Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2022-06-08
Page range: 301-332
Abstract views: 640
PDF downloaded: 32

An annotated checklist of the Vespidae (Hymenoptera: Vespoidea) of Bhutan with new records

Royal Society for Protection of Nature, Thimphu, Bhutan.
Sherubtse College, Royal University of Bhutan, Kanglung, Bhutan.
Naturalis Biodiversity Center, 2300 RA Leiden, Netherlands.
College of Natural Resources, Royal University of Bhutan, Lobesa, Bhutan.
National Biodiversity Centre, Ministry of Agriculture and Forest, Thimphu, Bhutan.
Hymenoptera checklist new records Vespidae Bhutan

Abstract

This checklist covers each of the 73 social and solitary wasps of the family Vespidae currently known from Bhutan. Of these, 11 species in the subfamily Eumeninae, five species in Polistinae and one species in Vespinae are reported as new records for the country.

 

References

  1. Archer, M.E. (1989) A key to the world species of the Vespinae (Hymenoptera). Research Monographs of the College of Ripon & York St. John, 2, 1–41.
    Archer, M.E. (1994) Taxonomy, distribution and nesting biology of the Vespa bicolor group (Hymenoptera, Vespinae). Entomologist's Monthly Magazine, 130, 149–158.
    Archer, M.E. (1999) Taxonomy and world distribution of the Euro-Asian species of Dolichovespula (Hymenoptera, Vespinae). Entomologist's Monthly Magazine, 135, 153–160.
    Archer, M.E. (2000) Taxonomy and distribution of the nocturnal hornets, Provespa (Hymenoptera, Vespinae). Entomologist's Monthly Magazine, 136, 127–130.
    Archer, M.E. (2006) Taxonomy, distribution and nesting biology of species of the genus Dolichovespula (Hymenoptera, Vespidae). Entomological Science, 9, 281–293. https://doi.org/10.1111/j.1479-8298.2006.00174.x
    Archer, M.E. (2012) Vespine Wasps of the World: Behaviour, Ecology & Taxonomy of the Vespinae. Monograph Series 4. SIRI Scientific Press, Manchester, 352 pp.
    Barthélémy, C., Lee, J.X.Q. & Kojima, J.I. (2014) Provisional distributional checklist of Hong Kong social wasps (Hymenoptera: Vespidae: Vespinae, Polistinae, Stenogastrinae). Hong Kong Entomological Bulletin, 6 (1), 3–14.
    Bingham, C.T. (1897) Fauna of British India, including Ceylon and Burma Hymenoptera. Vol. I. Wasps and Bees. Taylor and Francis, London, 579 pp.
    Bodlah, I., Naeem, M., Khan, M.R., Bodlah, M.A. & Akhter, T. (2012) Genus Delta de Saussure (Hymenoptera: Eumininae: Vespidae) from Punjab province of Pakistan. Pakistan Journal of Zoology, 44 (3), 759–764.
    Bui, H.T.Q., Tran, N.T., Bozdoğan, H. & Nguyen, L.T.P. (2020) Additional knowledge respecting taxonomy of the social wasp genus Ropalidia (Hymenoptera: Vespidae: Polistinae) from Vietnam, with new records of three species and an updated key to species. Zootaxa, 4722 (1), 034–040. https://doi.org/10.11646/zootaxa.4722.1.2
    Carpenter, J.M. (1996) Distributional checklist of species of the genus Polistes (Hymenoptera: Vespidae; Polistinae, Polistini). American Museum Novitates, 3188, 1–39.
    Carpenter, J.M. & Kojima, J.I. (1997) Checklist of the species in the subfamily Vespinae (Insecta: Hymenoptera: Vespidae). Natural History Bulletin of Ibaraki University, 1, 51–92.
    Castro, L. (2019) Una nueva introducción accidental en el género Vespa Linnaeus, 1758: Vespa bicolor Fabricius, 1787 en la provincia de Málaga (España). Revista gaditana de Entomología, X (1), 47–56.
    Choi, M.B. & Kwon, O. (2015) Occurrence of Hymenoptera (wasps and bees) and their foraging in the southwestern part of Jirisan National Park, South Korea. Journal of Ecology and Environment, 38 (3), 367–374. https://doi.org/10.5141/ecoenv.2015.038
    Daglio, A. (2019) Yellowjackets of the World Taxonomy, Distribution, and Nesting Biology. Lap Lambert Academic Publishing, Mauritius, 377 pp.
    Das, B.P. & Gupta, V.K. (1983) A catalogue of the families Stenogastridae and Vespidae from the Indian subregion (Hymenoptera: Vespoidea). Oriental Insects, 17 (1), 395–464. https://doi.org/10.1080/00305316.1983.10433698
    Das, B.P. & Gupta, V.K. (1989) The social wasps of India and the adjacent countries. Oriental Insects Monograph, 11, 1–292. https://doi.org/10.1080/00305316.1989.11835532
    Dorji, P., Gyeltshen, T., Klein, W. & Nidup, T. (2017) New records of social wasps (Hymenoptera: Vespinae: Vespa and Provespa) from Bhutan. Journal of Threatened Taxa, 9 (4), 10102–10108. https://doi.org/10.11609/jot.2423.9.4.10102-10108
    Dorji, P., Klein, W. & Nidup, T. (2016) Study of paper wasps (Hymenoptera: Vespidae: Polistinae) of Bhutan. Bio Bulletin, 2 (1), 128–136.
    Dorji, P., Klein, W. & Nidup, T. (2017a) A field guide to the common bees and wasps of Bhutan. National Biodiversity Centre, Ministry of Agriculture and Forest, Thimphu, 127 pp.
    Dorji, P., Klein, W. & Nidup, T. (2017b) Taxonomic study of social vespid wasps (Hymenoptera: Vespidae: Vespinae & Polistinae) in Bhutan. Journal of Insect Biodiversity and Systematics, 3 (2), 91–104.
    Dover, C. (1925) Further notes on the Indian Diplopterous wasps. The Journal of the Asiatic Society of Bengal, 20, 289–305.
    Dubatolov, V.V. & Dolgikh, A.M. (2009) Social wasps (Hymenoptera, Vespidae: Polistinae, Vespinae) of the Bolshekhekhtsirsky Nature Reserve (the Khabarovsk Suburbs), with notes on their distribution in the Lower Amur. Amurian Zoological Journal, I (1), 76–82.
    Dvořák, L. (2007) The social wasp fauna of Pakistan (Hymenoptera, Vespidae). Linzer Biologische Beiträge, 39 (1), 51–55.
    Dvořák, L. & Castro, L. (2007) New and noteworthy records of vespid wasps (Hymenoptera: Vespidae) from the Palaearctic region. Acta Entomologica Musei Nationalis Pragae, 47, 229–236.
    Fateryga, A.V. (2018) Wasps of the family Vespidae (Hymenoptera) of the Crimean Peninsula. Entomofauna, 39 (1), 193–233.
    Gawas, S.M. & Gupta, A. (2017) First record of Polistes (Polistella) dawnae Dover and Rao (Hymenoptera: Vespidae) from India. Journal of Biological Control, 31 (2), 79–81. https://doi.org/10.18311/jbc/2017/15914
    Gawas, S.M., Kumar, P.G., Pannure, A., Gupta, A. & Carpenter, J.M. (2020) An annotated distributional checklist of Vespidae (Hymenoptera: Vespoidea) of India. Zootaxa, 4784 (1), 1–87. https://doi.org/10.11646/zootaxa.4784.1.1
    Gawas, S.M., Kumar, P.G., Pannure, A. & Sureshan, P.M. (2019) Checklist of vespid wasps (Hymenoptera: Vespidae) of Goa, India, with new records and a key to species. Zootaxa, 4585 (2), 269–294. https://doi.org/10.11646/zootaxa.4585.2.3
    Ghosh, D., Kumar, P.G., Tripathy, B. & Chandra, K. (2020) An updated checklist of Vespidae (Insecta: Hymenoptera) from Arunachal Pradesh, India with new distributional records from the State. Halteres, 11, 36–43. https://doi.org/10.5281/zenodo.4042968
    Giordani Soika, A. (1975) Ergebnisse der Bhutan-Expedition des Naturhistorischen Museums in Basel, Hymenoptera: Fam. Eumenidae. Entomologica Basiliensia, 1, 387–393.
    Gusenleitner, J. (2006) Über Aufsammlungen von Faltenwespen in Indien (Hymenoptera: Vespidae). Linzer Biologische Beiträge, 38 (1), 677–695.
    Gusenleitner, J. (2011) Eine Aufsammlung von Faltenwespen aus Laos im Biologiezentrum Linz (Hymenoptera: Vespidae: Vespinae, Stenogastrinae, Polistinae, Eumeninae). Linzer Biologische Beiträge, 43 (2), 1351–1368.
    Husemann, M., Sterr, A., Maack, S. & Abraham, R. (2020) The northernmost record of the Asian hornet Vespa velutina nigrithorax (Hymenoptera, Vespidae). Evolutionary Systematics, 4, 1–4. https://doi.org/10.3897/evolsyst.4.47358
    Kim, J.K. (2017) Descriptions of three new species of the genus Symmorphus Wesmael, 1836 (Hymenoptera: Vespidae: Eumeninae) from South Korea, with an updated revised key to Far Eastern species. Zootaxa, 4350 (2), 234–250. https://doi.org/10.11646/zootaxa.4350.2.2
    Kim, J.K., Choi, M. & Moon, T.Y. (2006) Occurrence of Vespa velutina Lepeletier from Korea, and a revised key for Korean Vespa species (Hymenoptera: Vespidae). Entomological Research, 36, 112–115. https://doi.org/10.1111/j.1748-5967.2006.00018.x
    Kim, J.K., Moon, T.Y. & Yoon, I.B. (1994) Systematics of vespine wasps of Korea, I. Genus Vespa Linnaeus (Vespidae, Hymenoptera). Korean Journal of Entomology, 24 (2), 107–115.
    Kim, J.K. & Yamane, S. (2001) A revision of Eumenes Latreille (Hymenoptera: Vespidae) from the Far East Asia, with descriptions of one new species and one new subspecies. Entomological Science, 4 (2), 139–155.
    Kojima, J.I. & Carpenter, J.M. (1997) Catalog of species in the polistine tribe Ropalidiini (Hymenoptera: Vespidae). American Museum Novitates, 3199, 1–96.
    Kojima, J.I., Lambert, K., Nguyen, L.T.P. & Saito, F. (2007) Taxonomic notes on the paper wasps of the genus Ropalidia in the Indian subcontinent (Hymenoptera: Vespidae). Entomological Science, 10, 373–393. https://doi.org/10.1111/j.1479-8298.2007.00237.x
    Kumar, P.G. (2010) New distributional records of some species of the subgenus Polistes (Gyrostoma) Kirby (Hymenoptera: Vespidae) from various states of India and adjacent countries. Records of the Zoological Survey of India, 110 (4), 41–45.
    Kumar, P.G. (2012) Redescription and new distributional records of Delta esuriens (Fabricius) (Hymenoptera: Vespidae: Eumeninae) from Indian states. Records of the Zoological Survey of India, 112 (4), 55–60.
    Kumar, P.G. (2013a) A revision of the genus Subancistrocerus de Saussure (Hymenoptera: Vespidae: Eumeninae) from Indian subcontinent. In: Gupta, V. K. & Verma, A K. (Eds.), Animal Diversity, Natural History and Conservation. Vol. 2. Daya Publishing House, New Delhi, pp. 53–68.
    Kumar, P.G. (2013b) A taxonomic review of the Oriental genus Labus de Saussure, 1867 (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent. Biosystematica, 7 (2), 29–37.
    Kumar, P.G. (2013c) A taxonomic revision of Phimenes Giordani Soika (Hymenoptera: Vespidae: Eumeninae) of Indian subcontinent. Records of the Zoological Survey of India, 113 (3), 119–135.
    Kumar, P.G. & Carpenter, J.M. (2013) A taxonomic review of the genus Antodynerus de Saussure, 1855 (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent. Zootaxa, 3731 (2), 267–278. https://doi.org/10.11646/zootaxa.3731.2.7
    Kumar, P.G. & Carpenter, J.M. (2018) A taxonomic review of the genus Vespula Thomson (Hymenoptera: Vespidae: Vespinae) from the Indian subcontinent. Halteres, 9, 86–110. https://doi.org/10.5281/zenodo.1258043
    Kumar, P.G., Carpenter, J.M., Castro, L. & Sureshan, P.M. (2017) A taxonomic review of the Indian species of the genus Eumenes Latreille (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 4317 (3), 469–498. https://doi.org/10.11646/zootaxa.4317.3.3
    Kumar, P.G., Carpenter, J.M., Kishore, L. & Shareef, K.P.M. (2014) Additional notes on the genus Apodynerus Giordani Soika, 1993 (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent. Halteres, 5, 47–51.
    Kumar, P.G., Carpenter, J.M. & Sharma, G. (2014) A review of the genus Paraleptomenes Giordani Soika, 1970 (Hymenoptera: Vespidae: Eumeninae: Odynerini) from the Indian subcontinent, with the description of a new species from the eastern Himalayas. Zootaxa, 3802 (1), 131–143. https://doi.org/10.11646/zootaxa.3802.1.11
    Kumar, P.G., Carpenter, J.M., Srinivasan, G., Shareef, K.P.M. & Kishore, L. (2017) A taxonomic review of the genus Stenodyneriellus Giordani Soika (Hymenoptera: Vespidae: Eumeninae) from Indian subcontinent with descriptions of two new species. Insect Diversity and Taxonomy, 2017, 161–190.
    Kumar, P.G., Carpenter, J.M. & Sureshan, P.M. (2016a) Additions to the knowledge of the genus Allorhynchium van der Vecht from the Indian subcontinent with the description of a new species from Kerala (Hymenoptera: Vespidae: Eumeninae). Halteres, 7, 29–34. https://doi.org/10.5281/zenodo.54869
    Kumar, P.G., Carpenter, J.M. & Sureshan, P.M. (2016b) A taxonomic review of the genus Antepipona de Saussure, 1855 (Hymenoptera: Vespidae: Eumeninae) from India. Zootaxa, 4150 (5), 501–536. https://doi.org/10.11646/zootaxa.4150.5.1
    Kumar, P.G., Castro, L., Carpenter, J.M. & Sheikh, A.H. (2018) Additions to the knowledge of the genus Symmorphus Wesmael, 1836 (Hymenoptera: Vespidae: Eumeninae) in India with two new records of species. Graellsia, 74 (2), 1–7. https://doi.org/10.3989/graellsia.2018.v74.196
    Kumar, P.G., Kundu, B.G. & Kazmi, S.I. (2015) Insecta: Hymenoptera: Vespidae. Zoological Survey of India, Fauna of Uttar Pradesh, State Fauna Series, 22 (Part-2), 581–595.
    Kumar, P.G., Mazumdar, P.C. & Pham, P.H. (2015) New records of hornet wasps of the genus Vespa Linnaeus (Hymenoptera: Vespidae) from Indian states and Bangladesh. Records of the Zoological Survey of India, 115 (Part-2), 215–219.
    Kumar, P.G., Shareef, K.P.M., Kishore, L. & Carpenter, J.M. (2013) A taxonomic study on the Oriental genus Apodynerus Giordani Soika (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent. Biosystematica, 7 (1), 23–31.
    Kumar, P.G. & Sharma, G. (2014a) Taxonomic studies on vespid wasps (Hymenoptera: Vespoidea: Vespidae) of Sunderbans Biosphere Reserve, West Bengal, India. Part 1. Records of the Zoological Survey of India, 114 (Part-4), 563–580.
    Kumar, P.G. & Sharma, G. (2014b) Taxonomic studies on vespid wasps (Vespidae: Vespoidea: Hymenoptera: Insecta) of Rajasthan, India with six new records from the State. Journal on New Biological Reports, 3 (3), 240–258.
    Kumar, P.G. & Sharma, G. (2015a) A review of the genus Allorhynchium van der Vecht, 1963 (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent. Prommalia, III, 20–34.
    Kumar, P.G. & Sharma, G. (2015b) Taxonomic studies on vespid wasps (Hymenoptera: Vespoidea: Vespidae) of Chhattisgarh, India. Journal of Threatened Taxa, 7 (14), 8096–8127.
    Kumar, P.G. & Srinivasan, G. (2010) Taxonomic studies of hornet wasps (Hymenoptera: Vespidae) Vespa Linnaeus of India. Records of the Zoological Survey of India, 110 (Part-2), 57–80.
    Li, T.J., Barthélémy, C. & Carpenter, J.M. (2019) The Eumeninae (Hymenoptera, Vespidae) of Hong Kong (China), with description of two new species, two new synonymies and a key to the known taxa. Journal of Hymenoptera Research, 72, 127–176. https://doi.org/10.3897/jhr.72.37691
    Li, T.J. & Carpenter, J.M. (2018) A taxonomic account of the genus Labus de Saussure, 1867 (Hymenoptera, Vespidae, Eumeninae) with descriptions of three new species. Journal of Hymenoptera Research, 65, 23–46. https://doi.org/10.3897/jhr.65.26976
    Li, T.J. & Chen, B. (2014a) Description of two new Chinese Subancistrocerus de Saussure (Hymenoptera, Vespidae, Eumeninae), with a key to the Chinese species. Journal of Hymenoptera Research, 39, 47–57. https://doi.org/10.3897/jhr.39.7736
    Li, T.J. & Chen, B. (2014b) The taxonomic accounts of the genus Symmorphus Wesmael (Hymenoptera, Vespidae, Eumeninae) from China, with descriptions of three new species. ZooKeys, 389, 9–26. https://doi.org/10.3897/zookeys.389.7045
    Li, T.J. & Chen, B. (2016a) The taxonomic study of the genus Apodynerus Giordani Soika (Hymenoptera: Vespidae: Eumeninae) from China, with descriptions of two new species. Entomotaxonomia, 38 (2), 143–155.
    Li, T.J. & Chen, B. (2016b) Two newly recorded genera Stenodyneriellus and Lissodynerus with three new species from China (Hymenoptera, Vespidae, Eumeninae). Journal of Hymenoptera Research, 49, 111–127. https://doi.org/10.3897/jhr.49.7487
    Lou, L., Zhang, Q.H., Chen, B. & Li, T.J. (2020) Additions to the knowledge of the genus Allorhynchium van der Vecht, 1963 from China (Hymenoptera, Vespidae, Eumeninae). Journal of Hymenoptera Research, 77, 119–137. https://doi.org/10.3897/jhr.77.52309
    Ma, Z.X., Chen, B. & Li, T.J. (2016) A taxonomic account of the genus Stenodynerus from China, with descriptions of five new species (Hymenoptera, Vespidae, Eumeninae). ZooKeys, 595, 17–48. https://doi.org/10.3897/zookeys.595.7734
    Madl, M. (2012) Notes on the genus Provespa Ashmead, 1903 (Insecta: Hymenoptera: Vespidae: Vespinae) based on the material of the Naturhistorisches Museum Wien (Austria). Annalen des Naturhistorischen Museums in Wien. Serie B für Botanik und Zoologie, 114, 27–35.
    Mahmood, K., Ullah, M., Aziz, A., Hasan, S.A. & Inayatullah, M. (2012) To the knowledge of Vespidae (Hymenoptera) of Pakistan. Zootaxa, 3318 (1), 26–50. https://doi.org/10.11646/zootaxa.3318.1.2
    Nguyen, L.T.P. (2015a) Potter wasps of the genus Eumenes Latreille, 1802 (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species and key to species. Zootaxa, 3974 (4), 564–572. https://doi.org/10.11646/zootaxa.3974.4.7
    Nguyen, L.T.P. (2015b) Taxonomic Notes on the Genus Delta de Saussure (Hymenoptera: Vespidae: Eumeninae) from Vietnam. Animal Systematics, Evolution and Diversity, 31 (2), 95–100. https://doi.org/10.5635/ased.2015.31.2.095
    Nguyen, L.T.P. (2015c) Taxonomic notes on the species of the genus Anterhynchium de Saussure, 1863 (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Zootaxa, 3915 (1), 132–138. https://doi.org/10.11646/zootaxa.3915.1.7
    Nguyen, L.T.P. (2016) Contribution to the taxonomy of the genus Coeleumenes van der Vecht, 1963 (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Zootaxa, 4121 (2), 175–180. https://doi.org/10.11646/zootaxa.4121.2.7
    Nguyen, L.T.P. & Carpenter, J.M. (2016) Review of the Polistes (Polistella) "Stenopolistes" species-group (Hymenoptera: Vespidae: Polistinae) from Vietnam, with description of a new species and key to species. Zootaxa, 4088 (4), 583–593. https://doi.org/10.11646/zootaxa.4088.4.9
    Nguyen, L.T.P. & Carpenter, J.M. (2019) Taxonomic notes on the paper wasps of the subgenus Polistes (Gyrostoma) (Hymenoptera, Vespidae, Polistinae) occurring in Vietnam, with description of a new species. Journal of Hymenoptera Research, 74, 51–64. https://doi.org/10.3897/jhr.74.47795
    Nguyen, L.T.P. & Carpenter, J.M. (2020) Potter wasps of the genus Labus (Hymenoptera, Vespidae, Eumeninae) from Vietnam, with description of two new species. Journal of Hymenoptera Research, 77, 139–150. https://doi.org/10.3897/jhr.77.52797
    Nguyen, L.T.P., Dang, H.T., Kojima, J.I. & Carpenter, J.M. (2014) An annotated distributional checklist of solitary wasps of the subfamily Eumeninae (Hymenoptera: Vespidae) of Vietnam. Entomologica Americana, 120 (1), 7–17. https://doi.org/10.1664/13-RA-010.1
    Nguyen, L.T.P. & Kojima, J.I. (2013) Distribution of social wasps in Vietnam (Hymenoptera: Vespidae). Tap Chi Sinh Hoc, 35 (3), 16–25. https://doi.org/10.15625/0866-7160/v35n3se.3832
    Nguyen, L.T.P. & Kojima, J.I. (2014) Distribution and nests of paper wasps of Polistes (Polistella) in northeastern Vietnam, with description of a new species (Hymenoptera, Vespidae, Polistinae). ZooKeys, 368, 45–63. https://doi.org/10.3897/zookeys.368.6426
    Nguyen, L.T.P., Kojima, J.I. & Saito, F. (2011) Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species. Zootaxa, 3060 (1), 1–30. https://doi.org/10.11646/zootaxa.3060.1.1
    Nguyen, L.T.P., Kojima, J.I., Saito, F. & Carpenter, J.M. (2006) Vespidae (Hymenoptera) of Vietnam 3: Synoptic key to Vietnamese species of the polistine genus Ropalidia, with notes on taxonomy and distribution. Entomological Science, 9, 93–107. https://doi.org/10.1111/j.1479-8298.2006.00157.x
    Nguyen, L.T.P., Ngat, T.T., Hoa, D.T. & Thai, M.V. (2019) Potter wasps of the genus Stenodyneriellus Giordani Soika (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Raffles Bulletin of Zoology, 67, 396–402. https://doi.org/10.26107/RBZ-2019-0031
    Nguyen, L.T.P., Nguyen, D.D. & Carpenter, J.M. (2016) Additions to the knowledge of the genus Phimenes (Hymenoptera: Vespidae: Eumeninae) from Vietnam. Animal Systematics, Evolution and Diversity, 32 (1), 21–27. https://doi.org/10.5635/ased.2016.32.1.021
    Nguyen, L.T.P., Saito, F., Kojima, J.I. & Carpenter, J.M. (2006) Vespidae of Vietnam (Insecta: Hymenoptera) 2. Taxonomic Notes on Vespinae. Zoological Science, 23, 95–104. https://doi.org/10.2108/zsj.23.95
    Nguyen, L.T.P., Vu, T.T., Daglio, A. & Wiśniowski, B. (2019) Taxonomic review and distribution of the genus Vespula Thomson, 1869 (Hymenoptera: Vespidae: Vespinae) from Vietnam. Zootaxa, 4691 (3), 279–285. https://doi.org/10.11646/zootaxa.4691.3.8
    Nidup, T., Gyeltshen, T., Kumar, P.G., Klein, W. & Dorji, P. (2016) New records of petiolate potter wasps (Hymenoptera: Vespidae: Eumeninae) from Bhutan. Journal of Threatened Taxa, 8 (12), 9467–9472. https://doi.org/10.11609/jott.2774.8.12.9467-9472
    Nidup, T., Klein, W. & Dorji, P. (2017) New records of hover wasps (Hymenoptera: Vespidae: Stenogastrinae) from Bhutan. Journal of Threatened Taxa, 9 (7), 10487–10489. https://doi.org/10.11609/jott.3204.9.7.10487-10489
    Nidup, T., Klein, W., Kumar, P.G. & Dorji, P. (2018) Additional reports of solitary potter wasps (Hymenoptera: Vespidae: Eumeninae) in Bhutan. Journal of Threatened Taxa, 10 (5), 11686–11696. https://doi.org/10.11609/jott.3888.10.5.11686-11696
    Nugroho, H., Kojima, J.I. & Carpenter, J.M. (2011) Checklist of vespid species (Insecta: Hymenoptera: Vespidae) occuring in Indonesia Archipelago. Treubia, 38, 71–186.
    Nugroho, H., Kojima, J.I. & Ubaidillah, R. (2013) Review of potter wasps with a petiolate metasoma excluding so-called "Zethinae" (Hymenoptera: Vespidae: Eumeninae) in the Lesser Sunda Islands of the Indonesian Archipelago. Zootaxa, 3608 (1), 1–25. https://doi.org/10.11646/zootaxa.3608.1.1
    Nugroho, H., Kojima, J.I. & Ubaidillah, R. (2014) Synonymy of the potter wasp genus Philippodynerus Gusenleitner (Hymenoptera, Vespidae, Eumeninae) with Apodynerus Giordani Soika, with taxonomic notes on Apodynerus species. Journal of Hymenoptera Research, 36, 131–151. https://doi.org/10.3897/JHR.36.6830
    Pannure, A., Belavadi, V.V. & Carpenter, J.M. (2018) Taxonomic notes on poorly known species of potter wasps (Vespidae: Eumeninae: Zethini) from India. Oriental Insects, 53 (3), 395–410. https://doi.org/10.1080/00305316.2018.1521345
    Pham, P.H. (2014) A checklist of Ropalidiini wasps (Hymenoptera: Vespidae; Polistinae) in Indochina. Archives of Biological Sciences, 66 (3), 1061–1074. https://doi.org/10.2298/ABS1403061P
    Pham, P.H. & Kumar, P.G. (2016) Taxonomic notes on the genus Rhynchium Spinola (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with a first checklist of the species worldwide. Ecologica Montenegrina, 7, 530–537.
    Pham, P.H. & Li, T.J. (2015) A first list of vespid wasps from Vietnam (Hymenoptera: Vespidae). Russian Entomological Journal, 24 (2), 133–144. https://doi.org/10.15298/rusentj.24.2.05
    Porporato, M., Manino, A., Laurino, D. & Demichelis, S. (2014) Vespa velutina Lepeletier (Hymenoptera: Vespidae): A first assessment two years after its arrival in Italy. Redia, XCVII, 189–194.
    Qasim, M., Khan, I.A., Rafi, M.A., Khan, M.R., Amin, M., Sheik, M.K., Rasul, G. & Shah, M. (2018) Wasps fauna of (Subfamilies: Eumeninae, Vespinae and Polistinae) from district Muzaffarabad, Azad Jammu and Kashmir, Pakistan. Journal of Animal Sciences, 2 (4), 26–30.
    Rafi, M.A., Carpenter, J.M., Qasim, M., Shehzad, A., Zia, A., Khan, M.R., Mastoi, M.I., Naz, F., Ilyas, M., Shah, M. & Bhatti, A.R. (2017) The vespid fauna of Pakistan. Zootaxa, 4362 (1), 1–28. https://doi.org/10.11646/zootaxa.4362.1.1
    Rauf, S.A., Rafi, M.A., Qasim, M., Sheik, K., Mehmood, K., Khan, A. & Rahman, A. (2018) Wasp fauna of subfamilies (Eumeninae, Polistinae and Vespinae) from Dir, Khyber Pakhtunkhwa, Pakistan. Journal of Animal Sciences, 2 (1), 1–7.
    Rasool, M., Zahid, M. & Shah, M. (2017) Solitary Wasps (Hymenoptera: Vespidae: Eumeninae) of Swat Pakistan, with two species newly reported from the country and three unidentified species. Journal of Entomology and Zoology Studies, 5 (2), 648–653.
    Saito, F. & Kojima, J.I. (2007) A taxonomic revision of the hover wasp genus Eustenogaster van der Vecht (Insecta: Hymenoptera; Vespidae, Stenogastrinae). Zootaxa, 1556 (1), 1–30. https://doi.org/10.11646/zootaxa.1556.1.1
    Saito, F., Nguyen, L.T.P., Carpenter, J.M. & Kojima, J.I. (2006) A new Eustenogaster species (Hymenoptera: Vespidae; Stenogastrinae), the first hover wasp known to overwinter on the nest. American Museum Novitates, 3534, 1–11.
    Siddiqui, J.A., Bodlah, I., Carpenter, J.M., Naeem, M., Ahmad, M. & Bodlah, M.A. (2015) Vespidae (Hymenoptera) of the Pothwar region of Punjab, Pakistan. Zootaxa, 3914 (5), 501–524. https://doi.org/10.11646/zootaxa.3914.5.1
    Smith-Pardo, A.H., Carpenter, J.M. & Kimsey, L. (2020) The Diversity of Hornets in the Genus Vespa (Hymenoptera: Vespidae; Vespinae), Their Importance and Interceptions in the United States. Insect Systematics and Diversity, 4 (3), 1–17. https://doi.org/10.1093/isd/ixaa006
    Sora, K., Gawas, S.M., Gupta, A. & Shaw, S.S. (2019) Checklist of vespid wasps (Hymenoptera: Vespidae) of Arunachal Pradesh, India, with a new distribution record of Vespula flaviceps (Smith). Journal of Biological Control, 33 (4), 336–342. https://doi.org/10.18311/jbc/2019/24148
    Srinivasan, G. & Kumar, P.G. (2009) Taxonomic studies of the hover wasps (Hymenoptera: Vespidae; Stenogastrinae) of Arunachal Pradesh, India. Records of the Zoological Survey of India, 109 (3), 37–40.
    Srinivasan, G. & Kumar, P.G. (2010) New records of potter wasps (Hymenoptera: Vespidae: Eumeninae) from Arunachal Pradesh, India: five genera and ten species. Journal of Threatened Taxa, 2 (12), 1313–1322. https://doi.org/10.11609/JoTT.o2468.1313–22.
    Starr, C.K. (1992) The social wasps (Hymenoptera: Vespidae) of Taiwan. Bulletin of National Museum of the Natural Science, 3, 93–138.
    Tan, J.L., Carpenter, J.M. & Achterberg, C.V. (2018a) An illustrated key to the genera of Eumeninae from China, with a checklist of species (Hymenoptera, vespidae). ZooKeys, 740, 109–149. https://doi.org/10.3897/zookeys.740.22654
    Tan, J.L., Carpenter, J.M. & Achterberg, C.V. (2018b) Most northern Oriental distribution of Zethus Fabricius (Hymenoptera, Vespidae, Eumeninae), with a new species from China. Journal of Hymenoptera Research, 62, 1–13. https://doi.org/10.3897/jhr.62.23196
    Tan, J.L., Achterberg, C.V., He, Y.F. & Carpenter, J.M. (2018) Northeast Asian Allorhynchium van der Vecht (Hymenoptera: Vespidae), with a key to Oriental species. Zootaxa, 4434 (1), 049–064. https://doi.org/10.11646/zootaxa.4434.1.3
    Tan, J.L., Achterberg, C.V. & Chen, X.X. (2014) Pictorial key to species of the genus Ropalidia GuérinMéneville, 1831 (Hymenoptera, Vespidae) from China, with description of one new species. ZooKeys, 391, 1–35. https://doi.org/10.3897/zookeys.391.6606
    Tan, J.L., Achterberg, C.V., Duan, M.J. & Chen, X.X. (2014) An illustrated key to the species of subgenus Gyrostoma Kirby, 1828 (Hymenoptera, Vespidae, Polistinae) from China, with discovery of Polistes (Gyrostoma) tenuispunctia Kim, 2001. Zootaxa, 3785 (3), 377–399. https://doi.org/10.11646/zootaxa.3785.3.3
    Tan, Q.Q., Tan, J.L., Zhang, R.N., Tian, X.X. & Jian, J.K. (2018) Discovery of the nest of the yellow jacket Vespula structor (Smith) (Hymenoptera, Vespidae) from China with description of its immature stages. Journal of Hymenoptera Research, 67, 103–120. https://doi.org/10.3897/jhr.67.30059
    Tenzin, K. & Katel, O. (2019) Diversity and abundance of bees and wasps (Hymenoptera: Aculeata) in north central Bhutan. Bhutan Journal of Natural Resources and Development, 6 (1), 12–18. https://doi.org/10.17102/cnr.2019.02
    Ueno, T. (2014) Establishment of the invasive hornet Vespa velutina (Hymenoptera: Vespidae) in Japan. International Journal of Chemical, Environmental & Biological Sciences, 2 (4), 220–222.
    van der Vecht, J. (1957) The Vespinae of the Indo-Malayan and Papuan areas (Hymenoptera, Vespidae). Zoologische Verhandelingen, 34, 1–83.
    van der Vecht, J. (1959) Notes on Oriental Vespinae, including some species from China and Japan (Hymenoptera, Vespidae). Zoologische Mededelingen, XXXVI (13), 205–232.
    van der Vecht, J. (1963) Studies on Indo-Australian and East-Asiatic Eumenidae (Hymenoptera, Vespoidea). Zoologische Verhandelingen Leiden, 60, 1–116.
    van der Vecht, J. (1966) The East-Asiatic and Indo-Australian species of Polybioides buysson and Parapolybia (Hym., Vespidae). Zoologische Verhandelingen, 82, 1–45.
    van der Vecht, J. (1979) Notes on Vespoidea from Nepal. Entomologische Berichten, 39, 27–30.
    Yamane, S. (1974) On the genus Vespa (Hymenoptera, Vespidae) from Nepal. Kontyû, 42 (1), 29–39.
    Yamane, S. & Yamane, S. (1979) Polistine wasps from Nepal (Hymenoptera, Vespidae). Insecta Matsumurana, New Series, 15, 1–37.
    Yildirim, E. (2012) The distribution and biogeography of Vespidae (Hymenoptera: Aculeata) in Turkey. Turkish Journal of Entomology, 36 (1), 23–42.

  2.