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Type: Article
Published: 2022-03-18
Page range: 151-199
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Accidental monstrosities: Taxonomic chimeras in Ostracoda (Crustacea)

University of Graz, Institute of Earth Sciences (Geology & Palaeontology), Heinrichstrasse 26, 8010 Graz, Austria
University of Gdańsk, Faculty of Biology, Department of Evolutionary Genetics and Biosystematics, Laboratory of Biosystematics and Ecology of Aquatic Invertebrates, Wita Stwosza 59, 80-308 Gdańsk, Poland
School of Geography, Queen Mary University of London, Mile End Road, London E1 4NS, UK
Crustacea non-marine ostracods taxonomic artefacts comparative morphology geometric morphometrics Candona balatonica Candona levanderi Candoniella Fabaeformiscandona

Abstract

Taxonomic chimeras, artificial taxa created unintentionally by amalgamation of morphological traits belonging to different taxonomic units, can be found in 19th century to present day scientific literature. We recognise two types of such artefacts in Ostracoda. Chimera Type 1 is represented by species defined by morphological traits belonging to two (or more) different valid taxa at the rank of species. A thorough comparative analysis of carapace and limb characteristics of Fabaeformiscandona balatonica (Daday) sensu Bronshtein (1947) allows us to conclude that it is a chimera comprising F. balatonica and F. levanderi (Hirschmann), for which we provide new, expanded diagnoses. Chimera Type 2 refers to a genus defined by juvenile morphological traits that also occur in other genera. Analysis of Candoniella Schneider, 1956 shows it to be an artefact based on morphological traits belonging to juveniles of at least three genera: Pseudocandona Kaufmann, 1900, Fabaeformiscandona Krstić, 1972 and Neglecandona Krstić, 2006. Elimination of taxonomic artefacts is necessary to improve not only taxonomy but also adjacent domains of investigation like the ecology and geographical distribution of confused taxa. Considered in historical contexts, the creation and perpetuation of such accidental monstrosities may be attributed to social motivations as well as limitations of material, literature and communication.

 

References

  1. Absolon, A. (1970) Fossiles Vorkommen von Candona balatonica Daday, 1894 (Ostracoda). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historische Geologie, 10, 199–204.
    Absolon, A. (1978) Die Gattung Candona (Ostracoda) im Quartär von Europa. Rozpravy Československé Akademie Věd, Řada Matematických a Přirodnich Věd, 88 (5), 1–73.
    Aerts, D., Broekaert, J., Gabora, L. & Sozzo, S. (2016) Generalizing prototype theory: a formal quantum framework. Frontiers in Psychology, 7 (418), 1–21. https://doi.org/10.3389/fpsyg.2016.00418
    Agalarova, D.A., Kadyrova, Z.K. & Kulieva, S.A. (1961) Ostrakody pliocenovyh i postpliocenovyh otlozhenij Azerbaydzhana. Azerneshr, Baku, 420 pp. [in Russian]
    Akatova, N.A. (1971) O nekotoryh ostrakodah roda Candona iz Komi ASSR. Trudy Komi Filiala Akademii Nauk SSSR, 22, 106–109. [in Russian]
    Altman, D.G. & Bland, J.M. (1983) Measurement in medicine: the analysis of method comparison studies. The Statistician, 32 (3), 307–317.
    https://doi.org/10.2307/2987937
    Anderson, M.J. & Robinson, J. (2003) Generalized discriminant analysis based on distances. Australian and New Zealand Journal of Statistics, 45 (3), 301–318.
    https://doi.org/10.1111/1467-842X.00285
    Anderson, M.J., Gorley, R.N. & Clarke, K.R. (2008) PERMANOVA+ for PRIMER: Guide to Software and Statistical Methods. Primer-E, Plymouth, 214 pp.
    Baltanás, A. & Danielopol, D.L. (2011) Geometric Morphometrics and its use in ostracod research: a short guide. Joannea Geologie und Paläontologie, 11, 235–272.
    Beldescu, S. (1961) Ostracode din complexul de Balti Crapina-Jijila. Com. Academia Republicii Populare Române, 11 (1), 7–13. [in Romanian]
    Benson, R.H. (2003) A biomechanical theory of ostracode carapace morphology. The Paleontological Society Papers, 9, 89–100.
    https://doi.org/10.1017/S1089332600002163
    Bland, J. & Altman, D. (1986) Statistical methods for assessing agreement between two methods of clinical measurement. The Lancet, 327 (8476), 307–310. https://doi.org/10.1016/S0140-6736(86)90837-8
    Brady, G.S. (1864) On species of Ostracoda new to Britain. Annals and Magazine of Natural History, Series 3, 13, 59–64. https://doi.org/10.1080/00222936408681574
    Brady, G.S., Crosskey, H.W. & Robertson, D. (1874) A monograph of post-Tertiary Entomostraca of Scotland including species from England and Ireland. Monographs of the Palaeontological Society, 28, 1–232. https://doi.org/10.5962/bhl.title.84825
    Brady, G.S. & Robertson, D. (1870) The Ostracoda and Foraminifera of tidal rivers. Part 1. Annals and Magazine of Natural History, Series 4, 6 (1), 1–33. https://doi.org/10.1080/00222937008696200
    Bromley, R.G. (1979) Chalk and bryozoan limestone: facies, sediments and depositional environments. In: Birkelund, T. & Bromley, R.G. (Eds.), Cretaceous-Tertiary Boundary Events. I. The Maastrichtian and Danian of Denmark. University of Copenhagen, Copenhagen, pp. 16–32.
    Bronshtein, Z.S. (1947) Fauna SSSR. Rakoobraznye Tom 2, Vypusk 1: Ostracoda presnyh vod. Zoologicheskij Institut Akademii Nauk SSSR, Novaja Serija 31, Moskva/Leningrad, 339 pp. [in Russian, English translation: Bronshtein, Z.S. (1988) Freshwater Ostracoda, Fauna of the USSR, Crustaceans. A.A. Balkema, Rotterdam, 470 pp.]
    Clarke, J.A. (2004) Morphology, phylogenetic taxonomy and systematics of Ichthyornis and Apatornis (Avialae, Ornithurae). Bulletin of the American Museum of Natural History, 286, 1–179.
    https://doi.org/10.1206/0003-0090(2004)286<0001:MPTASO>2.0.CO;2
    Clerkin, P.J, Ebert, D.A. & Kemper, M. (2017) New species of Chimaera (Chondrichthyes: Holocephali: Chimaeriformes: Chimaeridae) from the Southwestern Indian Ocean. Zootaxa, 4312 (1), 1–37. https://doi.org/10.11646/zootaxa.4312.1.1
    Daday, J. (1894) Ujabb adatok a Balaton mikrofaunaìjaìnak ismereteìhez: Nematoda, Rotatoria, Entomostraca. Mathematikai és természettudományi Értesítő, 12 (1), 122–145. [in Hungarian]
    Daday, J. (1897) Crustaceen. Resultate der wissenschaftlichen Erforschung des Balatonsees, 2 (1), 163–193.
    Daday, J. (1900) Ostracoda Hungariae. Kiadja a Magyar Tudomanyos Akademia, Budapest, 320 pp. [in Hungarian]
    Danielopol, D.L. (1969) Recherches sur la morphologie de l’organe copulateur mâle chez quelques Ostracodes du genre Candona Baird (Fam. Cyprididae Baird). In: Neale, J.W. (Ed.), The taxonomy, morphology and ecology of Recent Ostracoda. Oliver & Boyd, Edinburgh, pp. 136–153.
    ‎Danielopol, D.L. (1980) Sur la biologie de quelques ostracodes Candoninae epigés et hypogés d’Europe. Bulletin du Musée National d’Histoire Naturelle de Paris, Series 4, 2 (2), 471–506.
    Danielopol, D.L., Gross, M., Piller, W.E. & Baltanás, A. (2008) Ostracods of the Paratethys Sea and Lake Pannon – Perspectives for renewal of cooperative projects. Senckenbergiana lethaea, 88 (1), 141−145.
    https://doi.org/10.1007/BF03043985
    Danielopol, D.L., Namiotko, T., von Grafenstein, U., Fuhrmann, R., Decrouy, L., Gross, M., Pichler, M. & Picot, L. (2015) The implementation of taxonomic harmonisation for Candoninae (Ostracoda, Cypridoidea): A heuristic solution for Fabaeformiscandona tricicosa (Diebel and Pietrzeniuk). Geo-Eco-Marina, 21, 111−158.
    https://doi.org/10.5281/zenodo.46362
    Diebel, K. & Pietrzeniuk, E. (1969) Ostracoden aus dem Mittelpleistozän von Süßenborn bei Weimar. Paläontologische Abhandlungen, Abteilung A: Paläozoologie, 3 (3–4), 463−488.
    Diebel, K. & Pietrzeniuk, E. (1975) Mittel- und jungpleistozäne Ostracodenfaunen des Raums Potsdam-Brandenburg-Rathenow in stratigraphischer und ökologischer Sicht. Zeitschrift für geologische Wissenschaften, 3 (9), 1197−1233.
    Dykan, N. (2006) Systematization of Quaternary Ostracoda of Ukraine (a reference-book). National Academy of Sciences of Ukraine, Institute of Geological Sciences, Kiev, 427 pp. [in Russian]
    Fischer, S. (1851) Abhandlung über das genus Cypris und dessen in der Umgebung von St. Petersburgund von Fall bei Reval vorkommended Arten. Mémoires présentés à l’Académie Impériale des Sciences de Saint-Pétersbourg, 7, 127–167.
    Foerster, H. von (2003) Understanding Understanding. Essays on cybernetics and cognition. Springer-Verlag, New York, 362 pp.
    Forró, L., Meisch, C., Petersen, H. & Martens, K. (1987) Ostracoda taxa described by E. Daday, together with a catalogue of pertinent material in the Hungarian Natural History Museum. Miscellanea Zoologica Hungarica, 4, 33–63.
    Fowler, J., Cohen, L. & Jarvis, P. (1998) Practical statistics for field biology. J. Wileys & Sons, Chichester, 272 pp.
    Fox, M.J. (2011) Prototype theory: An alternative concept theory for categorizing sex and gender? In: Smiraglia, R.P. (Ed.), Proceedings from North American Symposium on Knowledge Organization, Toronto, Canada, 3, pp. 151−159.
    https://doi.org/10.7152/nasko.v3i1.12799
    Fuhrmann, R. (2012) Atlas quartärer und rezenter Ostrakoden Mitteldeutschlands. Altenburger Naturwissenschaftliche Forschungen, 15, 1−320.
    Giavarina, D. (2015) Understanding Bland Altman analysis. Biochemia Medica, 25 (2), 141–151.
    https://doi.org/10.11613/BM.2015.015
    Goldman, A & O’Connor, C. (2021) Social epistemology. In: Zalta, E. (Ed.), The Stanford Encyclopedia of Philosophy. pp. 128 pp. Available from https://plato.stanford.edu/entries/epistemology-social/ (accessed 1 March 2021)
    Griffiths, H.I. (1995) European Quaternary freshwater Ostracoda: a biostratigraphic and palaeobiogeographic primer. Scopolia (Museum Historiae Naturalis Sloveniae), 34, 1–168.
    Griffiths, H.I. & Evans, J.G. (l995) An annotated checklist of British Pleistocene, Holocene and modern freshwater ostracods. Journal of Micropalaeontology, 14, 59–65. https://jm.copernicus.org/articles/14/59/1995/jm-14-59-1995.pdf
    Hanneman, S.K. (2008) Design, analysis and interpretation of method-comparison studies. AACN Advanced Critical Care, 19 (2), 223–234.
    https://pubmed.ncbi.nlm.nih.gov/18560291/
    Hartmann, G. (1959) Neue Ostracoden von Teneriffa. Zoologischer Anzeiger, 162 (5–6), 160–171.
    Hartwig, W. (1900) Eine neue Candona der Provinz Brandenburg: Candona reniformis nov. spec. Sitzungs-Berichte der Gesellschaft naturforschender Freunde zu Berlin, 4, 139–141.
    Hiller, D. (1972) Untersuchungen zur Biologie und zur Ökologie limnischer Ostracoden aus der Umgebung von Hamburg. Archiv für Hydrobiologie, Supplement 40 (4), 400–497.
    Hirschmann, N. (1912) Beitrag zur Kenntnis der Ostrakodenfauna des Finnischen Meerbusens. Zweite Mitteilung. Acta Societatis pro Fauna et Flora Fennica 36 (2), 1–68.
    Horne, D.J. (1982a) On Loxoconcha dimorpha. A Stereo Atlas of Ostracod Shells, 9 (5), 27–32.
    Horne, D.J. (1982b) On Loxoconcha linleyi Horne sp. nov. A Stereo Atlas of Ostracod Shells, 9 (6), 33–40.
    Horne, D.J., Danielopol, D.L. & Martens, K. (1998) Reproductive behaviour. In: Martens, K. (Ed.), Sex and parthenogenesis: evolutionary ecology of reproductive modes in non-marine ostracods. Backhuys Publishers, Leiden, pp. 157–197.
    Horne, D.J., Bunbury, J. & Whittaker, J. (2011) Taxonomic harmonisation and calibration of nonmarine ostracods for palaeoclimate applications: the case of Candona acutula Delorme, 1967. Joannea Geologie und Paläontologie, 11, 76–79. [extended abstract]
    Karanovic, I. (2012) Recent freshwater ostracods of the world. Springer, Berlin-Heidelberg, 608 pp. https://doi.org/10.1007/978-3-642-21810-1
    Kaufmann, A. (1900) Cypriden und Darwinuliden der Schweiz. Revue Suisse de Zoologie, 8, 209–423. https://doi.org/10.5962/bhl.part.10584
    Kempf, E.K. (1980) Index and bibliography of non-marine Ostracoda 1: Index A. Sonderveröffentlichungen des Geologischen Instituts der Universität zu Köln, 35, 1–188.
    Kempf, E.K. (1997) Index and bibliography of non-marine Ostracoda 6: Index A, Supplement 1. Sonderveröffentlichungen des Geologischen Instituts der Universität zu Köln, 109, 1–148.
    Kiss, A. (2007) Factors affecting spatial and temporal distribution of Ostracoda assemblages in different macrophyte habitats of shallow lake (Lake Feher, Hungary). Hydrobiologia, 585 (1), 89–98.
    https://doi.org/10.1007/s10750-007-0631-8
    Klie, W. (1937) Ostracoden und Harpacticoiden aus brackigen Gewässern an der bulgarischen Küste des Schwarzen Meeres. Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten Sofia, 10 (1), 1–42.
    Klie, W. (1938) Ostracoda, Muschelkrebse. In: Dahl, F. (Ed.), Die Tierwelt Deutschlands und der angrenzenden Meeresteile. Gustav Fischer Verlag, Jena, pp. 1–230.
    Konovalova, V.A. (2016) Representatives of the genus Fabaeformiscandona Krstić, 1972 (Crustacea, Ostracoda) from Quaternary deposits of Western Siberia. Revue de Micropaléontologie, 59 (2), 168–179.
    https://doi.org/10.1016/j.revmic.2015.11.002
    Kovalenko, A.L. (1976) Sovremennye ostrakody bassejna Dnestra. Izdatelstvo Shtiintsa, Kishinev, 180 pp. [in Russian]
    Kovalenko, A.L. (1983) Postembrionalnoe razvitie Candona parallela G.W. Müller (Crustacea, Ostracoda). Izvestija Akademii Nauk Moldavskoj SSR, Serija biologicheskih i himicheskih nauk, 1983 (1), 33–38.
    Kovalenko A.L. (1988). Kandonidy (Candonidae, Ostracoda) yugo-zapada SSSR. Izdatelstvo Shtiintsa, Kishinev, 175 pp. [in Russian]
    Krstić, N. (1972) Genus Candona (Ostracoda) from Congeria beds of southern Pannonian Basin. Monographs of the Serbian Academy of Sciences and Arts, Section Natural and Mathematical Sciences, 39, 1–145. [in Serbian with English summary]
    Krstić, N. (2006) Pliocene ostracodes of the Paludinian Beds in the Pannonian Plain, Serbian Part. Herald of the Nature History Museum Special Publication. Herald of the Nature History Museum, Belgrade, 411 pp.
    Lane, R. (2007) Taxonomy in Europe in the 21st Century. EDIT. Available from: http://ww2.bgbm.org/EditDocumentRepository/Taxonomy21report.pdf (accessed 1 April 2021)
    Linhart, J., Brauneis, W., Neubauer, W. & Danielopol, D.L. (2007) Morphomatica, Computer Program. Version 1.6.1. In ostracodology Available from: http://iewarchiv.uni-graz.at/palstrat/Ostracodology/ (accessed 1 April 2021)
    Löffler, H. (1969) Recent and subfossil distribution of Cytherissa lacustris (Ostracoda) in lake Constance. Mitteilungen Internationaler Verein für theoretische und angewandte Limnologie, 17, 240–251. https://doi.org/10.1080/05384680.1969.11903891
    Lord, A. (2020) Towards objectivity in Ostracoda species definition. Bulletin de la Société Géologique de France, 191 (1), 27. https://doi.org/10.1051/bsgf/2020029.
    Mallet, J. (2007) Species, concepts of. In: Levin, S.A. (Ed.), Encyclopedia of Biodiversity. Academic Press, Cambridge, Massachusetts, pp. 679–691. https://doi.org/10.1016/B978-0-12-384719-5.00131-3
    Martens, K. (Ed.) (1998) Sex and parthenogenesis. Evolutionary ecology of reproductive modes in non-marine ostracods. Backhuys Publishers, Leiden, 335 pp.
    Matoshko, A., Matoshko, A., de Leeuw, A. & Stoica, M. (2016) Facies analysis of the Balta Formation: Evidence for a large late Miocene fluvio-deltaic system in the East Carpathian Foreland. Sedimentary Geology, 343, 165–189. https://doi.org/10.1016/j.sedgeo.2016.08.004
    Meisch, C. (2000) Freshwater Ostracoda of Western and Central Europe. In: Schwoerbel, J. & Zwick, P. (Eds.), Süßwasserfauna von Mitteleuropa. 8 (3). Spektrum Akademischer Verlag, Heidelberg, pp. 1–522.
    Meisch, C., Smith, R.J. & Martens, K. (2019) A subjective global checklist of the extant non-marine Ostracoda (Crustacea). European Journal of Taxonomy, 492, 1–135.
    https://doi.org/10.5852/ejt.2019.492
    Müller, G.W. (1900) Deutschlands Süßwasser-Ostracoden. Zoologica, 12 (30), 1–112.
    Namiotko, T. & Martins M.J.F. (2008) Sex ratio of sub-fossil Ostracoda (Crustacea) from deep lake habitats in northern Poland. Palaeogeography, Palaeoclimatology, Palaeoecology, 264 (3–4), 330–337.
    https://doi.org/10.1016/j.palaeo.2007.06.024
    Namiotko, T., Danielopol, D.L., Meisch, C., Gross, M. & Mori, N. (2014) Redefinition of the genus Typhlocypris Vejdovský, 1882 (Ostracoda, Candonidae). Crustaceana, 87 (8–9), 952–984.
    https://doi.org/10.1163/15685403-00003338
    Negadaev-Nikonov, K.N. (1965) Fauna ostrakod verhnechetvertichnyh otlozhenii srednego Priputja. Izvestija Akademii Nauk Moldavskoj SSR, 1965 (8), 12–19. [in Russian]
    Neubauer, W. & Linhart, J. (2008) Approximating and distinguishing Ostracoda by the use of B-splines. In: Danielopol, D.L., Gross, M. & Piller, W.S. (Eds.), Contributions to Geometric Morphometrics. Berichte des Institutes für Erdwissenschaften, Karl-Franz-Universität Graz, 13, pp. 21–42.
    Ostermeyer, E. (1937) Über zwei neue Candona-Arten aus dem Saalekreis. Zoologischer Anzeiger, 119, 155-159.
    Padial, J.M., Miralles, A., De la Riva, I. & Vences, M. (2010) The integrative future of taxonomy. Frontiers in Zoology, 7 (16), 1–14.
    https://doi.org/10.1186/1742-9994-7-16
    Petkovski, T. (1977) Ostracodenfauna des Mindelsees (Südwest-Deutschland). Acta Musei Macedonici Scientiarum Naturalium, 15 (3), 49–94.
    Petkovski, T. & Karanovic, I. (2000) On Eucandona balatonica (Daday, 1894) and Eucandona levanderi (Hirschman, 1912), two rare and little-known ostracod species (Crustacea, Ostracoda). Mésogée, 58, 77–84.
    Pigliucci, M. (2003) Species as family resemblance concepts: the (dis-)solution of the species problem? BioEssays, 25 (6), 596–602.
    https://doi.org/10.1002/bies.10284
    Popova-Lvova, M.G. (1965) Ostrakody iz Pliotsenovyh otlozhenij Bashkiroskogo Preduralya. In: Kainozoy Bashkirskogo Preduralya. Part 2.Nedra, Moskva, pp. 210–315. [in Russian]
    Popova-Lvova, M.G. (1972) Ostrakody—Ostracoda. In: Yahimovich, V.L. (Ed) Fauna i flora Akkulaeva (opornyj razrez srednego Akchagyla –srednevo Apsherona Bashkirii). Akademiya Nauk SSSR, Bashkirskij Filial, Institut Geologii, Ufa, pp. 71–97. [in Russian]
    Popova-Lvova, M.G. (1977) Ostrakody. In: Goretskij, G.I. (Ed.), Fauna i flora Simbugino (opornyj razrez akchagyla i apsherona Bashkirii). Izdatelstvo Nauka, Moskva, pp. 86–93. [in Russian]
    Rohlf, F.J. & Slice, D.E. (1999) BIOMstat for Windows. Statistical software for biologists. Exeter Softwere, Setauket, 23 pp.
    Rohlf, F.J. & Sokal, R.R. (1995) Statistical Tables. W.H. Freeman and Co., New York, 199 pp.
    Rosch, E. (1978) Principles of categorisation. In: Rosch, E. & Lloyd, B.B. (Eds.), Cognition and categorisation. Laurence Erlbaum, Hilsdale, New York, pp. 27–48.
    Rowe, T., Ketcham, R.A., Denison, C., Colbert, M., Xu, X. & Currie, P.J. (2001) The Archaeoraptor forgery. Nature, 410, 539–540. https://doi.org/10.1038/35069145
    Saniotis, A. (2007) Social and genomic constructions of chimera. Journal of Futures Studies, 11 (3), 47–60.
    Sars, G.O. (1887) Nye bidrag til kundskaben om Middelhavets invertebratfauna. 4. Ostracoda mediterranea (Sydeuropæiske Ostracoder). Archiv for Mathematik og Naturvidenskab, 12, 173–324. [in Norwegian] https://doi.org/10.5962/bhl.title.10252
    Schneider, D.C. (1994) Quantitative ecology—Spatial and temporal scaling. Academic Press, San Diego, 395 pp. https://doi.org/10.1016/B978-0-08-092563-9.50018-2
    Schneider, G.F. (1956) Rod Candoniella Schneider gen. n. In: Suzin, A.V. (Ed.), Ostrakody tretichnih otlozhenij severnogo Predkavkazja. Izdatelstvo Gostopteh, Moskva, pp. 34–37. [in Russian]
    Scott, R.W., Wan, X., Wang, C. & Huang, Q. (2012) Late Cretaceous chronostratigraphy (Turonian-Maastrichtian): SK1 core Songliao Basin, China. Geoscience Frontiers, 3 (4), 357–367. https://doi.org/10.1016/j.gsf.2012.02.004
    Simon, H.A. (1962) The architecture of complexity. Proceedings of the American Philosophical Society, 106 (6), 467–482.
    Simundic, A.M. (2016) Statistical analysis in method comparison part one. acutecaretesting.org website. Available from: https://acutecaretesting.org/en/articles/statistical-analysis-in-method-comparison-studies-part-one (accessed 21 July 2021)
    Siveter, D.J. (2015) Imaging the microfossil world. In: Danielopol, D.L., Baltanás, A., Carbonel, P., Colin, J.-P., Crasquin, S., Decrouy, L., De Deckker, P., Gliozzi, E., Groos-Uffenorde, H., Horne, D.J., Iepure, S., Keyser, D., Kornicker, L.S., Lord, A., Martens, K., Matzke-Karasz, R., Miller, C.G., Oertli, H.J., Pugliese, N., Russo, R., Sames, B., Schön, I., Siveter, D.J., Smith, A., Viehberg, F.A., Wouters, K. & Yassini, I. (Eds.), From Naples 1963 to Rome 2013. A brief review of how the International Research Group on Ostracoda (IRGO) developed as a social communication system. Palaeogeography, Palaeoclimatology, Palaeoecology, 419, pp. 3–22. [Supplementary Material, 21, 1–6]
    https://doi.org/10.1016/j.palaeo.2014.08.016
    Smith, W. (1880) Dictionary of Greek and Roman biography and mythology. J. Murray, London, 1115 pp. (Digitised Edition 2009 – Google ID 4h5XAAAAMAAJ).
    Smith, R.J. & Kamiya, T. (2007) Copulatory behaviour and sexual morphology of three Fabaeformiscandona Krstić 1972 (Candoninae, Ostracoda, Crustacea) species from Japan, including descriptions of two new species. Hydrobiologia, 585, 225–248.
    https://doi.org/10.1007/s10750-007-0640-7
    Sohar, K. & Meidla, T. (2010) Changes in the Early Holocene lacustrine environment inferred from the subfossil ostracod record in the Varangu section, northern Estonia. Estonian Journal of Earth Sciences, 59 (3), 195–206.
    https://doi.org/10.3176/earth.2010.3.02
    Sokal, R.R. & Rohlf, F.J. (1995) Biometry. The principles and practice of statistics in biological research. W.H. Freeman and Co., New York, 859 pp.
    Spadi, M., Gliozzi, E., Boomer, I., Stoica, M. & Athersuch, J. (2019) Taxonomic harmonization of Neogene and Quaternary candonid genera (Crustacea, Ostracoda) of the Paratethys. Journal of Systematic Palaeontology, 17 (19), 1665–1698.
    https://doi.org/10.1080/14772019.2018.1545708
    Stearns, S.C. (1994) The evolutionary links between fixed and variable traits. Acta Palaeontologica Polonica, 38 (3–4), 215–232.
    Stracke, A. (2008) From the photography to the digitalized outline for Morphomatica. Berichte des Institutes für Erdwissenschaften Karl-Franzens-Universität Graz, 13, 69–74.
    Su, D. & Neale, J.W. (1991) Plio/Pleistocene Candonidae (Freshwater Ostracoda) from boreholes in The Nanning Area, Guangxi Province, Southern China. Journal of Micropalaeontology, 9 (2), 1569–166.
    https://doi.org/10.1144/jm.9.2.159
    Sywula, T. (1971) Notes on Ostracoda. XII. New and interesting species from Poland. Bulletin de l’Academie Polonaise des Sciences, Classe 2, Serie des sciences biologiques, 19 (10), 665–670.
    Sywula, T. (1974) Małżoraczki (Ostracoda). Fauna Słodkowodna Polski. Vol. 24. PWN, Warszawa, 315 pp.
    Tesakov, A.S., Titov, V.V., Simakova, A.N., Frolov, P.D., Syromyatnikova, E.V., Kurshakov, S.V., Trikhunkov, Y.I., Sotnikova, M.V., Kruskop, S.V., Zelenkov, N.V., Tesakova, E.M. & Palatov, D.M. (2017) Late Miocene (Early Turolian) vertebrate faunas and associated biotic record of the Northern Caucasus: Geology, palaeoenvironment, biochronology. Fossil Imprint, 73 (3–4), 383–444.
    https://doi.org/10.2478/if-2017-0021
    Toulmin, S. (1953) The philosophy of science. An introduction. Hutchinson’s Univ. Library, London, 176 pp.
    Van Baak, C.G.C., Mandic, O., Lazar, I., Stoica, M. & Krijgsman, W. (2015) The Slanicul de Buzau section, a unit stratotype for the Romanian stage of the Dacian Basin (Plio-Pleistocene, Eastern Paratethys). Palaeogeography, Palaeoclimatology, Palaeoecology, 440, 594–613.
    https://doi.org/10.1016/j.palaeo.2015.09.022
    Van Baak, C.G.C., Stoica, M., Grothe, A., Aliyeva, E. & Krijgsman, W. (2016) Mediterranean-Paratethys connectivity during the Messinian salinity crisis: The Pontian of Azerbaijan. Global and Planetary Change, 141, 63–81.
    https://doi.org/10.1016/j.gloplacha.2016.04.005
    Vejdovský, F. (1882) Thierische Organismen der Brunnenwässer von Prag. s.n., Prag, 70 pp.
    Viehberg, F.A. (2004) Paleolimnological study based on ostracods (Crustacea) in Late-Glacial and Holocene deposits of lake Krakower See (Mecklenburg-Vorpommern, NE Germany). Studia Quaternaria, 21, 109–115.
    Wagner, A. (2005) Robustness and evolvability in living systems. Princeton Univ. Press, Princeton, 368 pp.
    Wang, H., Li, S., Zhang, H., Cao, M. & Horne, D.J. (2019) Biostratigraphic and palaeoenvironmental significance of Campanian-early Maastrichtian (Late Cretaceous) ostracods from the Jiaozhou Formation of Zhucheng, Shandong, China. Cretaceous Research, 93, 4–21. https://doi.org/10.1016/j.cretres.2018.08.023
    West-Eberhardt, M.-J. (2008) Are genes good markers of biological traits? In: Weinstein, M., Vaupel, J.W. & Wachter, K.J. (Eds.), Biosocial Surveys. National Academies Press, Washington, D.C., pp. 175–193.
    Wetterich, S., Schirrmeister, L. & Pietrzeniuk, E. (2005) Freshwater ostracodes in Quaternary permafrost deposits in the Siberian Arctic. Journal of Paleolimnology, 34 (3), 363–376. https://doi.org/10.1007/s10933-005-5801-y
    Whatley, R. (2012) The “Out of India” hypothesis: further supporting evidence from the extensive endemism of Maastrichtian non-marine Ostracoda from the Deccan volcanic region of peninsular India. Revue de Paléobiologie, 31, 229–248.
    Yu, G., Harrison, S.P. & Xue, B. (2001) Lake status records from China: Data base documentation. Technical Reports Max-Planck-Institut für Biogeochemie, 4, 1–247.
    Zanina, I.E. (1968) Paleontologi Sovetskogo Sojuza: Spravochnik. Izdatelstvo Nauka—Leningradsko Otdelenie, Leningrad, 214 pp. [in Russian]
    Zhai, D., Xiao, R., Fan, J., Zhou, L., Wen, R. & Pang, Q. (2013) Spatial heterogeneity of the population age structure of the ostracode Limnocythere inopinata in Hulun Lake, Inner Mongolia and its implications. Hydrobiologia, 716, 29–46. https://doi.org/10.1007/s10750-013-1541-6

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