Abstract
Species of four lace bug genera of the subfamily Tinginae (Hemiptera: Heteroptera: Tingidae), namely Cysteochila Stål, 1873, Hurdchila Drake, 1953, Physatocheila Fieber, 1844, and Xynotingis Drake, 1948, occurring in Japan, are revised. A total of 13 species are identified and diagnosed. Previous records of Cysteochila consueta Drake, 1948 from Japan are considered to be the misidentification of C. chiniana Drake, 1942. Limnostatua gen. nov. is described to accommodate Leptodictya (currently Hurdchila) lewisi Scott, 1880 (type species of the new genus). A new synonymy is proposed: Limnostatua lewisi (Scott, 1880), comb. nov. = Cysteochila vota Drake, 1948 syn. nov. Three new combinations, namely Physatocheila chiniana (Drake, 1942) comb. nov., P. consueta (Drake, 1948) comb. nov., and P. euthenia (Drake & Ruhoff, 1965) comb. nov., all of which were transferred from Cysteochila, are proposed. Three species, namely C. abundantis Drake & Poor, 1937, P. consueta comb. nov. (previous records all pertaining to misidentifications) and P. euthenia comb. nov., are newly recorded from Japan. Finally, identification keys for Japanese tingid genera and subgenera as well as for species of Cysteochila and Physatocheila are provided.
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Tomokuni, M. (2006b) Additional material of Heteroptera (Insecta) collected at the Imperial Palace, the Akasaka Imperial Gardens, and the Tokiwamatsu Imperial Villa, Tokyo Japan. Memoirs of the National Science Museum, 43, 349–354. [in Japanese, English summary]
Tomokuni, M. (2014) Heteroptera from the Garden of the Imperial Palace, Tokyo, Japan. Memoirs of the National Science Museum, 50, 359–369. [in Japanese, English summary]
Vinokurov, N.N., Kanyukova, E.V. & Golub, V.B. (2010) The catalogue of bugs (Heteroptera) of Asian part of Russia. Nauka, Novosibirsk, 320 pp.
Yamada, K. & Ishikawa, T. (2016) Family Tingidae. In: Hayashi, M., Tomokuni, M., Yoshizawa, K. & Ishikawa, T. (Eds.), Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, pp. 429–435. [in Japanese]
Yamada, K. & Tomokuni, M. (2012) Family Tingidae Laporte, 1832. In: Ishikawa, T., Takai, M. & Yasunaga, T. (Eds.), A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, pp. 180–213, pls. 2–13. [in Japanese]
Yano, S., Takechi, L. & Kikuhara, Y. (2013) Heteroptera of Ehime Prefecture, Shikoku, Japan (Insecta, Hemiptera) I. Bulletin of Omogo mountain museum, 5, 23–52. [in Japanese]
Yazaki, M. (2009) Hemipterous insects confirmed by Yahagi River forest investigation. Yahagi River Research, 13, 29–38. [in Japanese]
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Tomokuni, M. (1979) Records of the Family Tingidae collected by Dr. Kintaro Baba in Niigata Prefecture (Hemiptera). Transactions of Essa Entomological Society, 50, 135–138. [in Japanese]
Tomokuni, M. (1982) The heteropterous fauna of the Ozegahara moor and its adjacent area. In: Hara, H., Asahina, S., Sakaguchi, Y., Hogetsu, K. & Yamagata, N. (Eds.), Ozegahara: Scientific Research of the Highmoor in Central Japan. Japan Society for the Promotion of Science, Tokyo, pp. 347–353.
Tomokuni, M. (1987) The Tingidae of Hokkaido, Japan (Insecta, Heteroptera). Memoirs of the National Science Museum, 20, 115–121. [in Japanese, English summary]
Tomokuni, M. (1988) [Two distributional records of Tingidae from Hokkaido, Japan]. Rostria, 39, 652. [in Japanese]
Tomokuni, M. (1994) Type materials of several Japanese and Korean Tingidae (Heteroptera), with designation of Lectotypes and notes on Matsumura’s collection. Japanese Journal of Entomology, 62 (4), 839–844.
Tomokuni, M., Hayashi, M. & Usui, T. (2000) Hemiptera excluding Sternorrhyncha from the Garden of the Imperial Palace, Tokyo, Japan. Memoirs of the National Science Museum, 36, 35–55. [in Japanese, English summary]
Tomokuni, M. (2005) Heteroptera (Insecta) from the Akasaka Imperial Gardens and the Tokiwamatsu Imperial Villa, Tokyo, Japan. Memoirs of the National Science Museum, 39, 397–408. [in Japanese, English summary]
Tomokuni, M. (2006a) A small collection of Tingidae (Insecta, Heteroptera) from Taiwan, with a checklist of the known species. Memoirs of the National Science Museum, 44, 59–69.
Tomokuni, M. (2006b) Additional material of Heteroptera (Insecta) collected at the Imperial Palace, the Akasaka Imperial Gardens, and the Tokiwamatsu Imperial Villa, Tokyo Japan. Memoirs of the National Science Museum, 43, 349–354. [in Japanese, English summary]
Tomokuni, M. (2014) Heteroptera from the Garden of the Imperial Palace, Tokyo, Japan. Memoirs of the National Science Museum, 50, 359–369. [in Japanese, English summary]
Vinokurov, N.N., Kanyukova, E.V. & Golub, V.B. (2010) The catalogue of bugs (Heteroptera) of Asian part of Russia. Nauka, Novosibirsk, 320 pp.
Yamada, K. & Ishikawa, T. (2016) Family Tingidae. In: Hayashi, M., Tomokuni, M., Yoshizawa, K. & Ishikawa, T. (Eds.), Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, pp. 429–435. [in Japanese]
Yamada, K. & Tomokuni, M. (2012) Family Tingidae Laporte, 1832. In: Ishikawa, T., Takai, M. & Yasunaga, T. (Eds.), A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, pp. 180–213, pls. 2–13. [in Japanese]
Yano, S., Takechi, L. & Kikuhara, Y. (2013) Heteroptera of Ehime Prefecture, Shikoku, Japan (Insecta, Hemiptera) I. Bulletin of Omogo mountain museum, 5, 23–52. [in Japanese]
Yazaki, M. (2009) Hemipterous insects confirmed by Yahagi River forest investigation. Yahagi River Research, 13, 29–38. [in Japanese]
Yazaki, Y. (2019) [Distributional records of Physatocheila nigrintegerrima from Nagano-ken and Niigata-ken, Honshu, Japan]. Gekkan-Mushi, 586, 3–4. [in Japanese]
Yonekura, K. & Kajita, T. (2003–2020) BG Plants Wamei-Gakumei Index (YList). Available from: http://ylist.info (accessed 23 February 2020) [in Japanese]