilous species of the family Pselaphidae (Coleoptera) from Japan. Kontyû, 54 (3), 498–504.
    Nomura, S. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1–462. https://doi.org/10.5109/2550
    Nomura, S. (2007) Two new species of the genus Batrisodes (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan. Elytra, 35 (1), 76–84.
    Nomura, S. & Aung, M.M. (2020) Inventory studies on the subfamily Pselaphinae (Coleoptera, Staphylinidae) of Myanmar part 2: A list of collected species in Tanintharyi Region in January 2017. Bulletin of the National Museum of Nature and Science, Series A (Zoology), 46 (4), 203–213.
    Nomura, S. & Idris, A.B. (2003) Faunistic notes on the batrisine species from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga, 8 (1–2), 55–72.
    Nomura, S. & Pham, H.T. (2019) List of pselaphine species (Insecta, Coleoptera, Staphylinidae) collected by light traps from North Vietnam in 2014 with supplements and corrections to the check-list of Nomura (2013). Bulletin of the National Museum of Nature and Science, Series A (Zoology), 45 (2), 73–83.
    Park, O. (1947) Observations on Batrisodes (Coleoptera: Pselaphidae), with particular reference to the American species east of the Rocky Mountains. Bulletin of the Chicago Academy of Sciences , 8 (3), 45–132, pls. 1–11.
    Park, O. (1948) Checklist of the genus Batrisodes (Coleoptera: Pselaphidae). Bulletin of the Chicago Academy of Sciences, 8 (4), 137–169.
    Qiu, J. (2008) The Third Pole. Nature, 454, 393–396. https://doi.org/10.1038/454393a
    QTP Expedition Group, Chinese Academy of Sciences (1981) Insects of Xizang (Tibet). Vols. 1–2. Science Press, Beijing, xi + 600 pp. & ix + 508 pp. [in Chinese, with English summary]
    Raffray, A. (1893) Révision des Pselaphides de Sumatra. Annales de la Société Entomologique de France, 61, 463–504, pl. 10.
    Raffray, A. (1894a) Voyage de M. E. Simon à l'île de Ceylan (janvier–février 1892), 6e Mémoire (1). Psélaphides. Annales de la Société Entomologique de France, 62, 443–462.
    Raffray, A. (1894b) Révision des Psélaphides des Iles de Singapore et de Penang. Revue d'Entomologie, 13, 197–282, pl. 1.
    Raffray, A. (1904) Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France, 72, 484–604.
    Raffray, A. (1909) Nouvelles espèces de Psélaphides. Annales de la Société Entomologique de France, 78, 15–52.
    Reitter, E. (1882) Neue Pselaphiden und Scydmaeniden aus Brasilien. Deutsche Entomologische Zeitschrift, 26 (1), 129–152, pl. 5.
    Schülke, M. & Smetana, A. (2015) Staphylinidae Latreille, 1802. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Revised and Updated Edition, Hydrophiloidea—Staphylinoidea. 2 Vols. Brill, Leiden/Boston, pp. 304–1134. https://doi.org/10.1163/9789004296855_004
    Sharp, D. (1874) The Pselaphidae and Scydmaenidae of Japan. Transactions of the Entomological Society of London, 1874 (1), 105–130. https://doi.org/10.1111/j.1365-2311.1874.tb00160.x
    Shen, Q. & Yin, Z.-W. (2020) Five new species and a new record of Sathytes Westwood from eastern China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4751 (2), 386–394. https://doi.org/10.11646/zootaxa.4751.2.13
    Sugaya, H. & Maruyama, M. (2003) Biological notes on Tribasodites picticornis, with new locality records in the Ryukyus, Japan (Coleoptera: Staphylinidae: Pselaphinae). Entomological Review of Japan, 58 (1), 7–8.
    Wang, D. & Yin, Z.-W. (2015) Two new species of Batriscenellus Jeannel (Coleoptera: Staphylinidae: Pselaphinae) from China. Coleopterists Bulletin, 69 (3), 405–409. https://doi.org/10.1649/0010-065X-69.3.405
    Wang, D. & Yin, Z.-W. (2016) New species and records of Batraxis Reitter (Coleoptera: Staphylinidae: Pselaphinae) in continental China. Zootaxa, 4147 (4), 443–465. https://doi.org/10.11646/zootaxa.4147.4.6
    Wang, X.D., Zhong, X.H., Liu, S.Z., Liu, J.G., Wang, Z.Y. & Li, M.H. (2008) Regional assessment of environmental vulnerability in the Tibetan Plateau: Development and application of a new method. Journal of Arid Environments, 72, 1929–1939. https://doi.org/10.1016/j.jaridenv.2008.06.005
    Weise, J. (1877) Japanische Staphilinidae [sic] und Pselaphidae. Deutsche Entomologische Zeitschrift, 21 (1), 88–100.
    Westwood, J.O. (1870) Descriptions of twelve new exotic species of the Coleopterous family Pselaphidae. Transactions of the Royal Entomological Society of London, 1870, 125–132. https://doi.org/10.1111/j.1365-2311.1870.tb01868.x
    Yao, T., Thompson, L.G., Mosbrugger, V., Zhang, F., Ma, Y., Luo, T., Xu, B., Yang, X., Joswiak, D.R., Wang, W., Joswiak, M.E., Devkota, L.P., Tayal, S., Jilani, R. & Fayziev, R. (2012) Third Pole environment (TPE). Environmental Development, 3, 52–64. https://doi.org/10.1016/j.envdev.2012.04.002
    Yin, Z.-W. (2017) Lisubatrus dongzhiweii gen. et sp. nov. from the Gaoligong Mountains, China (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae, 57 (2), 593–601. https://doi.org/10.1515/aemnp-2017-0091
    Yin, Z.-W. (2018a) Loeblibatrus Yin, a new genus of myrmecophilous Pselaphinae (Coleoptera: Staphylinidae) from Southern China. Coleopterists Bulletin, 72 (2), 233–240. 2018. https://doi.org/10.1649/0010-065X-72.2.233
    Yin, Z.-W. (2018b) Cuccodorodes gen. nov., an apterous Batrisitae from the tropical Himalayas (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae, 58 (2) , 321–330. https://doi.org/10.2478/aemnp-2018-0027
    Yin, Z.-W. (2018c) First record of Arthromelodes Jeannel in China, with description of a new species (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys, 744, 43–47. https://doi.org/10.3897/zookeys.744.23318
    Yin, Z.-W. (2019) Two new species and new records of the Pselaphodes-complex of genera from China (Coleoptera: Staphylinidae: Pselaphinae). Revue suisse de Zoologie, 126 (2), 197–202. https://doi.org/10.5281/zenodo.3463447
    Yin, Z.-W. (2021) A new species of the rare myrmecophilous genus Myrmicophila Yin and Li from Xizang, China (Coleoptera: Staphylinidae: Pselaphinae). Coleopterists Bulletin, 75 (3), 684–688. https://doi.org/10.1649/0010-065X-75.3.684
    Yin, Z.-W., Hlaváč, P. & Li, L.-Z. (2013a) Further studies on the Pselaphodes complex of genera from China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys, 275, 23–65. https://doi.org/10.3897/zookeys.275.4571
    Yin, Z.-W. & He, L. (2020) New cavernicolous Pselaphinae from Sichuan, China (Coleoptera: Staphylinidae). Coleopterists Bulletin, 74 (4), 827–836. https://doi.org/10.1649/0010-065X-74.4.827
    Yin, Z.-W. & Li, L.-Z. (2013a) A new species of Tangius from north India (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys, 346, 23–27. https://doi.org/10.3897/zookeys.346.6233
    Yin, Z.-W. & Li, L.-Z. (2013b) Batrisodes (Excavodes) simplex sp. n. (Coleoptera: Staphylinidae: Pselaphinae): a second species of the genus from continental China. Deutsche Entomologische Zeitschrift, 60 (2), 261–266. https://doi.org/10.1002/mmnd.201300033
    Yin, Z.-W. & Li, L.-Z. (2015a) Review of the Himalayan genus Hingstoniella Jeannel, and description of Besuchetaceus gen. n. from Nepal (Coleoptera: Staphylinidae: Pselaphinae). Revue suisse de Zoologie, 122 (1), 165–180. https://doi.org/10.5281/zenodo.14576
    Yin, Z.-W. & Li, L.-Z. (2015b) Review of some species in the genera Pselaphodes Westwood and Labomimus Sharp (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4040 (3), 331–344. http://dx.doi.org/10.11646/zootaxa.4040.3.5
    Yin, Z.-W., Li, L.-Z. & Zhao, M.-J. (2011a) Contributions to the knowledge of the myrmecophilous pselaphines (Coleoptera, Staphylinidae, Pselaphinae) from China. VIII. Myrmicophila tangliangi, a new genus and species of Batrisini from Yunnan. Sociobiology, 58 (1), 175–183.
    Yin, Z.-W., Li, L.-Z. & Zhao, M.-J. (2011b) Batriscenellus Jeannel (Coleoptera, Staphylinidae, Pselaphinae) redefined, with notes on the Chinese species. Zootaxa, 3016 (1), 37–50. https://doi.org/10.11646/zootaxa.3016.1.3
    Yin, Z.-W., Li, L.-Z. & Zhao, M.-J. (2011c) Contributions to the knowledge of the myrmecophilous pselaphines (Coleoptera, Staphylinidae, Pselaphinae) from China. VI. Hingstoniella Jeannel is a myrmecophile, with notes on its definition and systematic position (Coleoptera: Staphylinidae: Pselaphinae). Sociobiology, 57 (2), 389–396.
    Yin, Z.-W., Li, L.-Z. & Zhao, M.-J. (2011d) Discovery in the caves of Guangxi, China: three new troglobitic species of Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 3065 (1), 49–59. https://doi.org/10.11646/zootaxa.3065.1.5
    Yin, Z.-W., Li, L.-Z. & Zhao, M.-J. (2011f) Taxonomical study on the genus Pselaphodes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part II. Annales Zoologici, 61 (3), 463–481. https://doi.org/10.3161/000345411X603337
    Yin, Z.-W., Li, L.-Z. & Zhao, M.-J. (2012a) Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History, 46 (13–14), 831–857. https://doi.org/10.1080/00222933.2011.651651
    Yin, Z.-W., Li, L.-Z. & Zhao, M.-J. (2012b) Contributions to the knowledge of the myrmecophilous pselaphines (Coleoptera, Staphylinidae, Pselaphinae) from China. I. Tangius glabellus gen. et sp. nov. from Tibet. Zootaxa, 3397 (1), 55–61. https://doi.org/10.11646/zootaxa.3397.1.4
    Yin, Z.-W. & Lin, Y.-J. (2020) Tibetyrus gen. nov., a new myrmecophilous Tyrini from Xizang, China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4786 (1), 131–137. https://doi.org/10.11646/zootaxa.4786.1.10
    Yin, Z.-W. & Nomura, S. (2011) Taxonomic study on Batrisodellus Jeannel of China, with discussion on the systematic position of Batrisodellus callissimus Nomura & Wang, 1991. Spixiana, 34 (1), 33–38.
    Yin, Z.-W., Nomura, S. & Li, L.-Z. (2013b) Redescription of Tyrus Aubé and T. sinensis Raffray (Coleoptera: Staphylinidae: Pselaphinae), and two new species from Sichuan, Southwest China. Zootaxa, 3637 (1), 47–57. http://dx.doi.org/10.11646/zootaxa.3637.1.5
    Yin, Z.-W., Nomura, S. & Li, L.-Z. (2015b) Ten new species of cavernicolous Tribasodites from China and Thailand, and a list of East Asian cave-inhabiting Pselaphinae (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 55 (1), 105–127.
    Yin, Z.-W. & Nomura, S. & Zhao, M.-J. (2011e) Contributions to the knowledge of the myrmecophilous pselaphines (Coleoptera, Staphylinidae, Pselaphinae) from China. VII. Liuyelis gen. n., a rare myrmecophilous Trichonychini associated with Camponotus ants (Formicidae). Sociobiology, 57 (3), 587–596.
    Yin, Z.-W. & Nomura, S. & Zhao, M.-J. (2012c) Batrisoplisus Raffray, 1908, a new synonymy of Trisinus Raffray, 1894 (Coleoptera, Staphylinidae, Pselaphinae), with description of three new Chinese species. Zootaxa, 3222 (1), 61–68. https://doi.org/10.11646/zootaxa.3222.1.4
    Yin, Z.-W., Shen, J.-W. & Li, L.-Z. (2015a) New species and new combinations of Asian Batrisodes Reitter (Coleoptera, Staphylinidae, Pselaphinae), and synonymy of Batrisodellus Jeannel with Batrisodes. Deutsche Entomologische Zeitschrift, 62 (1), 45–54. https://doi.org/10.3897/dez.62.4539
    Yin, Z.-W. & Shen, Q. (2020) Fifteen new species of Sathytes Westwood from China (Coleoptera, Staphylinidae, Pselaphinae), with an updated checklist of world species. European Journal of Taxonomy, 722, 37–74. https://doi.org/10.5852/ejt.2020.722.1133
    Yin, Z.-W. & Zhou, G.-C. (2018) Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa, 4457 (4), 589–594. https://doi.org/10.11646/zootaxa.4457.4.9
    Zhang, Q., Yuan, R., Singh, V.P., Xu, C.-Y., Fan, K., Shen, Z., Wang, G. & Zhao, J. (2022) Dynamic vulnerability of ecological systems to climate changes across the Qinghai-Tibet Plateau, China. Ecological Indicators, 134, 108483. https://doi.org/10.1016/j.ecolind.2021.108483
    Zhang, W.-X. & Yin, Z.-W. (2021) Two new species of Batrisodes Reitter from Shaanxi, China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4938 (2), 243–250. https://doi.org/10.11646/zootaxa.4938.2.5
    Zhang, Y.-Q., Li, L.-Z. & Yin, Z.-W. (2019) Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4554 (2), 497–531. https://doi.org/10.11646/zootaxa.4554.2.7
    Zhang, W.-X. & Yin, Z.-W. (2022) Two new species of myrmecophilous Pselaphinae from China (Coleoptera, Staphylinidae). Revue Suisse de Zoologie. [in press]
    Zhao, M.-J., Yin, Z.-W. & Li, L.-Z. (2010) Description of a new species of the genus Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae) from East China with a key to world species. ZooKeys, 64, 25–31. https://doi.org/10.3897/zookeys.64.45

  2.  

  3.