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Type: Article
Published: 2022-06-07
Page range: 151-188
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A rich and diverse fauna: an illustrated guide to the cetoniines of Ghana with comparisons to Ivory Coast and Benin (Coleoptera: Scarabaeidae: Cetoniinae)

Systematics and Evolution Laboratory, Department of Biology, Western Kentucky University, Bowling Green, KY 42101-3576 USA.
3726 Southeast Liebe St., Portland, OR 97202-4032 USA.
Insect Collection, Agricultural Science and Production Department, Zamorano University (Escuela Agrícola Panamericana), Zamorano, Honduras.
Systematics and Evolution Laboratory, Department of Biology, Western Kentucky University, Bowling Green, KY 42101-3576 USA
Am Steigberg 25, D-97502 Eurbach, Germany.
272, Chemin du Prince, B 7050, Jurbise, Belgium.
Coleoptera Fruit Chafers West Africa Guinean Forests Conservation

Abstract

The currently known cetoniine fauna of Ghana is discussed and illustrated. It now includes 60 genera and 148 taxa, including 9 subspecies. Comparison to the adjacent countries of Ivory Coast and nearby Benin gives evidence that the faunas of all three countries surprisingly may still be grossly under-sampled and hence poorly known for what is considered a relatively well-known group of beetles. At least 22 species known from both Ivory Coast and Benin are expected to be found in Ghana but still have yet to be reported. Notes on several species concepts and the alternate use of names are also given to clarify the taxonomy of the taxa known from this area.

 

References

  1. Allard, V. (1991) Goliathini 4. The Beetles of the World. Vol. 11. Les Coléoptères du Monde. Sciences Nat, Venette, 142 pp.
    Antoine, P. (1989) Contribution à l’étude des Cetoniinae de Côte d’Ivoire. Revue Française d’Entomologie, New Sereis, 11 (1), 29–35.
    Antoine, P. (1998) Contribution à la connaissance des Cetoniidae du Cameroun. Entomologia Africana, 3 (1), 11–18.
    Antoine, P. (2002) Contribution à la connaissance des Cetoniidae africains. Coléoptères, 8 (4), 65–92.
    Antoine, P. (2012) Contribution à la connaissance des Cetoniidae Africains. V. Cetoniimania NS, 3, 3–5.
    Bánki, O., Roskov, Y., Vandepitte, L., DeWalt, R. E., Remsen, D., Schalk, P., Orrell, T., Keping, M., Miller, J., Aalbu, R., Adlard, R., Adriaenssens, E., Aedo, C., Aescht, E., Akkari, N., Alonso-Zarazaga, M.A., Alvarez, B., Alvarez, F., Anderson, G., Angel, M., Appeltans, W., Arango, C. Artois, T., Arvanitidis, C., Atkinson, S., Auffenberg, K., Baeyens, W., Bailly, N., Baker, E., Bamber, R., Bank, R., Barber, A., Barber-James, H., Barbosa, J.P., Barthe, J.-F., Bartolozzi, L., Bartsch, I., Beccaloni, G., Beccaloni, G.W., Bellamy, C.L., Bellan-Santini, D., Bellinger, P.F., Ben-Dov, Y., Bernot, J., Bezerra, T.N., Bieler, R., Bisby, F., Bitner, M.A., Blasco-Costa, I., Bock, P., Bouchard, P., Bouchet, P., Bourgoin, T., Boury-Esnault, N., Boxshall, G., Boyko, C., Brandão, S., Braun, H., Bray, R., Brock, P.D., Bruce, N., Burckhardt, D., Bush, L., Böttger-Schnack, R., Büscher, T., Błażewicz-Paszkowycz, M., Cairns, S., Calonje, M., Carballo, J.L., Cardoso, L., Carrera-Parra, L., Castilho, R.C., Catalano, S., Chatrou, L.W., Chevillotte, H., Christiansen, K.A., Cigliano, M.M., Coleman, C.O., Collins, A., Corbari, L., Cordeiro, R., Cornils, A., Costello, M., Costello, M.J., Crosby, M.R., Cruz-López, J.A., Culham, A., Cárdenas, P., Daly, M., Daneliya, M., Dauvin, J.C., Davie, P., Davison, A.J., De Broyer, C., De Prins, J., De Prins, W., DeSalle, R., DeWalt, R.E., W., Deem, L.S., Defaye, D., Dellapé, P.M., Den Heyer, J., Deneudt, K., Deprez, T., Dmitriev, D.A., Dohrmann, M., Doner, S., Dorkeld, F., Downey, R., Dutilh, B., Díaz, M.-C., Döring, M., Eades, D.C., Eibye-Jacobsen, D., Eisendle, U., Eitel, M., El Nagar, A., Emig, C., Emig, C.C., Engel, M.S., Enghoff, H., Evans, G.A., Evenhuis, N.L., Faber, M., Farjon, A., Fauchald, K., Fautin, D., Favret, C., Fernández-Rodríguez, V., Figueroa, D., Finn, J., Fišer, C., Forró, L., Francis, A., Froese, R., Fuchs, A., Furuya, H., Gabelle, C., Gao, Y., Garcia-Alvarez, O., Gardner, M., Garic, R., Garnett, S., Gasca, R., Gattolliat, J.-L., Gaviria-Melo, S., Gerken, S., Gheerardyn, H., Gibson, D., Gibson, R., Gielis, C., Giribet, G., Gittenberger, A., Glasby, C., Glover, A.G., Gofas, S., Govaerts, R., Grabowski, M., Granado, A. de A., Gray, A., Grimaldi, D.A., Gross, O., Guerra-García, J.M., Guglielmone, A., Guilbert, E., Gusenleitner, J., Gómez-Noguera, S.E., Haas, F., Hadfield, K.A., Hagborg, A., Hajdu, E., Harrach, B., Harris, L., Harrison, R., Hassler, M., Hayward, B.W., Heads, S.W., Hendrycks, E., Henry, T.J., Herbert, D., Hernandes, F.A., Hernandez, F., Hernández-Crespo, J.C., Herrera Bachiller, A., Hine, A., Ho, J.-S., Hobern, D., Hodda, M., Hoeksema, B., Hoenemann, M., Holleman, A., Holovachov, O., Holstein, J., Hooge, M., Hooper, J., Hopcroft, R., Hopkins, H., Horak, I., Horton, T., Hosoya, T., Houart, R., Hošek, J., Hughes, L., Huijbers, C., Häuser, C., Iniesta, L.F.M., Ivanenko, S., Janssen, R., Janssens, F., Jarms, G., Jaume, D., Jazdzewski, K., Ji, L.-Q., Johnson, K.P., Junglen, S., Jóźwiak, P., Kabat, A., Kamiński, M.J., Kanda, K., Kantor, Y., Karanovic, I., Kathirithamby, J., Kelly, M., Kim, Y.-H., King, R., Kirk, P.M., Kirk, P., Kitching, I., Klautau, M., Knowles, N.J., Koenemann, S., Korovchinsky, N.M., Kotov, A., Kouwenberg, J., Krapp-Schickel, T., Krishna, V., Kristensen, R.M., Kroh, A., Kroupa, A.S., Krupovic, M., Kuhn, J.H., Kury, A.B., Kury, M.S., Kvaček, J., Köhler, F., Lado, C., Lambert, G., Lazarus, D., Le Coze, F., LeCroy, S., Leduc, D., Leermakers, M., Lefkowitz, E.J., Lichtwardt, R. (†), Lobanov, A., Lohrmann, V., Lojen, S., Londoño-Mesa, M., Longhorn, S.J., Lorenz, W., Lowry, J., Lujan-Toro, B.E., Lumen, R., Lyal, C., Lyal, C.H.C., Lyangouzov, I., Lörz, A.-N., Macklin, J.A., Madin, L., Maehr, M.D., Maehr, M.D., Magill, R.E., Magnien, P., Mah, C., Mal, N., Mamo, B., Mamos, T., Manconi, R., Marek, P., Marshall, B., Martin, J.H., Martin, S.L., Martínez-Muñoz, C.A., Maslin, B., McFadden, C., McKamey, S., McMurtry, J.A., Medvedev, S., Mees, J., Meidla, T., Mendes, A.C., Merrin, K., Mesa, N.C., Messing, C., Migeon, A., Miller, D.R., Mills, C., Minelli, A., Mokievsky, V., Molodtsova, T., Mooi, R., Morandini, A., Moreira da Rocha, R., Morrow, C., Mushegian, A., Narita, J.P.Z., Nealova, L., Neu-Becker, U., Neubauer, T.A., Neubert, E., Neuhaus, B., Newton, A., Ng Kee Lin, P., Nguyen, A., Nibert, M.L., Nicolson, D., Nijhof, A., Nishikawa, T., Norenburg, J., Noyes, J., O'Hara, T., Ochoa, R., Oosterbroek, P., Opresko, D., Osborne, R., Osigus, H.-J. Oswald, J.D., Ota, Y., Otte, D., Ouddane, B., Ouvrard, D., Ower, G., Pape, T., Paulay, G., Pauly, D., Paxton, H., Perrier, V., Petrusek, A., Peña Santiago, R., Picton, B., Pisera, A., Pitkin, B., Poore, G., Pulawski, W.J., Pyle, R., Páll-Gergely, B., Rainer, H., Raz, L., Read, G., Rees, T., Rein, J.O., Reip, H., Reuscher, M., Richling, I., Rius, M., Robertson, T., Robinson, G., Robinson, G.S.(†), Rogacheva, A., Romani, L., Rosenberg, G., Roskov, Y., Rubino, L., Ruggiero, M., Ríos, P., Rützler, K., Sabanadzovic, S., Salazar-Vallejo, S., Sanborn, A., Sartori, M., Saucède, T., Schierwater, B., Schilling, S., Schmid-Egger, C., Schmidt-Rhaesa, A., Schneider, S., Schoolmeesters, P., Schotte, M., Schuchert, P., Schuh, R.T., Schönberg, C., Scoble, M., Segers, H., Senna, A., Serejo, C., Sforzi, A., Sharma, J., Shear, W., Shenkar, N., Short, M., Siciński, J., Siddell, S., Siegel, V., Sierwald, P., Silva, E.S., Simmonds, P., Simmons, E., Simonsen, T., Sket, B., Smith, A.D., Smith, V.S., Smol, N., Soulier-Perkins, A., South, E.J., Souza-Filho, J.F., Spearman, L., Spelda, J., Stampar, S., Steiner, A., Stemme, T., Sterrer, W., Stevenson, D., Stiewe, M.B.D., Stjernegaard Jeppesen, T., Stoev, P., Strand, M., Stueber, G., Stöhr, S., Suárez-Morales, E., Swalla, B., Swedo, J., Szumik, C., Sánchez-Ruiz, M., Söderström, L., Taiti, S., Takiya, D.M., Tandberg, A.H., Tang, D., Tavakilian, G., Taylor, J., Tchesunov, A., Thessen, A., Thomas, J.D., Thomas, P., Thuesen, E., Thurston, M., Thuy, B., Timm, T., Todaro, A., Turiault, M., Turon, X., Tyler, S., Uetz, P., Ulmer, J.M., Uribe-Palomino, J., Vacelet, J., Vachard, D., Vader, W., Vanhoorne, B., Vanreusel, A., Varsani, A., Venekey, V., Vinarski, M., Vonk, R., Vos, C., Väinölä, R., Walker, P.J., Walker-Smith, G., Walter, T.C., Wambiji, N., Wartel, M., Warwick, S., Watling, L., Weaver, H., Webb, J., Welbourn, W.C., Wesener, T., Whipps, C., White, K., Wieneke, U., Wilson, G.D.F., Wilson, R., Wirth, C.C., Yesson, C., Yoder, M., Yu, D.S.K., Yunakov, N., Zahniser, J., Zanol, J., Zarucchi, J., Zeidler, W., Zerbini, F.M., Zhang, Z.Q., Zhao, Z., Zinetti, F., d'Hondt, J.-L., de Moraes, G.J., de Reus, M., de Voogd, N., ten Hove, H., ter Poorten, J.J., van Nieukerken, E.J., van Soest, R., van Tol, J., von Konrat, M. (2021) Catalogue of Life Checklist. Version 2021-09-21. In: Bailly, N., Bourgoin, T., Cigliano, M.M.Costello, M., de Reus, M., Decock, W., Fuchs, A., Garnett, S., Gray, A., Hernandez, F., Döring, M., Hobern, D., Holleman, A., Hosoya, T., Huijbers, C., Ji, L.-Q., Kathirithamby, J., Nicolson, D., Ower, G., Pape, T., Pyle, R., Raz, L., Rees, T., Robertson, T., Ruggiero, M., Stjernegaard Jeppesen, T., Swedo, J., Vanhoorne, B., Wambiji, N., Weaver, H. & Yoder, M. (Contrib.), Catalogue of Life. Leiden. https://doi.org/10.48580/d4sv
    Bates, H.W. (1884) List of Coleoptera of the Families Carabidae and Scarabaeidae collected by the late W. A. Forbes on the Lower Niger. Proceedings of the Zoological Society of London, 1884, 404–406. https://doi.org/10.1111/j.1096-3642.1884.tb02845.x
    Beinhundner, G. (2017) The Cetoniinae of Africa. Privately published, Euerbach, 1174 + XXIV pp.
    Bouyer, J., Sana Y., Samandoulgou, Y., Cesar, J., Guerrini, L., Kabore-Zoungrana, C. & Dulieu, D. (2007) Identification of ecological indicators for monitoring ecosystem health in the transboundary W Regional Park: a pilot study. Biological Conservation, 138, 73–88. https://doi.org/10.1016/j.biocon.2007.04.001
    Brown, K.S.J. (1991) Conservation of insects and their habitats: insects as indicators. In: Collins, N.M. & Thomas, J.A. (Eds.), The conservation of insects and their habitats. Academic Press, London, pp. 350–404. https://doi.org/10.1016/B978-0-12-181370-3.50020-8
    CIA Worldfact Book (2021) Available from: https://www.cia.gov/the-world-factbook/countries/ghana/ (accessed 21 September 2021)
    ClientEarth (2020) Protecting Ghana’s Atewa Range Forest Reserve from Bauxite mining. Available from: https://www.clientearth.org/latest/latest-updates/news/protecting-ghana-s-atewa-range-forest-reserve-from-bauxite-mining/ (accessed 28 January 2021)
    Conservation International. (2021) Biodiversity Hotspots. Available from: https://www.conservation.org/priorities/biodiversity-hotspots (accessed 10 April 2021)
    De Palma, M. & Frantz, S. (2010) Taxonomic Revison of Megalorhina Westwood and subgeneric classification of Mecynorhina Hope (Coleoptera: Scarabaeidae: Cetoniinae). Natura Edizioni Scientifiche, Bologna, 80 pp., 16 pls.
    De Palma, M. (2009) Taxonomic Revision of Eudicella White (Coleoptera: Cetoniinae) and Iconographic Catalog. Natura Edizione Scientifique, Bologna, 48 pp., 32 pls.
    De Palma, M. (2018) Taxonomic revision of genus Fornasinius Bertoloni, 1853 with description of new taxa Entomologia Africana, 23 (1), 2–18
    De Palma, M., Takano, H., Léonard, P. & Bouyer. T. (2020) Barcoding analysis and taxonomic revision of Goliathus Lamarck, 1802 (Scarabaeidae, Cetoniinae). Entomologia Africana, 25 (1), 11–32.
    De Palma, M., Takano, H., Léonard, P. & Bouyer. T. (2021) Taxonomic revision of the Eudicella White, 1839 complex, with description of three new subgenera. Entomologia Africana, 26 (2), 11–24.
    Devecis, J. (1997) Nouvelles espèces et sous-espèces de Cetoniidae Africains. Lambillionea, 97 (1), 134–140
    Devecis, J. (2001) Contribution á la connaissance des Cetoines du Mali. Cetoniimania, 1, 114–119.
    Endrödi, S. (1973) Entomological explorations in Ghana by Dr. S. Endrödy-Younga 14, Lamellicornia (Coleoptera) I. Annales Historico - Naturalis Musei Nationalis Hungarici, 65, 195–229.
    Endrödi, S. (1976) Lamellicornia (Coleoptera) aus Ghana, II. Annales Historico—Naturalis Musei Nationalis Hungarici, 68, 55–164.
    FAO, (2021) Introduction & Status of the Forestry Sector in Ghana. Available from: http://www.fao.org/3/ab567e/AB567E02.htm (accessed 27 November 2020)
    Fédière, G., Lery, X., Herder, S. & Kouassi, N.K. (1989) Cétoines de Côte d’Ivoire (Coleoptera, Cetoniidae, Cetoniinae). Institut International De Resherche Por Le Development a Adiopodoume (IIRSDA), Abidjan, Côte d’Ivoire, number of pages
    Holm, E. & Marais, E. (1992) Fruit Chafers of Southern Africa. Ekogilde, Hartebeespoort, 326 pp.
    Janson, O. E. (1907a) List of the Cetoniidae collected by the late Leonardo Fea in West Africa. Annali del Museo Civico di Storia Naturale di Genova, 3a, 3 (43), 326–338.
    Janson, O. E. (1907b.) Description of two new species of the genus Coryphocera. Notes from the Leyden Museum, 28, 149–152.
    Johannsmeier, M.F. (1980) Beetles that eat brood in South Africa, South African Bee Journal, 52, 3–5.
    Joly, C. (2001) Contribution to the knowledge of Cetoniinae of Ghana (Coleoptera, Scarabaeidae, Cetoniinae). Lambillonea, CI, 1–10.
    Joly, C. (2011) Compte rendu d’une expédition entomologique au Ghana. Entomologie faunistique – Faunistic Entomology, 63, 207–211.
    Joly, C. (2018) Deuxième Contribution à la connaissance des Cetoniidae du Ghana, Sous - familles des Cetoniinae et des Trichinae (Coleoptera, Scarabaeidae). Lambillionea, 118 (1), 28–89.
    Krajčík, M. (1998) Cetoniidae of the world catalogue. Part 1. Typos Studio, Most, 96 + xxxvi pp.
    Krajčík, M. (1999) Cetoniidae of the world catalogue. Part 2. Typos Studio, Most, 72 + xxiii pp.
    Krikken , J. (1984) A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated lists of the known genera. Zoologische Verhandelingen, 210, 1–75 .
    Le Gall, P. (2010) Affinités biogégraphiques des Insectes du “Dahomey Gap”: présence d’une population de Goliathus goliatus Linné, 1771, au Bénin (Coleoptera, Scarabaeidae, Cetoniinae). Bulletin of the Entomological Society of France, 115 (1), 17–21. https://doi.org/10.3406/bsef.2010.2821
    Le Gall, P. (2015) La faune entomologique du Bénin: Les Cetoniinae (Coleoptera, Cetoniidae: Cetoniinae). Cetoniimania NS, 8, 25–53.
    Le Gall, P., Goergen, G. & Neuenschwander, P. (2002) Les Insectes et le Sillon Dahomeen: Fragmentation et Refuges Forestiers. Biosystema, 20, 73–80.
    Malec, P. (2015) The Cetoniine fauna of Sierra Leone and the basic guidelines to captive breeding of these beetles (Coleoptera: Scarabaeidae: Cetoniinae). Cetoniimania NS, 7, 35–76.
    Maquart, P.O. & Malec, P. (2017) On the distribution and natural history of a rarely encountered species: Goliathus (Fornasinius) klingbeili Zöller, Fiebig, & Schulz, 1995 (Coleoptera, Scarabaeidae, Cetoniinae). Zootaxa, 4341 (3), 441–444. https://doi.org/10.11646/zootaxa.4341.3.12
    Moretto, P. & Rojkoff, S. (2021) Coleoptera, Cetoniidae (Annexe 3c). In: Moretto, P., Cosson, B., Takano, H., Basquin, B., Bordat, P., Boucher, S., Bouyer, T., Danflous, S., Dérozier, V., Eitschberger, U., Juhel, P., Leroy, É., Limbourg, P., Massa, B., Meunier, J.-Y., Miles, W., Orbach, E., Robiche, G., Rojkoff, S. & Silvestre, G., A threatened forest refuge: The mountain forest at Parinari of Mont Tonkoui in Côte d’Ivoire. Assessment of entomological, botanical and ornithological biodiversity. Site conservation values. Catharsius La Revue, Hors-Série, 1, pp. 1–123.
    Mudge, A.D., Orozco, J., Philips, T.K. & Antoine, P. (2012) The cetoniine fauna of the Upper Guinean forests and savannas of Ghana (Coleoptera: Scarabaeidae: Cetoniinae). Terrestrial Arthropod Reviews, 5, 113–174. https://doi.org/10.1163/187498312X635319
    Myers, N., Mittermeier, R.A., Mittermeier, C.G., da Fonesca, G.A.B. & Kent, J. (2000) Biodiversity hotspots for conservation priorities. Nature, 403, 853–858. https://doi.org/10.1038/35002501
    Perissinotto, R., Beinhundner, G. & Strümpher, W. (2021) Description of a new species of Anoplocheilus MacLeay, 1838 from South Africa, with review of the genus. Fragmenta entomologica, 53 (2), 367–376.
    Perissinotto, R. & Beinhundner, G. (2022) Review of Mausoleopsis amabilis (Schaum, 1844) (Scarabaeidae, Cetoniinae), with Description of a New Species and a New Subspecies. Diversity, 2022, 14, 248 https://doi.org/10.3390/d14040248
    Puker, A., Ad’Vincula, E., Korasaki, V., Ferreira, F.N.F. & Orozco, J. (2014) Biodiversity of Cetoniinae beetles (Coleoptera: Scarabaeidae) in introduced and native habitats in the Brazilian Atlantic Forest. Entomological Science, 17, 309–315. https://doi.org/10.1111/ens.12069
    Rojkoff, S. (2012) 1ère Contribution à l’étude des Coenochilus africains. Révision du groupe <strigatus> sensu Schein et description d’une nouvelle espèce (Coleoptera, Cetoniidae, Cetoniinae, Cremastocheilini). Cetoniimania NS, 3, 20–28.
    Rojkoff, S. (2013) Contribution à l'étude des Cremastocheilini africains. Cetoniimania NS, 5, 19–29.
    Rojkoff, S. & Perissinotto, R. (2015) Review of the genera Anelaphinis Kolbe, 1892 and Atrichelaphinis Kraatz, 1898. ZooKeys, 482, 91–142. https://doi.org/10.3897/zookeys.482.8343
    Ruter, G. (1948) Liste des Coléoptères Cétonides et Trichiides recoltés dans la réserve du Banco (Côte d'Ivoire) par la mission Paulian-Delamare Bulletin de la Société entomologique de France, 53 (3–4), 54–55. https://doi.org/10.3406/bsef.1948.16022
    Ruter, G. (1954) La réserve naturelle intégrale du Mont Nimba Fasc II 18 Coléoptères Cétonides, Memoires de l’Institut francais d’Afrique noire, 40 (2), 213–232.
    Ruter, G. (1958) La réserve naturelle intégrale du Mont Nimba Fasc IV 7 Coléoptères Cétonides, Memoires de l’Institut francais d’Afrique noire, 53, 191–198.
    Ruter, G. (1963) Contribution à l’étude des Cetoniinae africains du genre Pachnoda Burmeister Bulletin de l’Institut fondamental d’Afrique noire, 25 (A4), 1127–1143.
    Ruter, G. (1966) Liste des Cetoniinae capturés dans la Rep. de Guinéé. Acta Musei Moraviae, Scientiae biologicae, 51, 285–288.
    Ruter, G. (1969) Contributions à la connaissance de la Faune Entomologique de la Côte-d’Ivoire. Annales du Musee royal du Congo Belge, Series 8 Sciences Zoologiques, 175, 81–100.
    Ruter, G. (1975a) Deux Cétonides Crémastochilides nouveaux du Zaïre. Revue de Zoologie africaine, 89 (4), 969–973.
    Ruter, G. (1975b) Contribution à l’étude biologique du Sénégal septentrional 28. Bulletin de l’Institut fondamental d’Afrique noire, 37 (3), 661–668.
    Sakai, K. & Nagai, S. (1998) The cetoniine beetles of the world. Mushi-Sha, Tokyo, 421 pp.
    Schenkling S. (1921) Coleopterorum Catalogus auspiciis et auxilio, W. Junk. Scarabaeidae: Cetoninae editus S. Schenkling Pars 72. W. Junk, Berlin, 431 pp. https://doi.org/10.1007/978-94-011-9697-0_1
    Schoolmeesters, P. (2021) SCARABS: World Scarabaeidae Database. In: Bánki, O., Roskov, Y., Vandepitte, L., DeWalt, R.E., Remsen, D., Schalk, P., Orrell, T., Keping, M., Miller, J., Aalbu, R., Adlard, R., Adriaenssens, E., Aedo, C., Aescht, E., Akkari, N., Alonso-Zarazaga, M.A., Alvarez, B., Alvarez, F., Anderson, G. et al, Catalogue of Life Checklis. Version 2021-09-01. Catalogue of Life. Leiden. https://doi.org/10.48580/d4sl-38g
    Serrano, A.R.M., Capel, R. A., Nunes, T., Van-dúnem, C. & Santos, N. (2020) The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes. Zootaxa, 4776 (1), 1–130. https://doi.org/10.11646/zootaxa.4776.1.1
    Spector, S. (2006) Scarabaeine dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae): An invertebrate focal taxon for biodiversity research and conservation. Coleopterists Society Monographs, 5, 71–83. https://doi.org/10.1649/0010-065X(2006)60[71:SDBCSS]2.0.CO;2
    Tchibozo, S., Aberlenc, H., Ryckewaert P. & Le Gall, P. (2008) Première évaluation de la bidoversité des Odonates, des Cétoines et des Rhopalocères de la forêt marécageuse de Lokoli (Sud Bénin). Bulletin de la Société Entomologique de France, 113 (4), 497–509. https://doi.org/10.3406/bsef.2008.3046
    Touroult, J. & Delahaye, N. (2007) Apercu des cetoines de Zambie (Coleoptera, Scarabaeiodea, Cetoniidae). Cetoniimania, 3–4, 87–95.
    Touroult, J. & Le Gall, P. (2001) Les Cétoines du Sud-Bénin. Etude comparative du peuplement de différents biotopes. Cetoniimania, 1, 29–39.
    Touroult, J. & Le Gall, P. (2012) Fruit feeding Cetoniinae community structure in an anthropogenic landscape in West Africa. Journal of Insect Conservation, 17, 23–34. https://doi.org/10.1007/s10841-012-9483-2
    U.S. Geological Survey (2021) The Deforestation of the Upper Guinean Forest. Available from: https://eros.usgs.gov/westafrica/land-cover/deforestation-upper-guinean-forest (accessed 15 June 2021)
    Weisse, M. & Goldman, E.D. (2019) The World Lost a Belgium-sized Area of Primary Rainforests Last Year. World Resources Institute. Available from: https://www.wri.org/insights/world-lost-belgium-sized-area-primary-rainforests-last-year (accessed 20 February 2021)

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