Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2022-06-20
Page range: 61-86
Abstract views: 482
PDF downloaded: 24

A review of the Mutillidae (Hymenoptera) of Azerbaijan

Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia.
Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia.
Institute of Bioresources of Nakhchivan Branch of National Academy of Sciences of Azerbaijan, Nakhchivan, AZ 7000, Azerbaijan.
Hymenoptera

Abstract

Fifty-eight species in 19 genera are recorded from Azerbaijan. A new synonymy is proposed for Mutilla sinuata Olivier, 1811 (=M. binotata Radoszkowski, 1879, syn. nov.; M. araratica Radoszkowski, 1890, syn. nov.) and Smicromyrme atrithorax (André, 1902) (=S. kurdus Skorikov, 1935, syn. nov.). The lectotypes of seven species: Agama caucasica Radoszkowski, 1885, ♂; Ephutomma radoszkovskii Skorikov, 1935, ♂; Mutilla armeniaca Kolenati, 1846, ♀; M. elongata Radoszkowski, 1885, ♂; M. binotata Radoszkowski 1879, ♀; M. araratica Radoszkowski 1890, ♀; and M. villosa Klug 1829, ♀ are designated. Fourteen species are recorded for the first time from Azerbaijan: Dasylabris (Inbaltilla) regalis (Fabricius, 1793); D. (I.) popovi Skorikov, 1935; D. (I.) shelkovnikovi Lelej, 1985; Dentilla erronea (André, 1900); Myrmilla (M.) calva (Villers, 1789); M. (Pseudomutilla) glabrata (Fabricius, 1775); M. (P.) vutshetitshi Skorikov, 1927; Physetopoda daghestanica (Radoszkowski, 1885); Pseudophotopsis armeniaca (Skorikov, 1935); Smicromyrme (Astomyrme) ausonius Invrea 1950; S. (Eremotilla) novaki Invrea, 1954; S. (Smicromyrme) ruficollis ruficollis (Fabricius, 1793); S. (S.) rufipes (Fabricius, 1787); and S. (S.) tristis Lelej, 1984. Four species are excluded from the list of Azerbaijan fauna. An updated key to three Palaearctic species of Macromyrme Lelej, 1984 is given and new combination and resurrected status are proposed for Macromyrme villosa (Klug, 1829). A cluster analysis of faunal similarities among Azerbaijan, Armenia, Georgia, Turkey and three regions of Russia (Crimea, North Caucasus, south of European part) for Mutillidae produce two major clusters (index similarity 0.4): Azerbaijan, Armenia, Turkey and Georgia (bootstrap probability 45 %), and three regions of Russia (bootstrap probability 87 %). This demonstrates the important boundary between two large biogeographical subregions of Palaearctic: Mediterranean and Euro-Siberian. The Azerbaijan fauna, especially Nakhchivan Autonomous Republic, is most closely related with the Armenian fauna.

 

References

  1. Aliev, A. & Lelej, A.S. (1999[“1998”]) The Aculeate Hymenoptera in the collection of the Institute of Zoology of the Academy of Sciences of Azerbaijan, Baku. Part 1. The mutillid wasp (Mutillidae). Russian Entomological Journal, 7 (1–2), 43–50.
    André, E. (1893) Pour servir à la connaissance des mutilles paléarctiques et description de quelques espèces nouvelles. Mèmoirs de la Société Zoologique de France, 6, 286–296.
    André, E. (1896) Notes pour servir à la connaissance de mutilles paléarctiques et description de quelques espèces nouvelles. Deuxieme partie. Mémoires de la Société Zoologique de France, 9, 261–277.
    André, E. (1899–1902) Species de Hyménoptères d’Europe & d’Algérie. Tome Huitième. Les Mutillides. Vve Dubosclard and A. Hermann, Paris, 480 pp. + 15 pls. [pp. 1–64, pls. I–VIII (1899b) + 65–144 (1900) + 145–304 (1901) + 305–384 (1902a) + 385–478 (1902a) + pls. IX–XV (dates?)]
    André, E. (1902b [“1903”]) Fam. Mutillidae. In: Wytsman, P., Genera Insectorum. Fasc. 11. L. Desmet-Verteneuil, Bruxelles, pp. 1–77, 3 pls.
    André, E. (1910) Revision monographique des Mutillides de l’Egypte. Mémoires de la Société Entomologique d’Egypte, 1 (2), 1–94, 3 pls.
    Ashmead, W.H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45–60.
    Ashmead, W.H. (1900–1904) Classification of the fossorial, predaceous and parasitic wasps, or the superfamily Vespoidea. The Canadian Entomologist, 32, 34, 35 & 36, 145–155 + 185–188 + 295–296 (1900), 79–88 + 131–137 + 163–166 + 203–210 + 219–231 + 268–273 + 287–291 (1902), 3–8 + 39–44 + 95–107 + 155–158 + 199–205 + 303–310 + 323–332 (1903) & 5–9 (1904). https://doi.org/10.4039/Ent32145-5 https://doi.org/10.4039/Ent3479-4 https://doi.org/10.4039/Ent353-1
    Bischoff, H. (1920–1921) Monographie der Mutilliden Afrikas. Archiv für Naturgeschichte, 86A (1–3 & 4–5), 1–480 (1920) & 481–830 (1921).
    Brothers, D.J. & Lelej, A.S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1–97. https://doi.org/10.3897/jhr.60.20091
    Costa, A. (1858–1860) Famiglia de’Mutillidei; Mutillidea. In: Fauna del Regno di Napoli ossia Enumerazione di Tutti gli Animali che abitano le Diverse Regioni di questo Regno e le Acque che le bagnano e Descrizione de’ Nuovi o poco esattemente conosciuti con Figure ricevute da Originali viventi e dipinte al Naturale. Imenotteri Aculeati. Gaetano Sautto, Napoli, pp. 1–16 (1858) + pp. 17–28, tabs. 21–24 (1859) + pp. 29–36, tab. 25 (1860).
    Dalla Torre, C.G. de. (1897) Catalogus Hymenopterorum hucusque Descriptorum Systematicus et Synonimicus. Vol. VIII. Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], [4] + viii + 750 pp.
    El-Torkey, A.M., Oshaibah, A.D.A., Salem, M.M.H., Hosni, M.T., Abdel-Rhman, A.-R. G. & Soliman, A.M. (2011) Review of the Palaearctic species of Macromyrme Lelej 1984 (Hymenoptera: Mutillidae) with description of a new species from Egypt. Zootaxa, 2863 (1), 63–68. https://doi.org/10.11646/zootaxa.2863.1.4
    Fabricius, J.C. (1775) Systema entomologiae, systens Insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. In Officina Libraria Kortii, Flensburgi et Lipsiae, 832 pp. https://doi.org/10.5962/bhl.title.36510
    Fabricius, J.Ch. (1787) Mantissa insectorum sistens eorum species nuper detecta, adjectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tome 1. Impensis C. G. Proft, Hafniae [Copenhagen], 348 pp. https://doi.org/10.5962/bhl.title.36471
    Fabricius, J.C. (1793) Entomologca Systematica emendata et aucta, secundum Classes, Ordines, Genera, Species adjectis Synonymis, Locis, Observationibus, Descriptionibus. Tome 2. Christ. Gottl. Proft, Hafniae [Copenhagen], viii + 519 pp.
    Fateryga, A.V., Proshchalykin, M.Yu., Aliyev, Kh.A. & Maharramov, M.M. (2019) To the knowledge of eumenine wasps (Hymenoptera: Vespidae: Eumeninae) of Nakhchivan Autonomous Republic of Azerbaijan. Far Eastern Entomologist, 379, 25–32. https://doi.org/10.25221/fee.379.2
    Fateryga, A.V., Proshchalykin, M.Yu., Maharramov, M.M. & Astafurova, Yu. V. (2021) New records of solitary vespid wasps (Hymenoptera: Vespidae: Masarinae and Eumeninae s. l.) from the Nakhchivan Autonomous Republic of Azerbaijan. Zootaxa, 5027 (1), 36–60. https://doi.org/10.11646/zootaxa.5027.1.2
    Hammer, Ø., Harper, D.A.T. & Ryan, P.D. (2006) Paleontological statistics. Version 1.57. Available from: https://past.en.lo4d.com/download (accessed 23 March 2022)
    Invrea, F. (1950) Mutillidi nuovi o notevoli del Bacino Mediterraneo (Hymenoptera, Mutillidae). Memorie della Società Entomologica Italiana, 29, 19–27.
    Invrea, F. (1954) Mutillidi della Dalmazia e del litorale Montenegrino. Nota prima. Memorie della Società Entomologica Italiana, 32, 143–159.
    Invrea, F. (1963) Una nuova Myrmosa delle Serbia diagnosi preventiva. Bolletino della Società Entomologica Italiana, 93, 67.
    Invrea, F. (1965) Studi sugli Apterogynidi e Mutillidi della Palestina (Hymenoptera). Memorie della Società Entomologica Italiana, 44, 53–93.
    Klug, J.C.F. (1829) Symbolae Physicae seu Icones et Descriptiones Insectorum quae ex Itinere per Africam Borealem et Asiam Occidentalem Friderici Guilelmi Hemprich et Cristiani Godofredi Ehrenberg Studio Novae aut Illustratae redierunt. III, Insectorum. Hymenoptera, Decas Prima & Symbolae Physicae seu Icones et Descriptiones Insectorum quae ex Itineribus per Libyam, Aegyptum, Nubiam, Dongalam, Syriam Arabiam et Habbesiniam Publico Institutis sumptu Friderici Guilelmi Hemprich et Cristiani Godofredi Ehrenberg Medicinae et Chirurgiae Doctorum, Studio Annis MDCCCXX–MDCCCXXV redierunt. III. Insectorum. Hymenoptera, Decas Prima. Ex Officina Academica, venditur a Mittlero, Berolini, 17 pp. & 10 pls.
    Kolenati, F.A. (1846) Meletemata Entomologica V. Insecta Caucasi. Coleoptera, Dermaptera, Lepidoptera, Neuroptera, Mutillidae, Aphaniptera, Anoplura. Academia Sciences, Petropolis, iii + 170 pp.
    Kuhlmann, M. & Proshchalykin, M. Yu. (2016) The bees of the genus Colletes Latreille Hymenoptera: Colletidae) of the Caucasus region. Zootaxa, 4161 (3), 367­–385. http://doi.org/10.11646/zootaxa.4161.3.5
    Latreille, P.A. (1810) Considérations Générales sur l’Ordre Naturel des Animaux composant les Classes des Crustacés, des Arachnides et des Insectes; avec un Tableau Méthodique de leurs Genres, disposés en Familles. F. Schoell, Paris, [2] + 444 pp.
    Legendre, L. & Legendre, P. (1998) Numerical ecology. Developments in enviromental modeling, 2nd edn. Elsevier Scientific Publishing Co., Amsterdam, Oxford & New York, 853 pp.
    Lelej, A.S. (1976) Wasps of the genus Dasylabris Rad. (Hymenoptera, Mutillidae) of the fauna of the USSR. Entomologicheskoe Obozrenie, 55 (1), 189–204. [in Russian, English translation: (1976) Entomological Review, 55 (1), 136–146]
    Lelej, A.S. (1980) The genus Pseudophotopsis André, 1896 (Hymenoptera, Mutillidae) from the USSR and neighbouring countries. Entomologicheskoe Obozrenie, 59 (3), 634–649. [in Russian, English translation: (1982) Entomological Review, 59 (3), 112–126]
    Lelej, A.S. (1984a) The velvet ants of the genus Paramyrme (Hymenoptera, Mutillidae). In: Lehr, P.A. (Ed.), Systematic of the insects of the Far East. DVNTS AN SSSR, Vladivostok, pp. 101–115. [in Russian]
    Lelej, A.S. (1984b) New and little-known species of velvet ants of the genus Smicromyrme Thomson (Hymenoptera, Mutillidae). In: Savtshenko, E.N. (Ed.), Taxonomy and zoogeography of insects. Naukova Dumka, Kiev, pp. 7–89. [in Russian]
    Lelej, A.S. (1984c) New Palaearctic genera and species of velvet ants (Hymenoptera, Mutillidae). Entomologicheskoe Obozrenie, 63 (4), 826–829. [in Russian, English translation: (1985) Entomological Review, 64 (1), 101–104]
    Lelej, A.S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]
    Lelej, A.S. (1992) Two interesting species of velvet ants from China (Hymenoptera, Mutillidae). Entomofauna, 13 (19), 281–288.
    Lelej, A.S. (1996) To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species. Far Eastern Entomologist, 30, 1–24.
    Lelej, A.S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic Region. Dalnauka, Vladivostok, 172 pp.
    Lelej, A.S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp.
    Lelej, A.S. (2007) Biogeography of mutillid wasps (Hymenoptera, Mutillidae). In: Rasnitsyn, A.P. & Gokhman, V.E. (Eds.), Studies on Hymenopterous Insects. Collection of Scientific Papers. KMK Scientific Press Ltd., Moscow, pp. 82–111. [in Russian]
    Lelej, A.S. (2017) Family Mutillidae—Velvet ants. In: Lelej, A.S., Proshchalykin, M. Yu. & Loktionov, V.M. (Eds.), Annotated catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute Russian Academy of Sciences. Supplement 6. Zoological Institute RAS, St Petersburg, pp. 152–158.
    Lelej, A.S. (2020) To the knowledge of velvet ants of the genera Artiotilla Invrea, Radoszkowskitilla Lelej and Taiwanomyrme Tsuneki (Hymenoptera: Mutillidae). Far Eastern Entomologist, 440, 1–10. https://doi.org/10.25221/fee.410.1
    Lelej, A.S., Gharali, B. & Lotfalizadeh, H. (2008) New records of velvet ants (Hymenoptera: Mutilidae) from the West of Iran. Far Eastern Entomologist, 191, 1–7.
    Lelej, A.S. & Kabakov, O.N. (1980) On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan. Entomologicheskoe Obozrenie, 59 (1), 181–196. [in Russian, English translation: (1981) Entomological Review, 59, 141–156]
    Lelej, A.S. & Yildirim, E. (2009) A review of the Mutillidae (Hymenoptera) of Turkey. Zootaxa, 2160 (1), 1–28. https://doi.org/10.11646/zootaxa.2160.1.1
    Lepeletier de Saint-Fargeau, A. (1845) Histoire Naturelle des Insectes. Hyménoptères. Tome Troisième. Roret, Paris, [4] + 646 pp.
    Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. Editio Decima Reformata. Laurentii Salvii, Holmiae [Stockholm], [4] + 823 + [5] pp. https://doi.org/10.5962/bhl.title.542 
    Linnaeus, C. (1767) In: Systema Naturae per Regna Tria Naturae secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. Pars II. Editio Duodecima Reformata. Laurentii Salvii, Holmiae [Stockholm], pp. [2] + 533–1327 + [37].
    Lo Cascio, P., Romano, M. & Grita, F. (2012) New species and new records of mutillid wasps from the Socotra Archipelago (Hymenoptera: Mutillidae). Acta Entomologica Musei Nationalis Pragae, 52 (Supplement), 525–544.
    Lucas, H. (1849) Exploration Scientifique de l’Algérie pendant des Années 1840, 1841, 1842. Sciences Physiques, Zoologie. III. Histoire Naturelle des Animaux Articulés, Troisieme Partie, Insectes & Zoologie. IV. Histoire Naturelle des Animaux Articulés, Atlas. Imprimerie Nationale, Paris, [6] + 527 pp. & 122 pls.
    Maharramov, M.M., Mokrousov, M.V. & Proshchalykin, M.Yu. (2020) New distributional records of the family Sphecidae (Hymenoptera) in Azerbaijan. Caucasian Entomological Bulletin, 16 (1), 43–47. https://doi.org/10.23885/181433262020161-4347
    Mokrousov, M.V., Proshchalykin, M.Yu., Aliyev, Kh.A. & Maharramov, M.M. (2019) To the knowledge of digger wasps (Hymenoptera: Crabronidae) of Nakhchivan Autonomous Republic of Azerbaijan. Far Eastern Entomologist, 394, 1–24. https://doi.org/10.25221/fee.394.1
    Morawitz, F. (1893) Catalog der von D. Glasunov in Turkestan gesammelten Hymenoptera Fossoria. Horae Societatis Entomologicae Rossicae, 27 (3–4), 391–428.
    Nagy, C.G. (1972) Two new species of Mutillidae (Hymenoptera) from Asia Minor and description of female Mutilla erronea. Bulletin of the Iraq Natural History Museum, 5 (2), 4–9.
    Nonveiller, G. (1959). Une nouvelle espece du groupe Smicromyrme perisii S.-R. Zastita Bilja, 54, 105–116.
    Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societé Entomologique de France, New Series, 31 (4), 349–368.
    Olivier, [A.G.] (1811) Mutille. Encyclopédie Methodique. Histoire Naturelle. Insectes, 8, 49–66.
    Olson, D.M., Dinerstein, E., Wikramanayake, E.D., Burgess, N.D., Powell, G.V.N., Underwood, E.C., D’Amico, J.A., Itoua, I., Strand, H.E., Morrison, J.C., Loucks, C.J., Allnutt, T.F., Ricketts, T.H., Kura, Y., Lamoreux, J.F., Wettengel, W.W., Hedao, P. & Kassem, K.R. (2001) Terrestrial ecoregions of the world: A new map of life on Earth. BioScience, 51 (11), 933–938. https://doi.org/10.1641/0006-3568(2001)051[0933:TEOTWA]2.0.CO;2
    Ovtchinnikov, S.V. (2002) Materials to the fauna of velvet ants (Hymenoptera, Mutillidae) of Kyrghizia. In: Zonstein, S.L. (Ed.), Entomological Investigations in Kyrgystan. Vol. 22. Kyrghyz Society of Entomology, Bishkek, pp. 79–104. [in Russian]
    Özbek, H., Yıldırım, E. & Osten, T. (1999) A contribution to the knowledge of the fauna of the families Myrmosidae and Mutillidae (Hymenoptera, Aculeate) in Türkiye. Türkiye Entomoloji Dergisi, 23 (1), 15–26.
    Pagliano, G., Brothers, D.J., Cambra, R., Lelej, A.S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P.L., Williams, K.A. & Romano, M. (2020 [“2018”]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torino, 36 (1–2), 5–425.
    Pate, V.S.L. (1947 [“1946”]) A minute on Paramyrmosa Saussure (Hymenoptera, Tiphiidae). The Canadian Entomologist, 78 (11–12), 196–197. https://doi.org/10.4039/Ent78196-11
    Pallas, P.S. (1773) Reisen durch verschiedene Provinzen des Russischen Reichs. Zweiter Theil. Zweites Buch vom Jahr 1771. Kayserlichen Academie der Wissenschaften, St. Petersburg, [12] + 744 pp.
    Petagna, V. (1787) Specimen Insectorum Ulterioris Calabriae. Perger, Neapoli, 46 pp., 1 pl.
    Petersen, B. (1988) The Palaearctic Mutillidae of I.C. Fabricius and some related material (Insecta, Hymenoptera, Aculeata). Steenstrupia, 14 (6), 129–224.
    Radoszkowski, O. (1865) Les mutilles russes. Bulletin de la Société Impériale des Naturalistes de Moscou, 38 (2), 422–467, tabs. 7–9.
    Radoszkowski, O. (1877) Chrysidiformes, Mutillidae et Sphegidae. In: Puteshestvie v Turkestan A.P. Fedtshenko [Voyage au Turkestan d’Alexis Fedtschenko], Sankt-Petersburg, Series 14, 2 (5), pp. 1–87, 3 pls. [in Russian]
    Radoszkowski, O. (1879) Les Chrysides et Sphegides du Caucase. Horae Societatis Entomologicae Rossicae, 15 (2–3), 140–156.
    Radoszkowski, O. (1885) Revision des armures copulatrices des mâles de la famille de Mutillides. Horae Societatis Entomologicae Rossicae, 19 (1–2), 3–49, 9 pls.
    Radoszkowski, O. (1888) Etudes hyménoptèrologique. Revision des armures copulatrices des mâles. Horae Societatis Entomologicae Rossicae, 22 (3–4), 315–337, tabs. 12 + 14 + 15.
    Radoszkowski, O. (1890) Hyménoptères recoltes sur le mont Ararat. Horae Societatis Entomologicae Rossicae, 24 (3–4), 502–510.
    Reitter, E. (1890) Eine neue mit Athous verwandte Elateriden. Gattung aus russisch Armenien. Entomologische-Nachrichten, 16, 247–248.
    Reitter, E. (1905) Neun neue Coleopteren aus der palaearktischen Fauna. Wiener Entomologische Zeitung, 24 (5–6), 201–206.
    Rossi, P. (1792–1794) Mantissa insectorum exhibens species nuper in Etruria collectas adjectis Faunae Etruscae illustrationibus ac emendationibus, Pisa, 1 & 2, 11–148 (1792) & 1–154 pp., 8 pls. (1794).
    Saussure, H.de. (1880) Scolidae. In: Puteshestvie v Turkestan A.P. Fedtshenko [Voyage au Turkestan d‘Alexis Fedtschenko], Sankt-Petersburg, Series 16, 2 (5), pp. 1–46, pls. 1–2. [in Russian]
    Schuster, R.M. (1949) Contribution towards a monograph of the Mutillidae of the Neotropical region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, New Series, 29 (3–4), 59–140.
    Semenov Tian-Shanskij, A. (1935) Les limites et les subdivisions zoogeographiques de la région paléarctique pour les animaux terrestres, basées sur la distribution géographique des insectes Coléoptères. (Avec une carte géographique). Travaux del’lnstitut Zoologique de I’Academie des Sciences de I’URSS, 2–3, 397–410, 1 map.
    Sichel, J. & Radoszkowski, O. (1869–1870) Essai d’une monographie des Mutilles de l’ancien continent. Horae Societatis Entomologicae Rossicae, 6 (3 & 4), 139–172 + pls. 6–8 (1869) & 6 (4), 173–309, pls. 9–11 (1870).
    Skorikov, A.S. (1927) Sur la sousfam. des Myrmillini (Myrmosidae, Hymenoptera) dans la region palearctique. Ezhegodnik Zoologicheskogo Muzeya Akademii Nauk SSSR, 28 (1), 33–47. [in Russian]
    Skorikov, A.S. (1935) Zur Mutilliden-Fauna Zentralasiens. Trudy Tadzhikskoi Basy Akademii Nauk SSSR, 5, 257–349, 7 pls. [in Russian]
    Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Trustees of the British Museum, London, [4] + 206 pp., pls. A + 1–5. https://doi.org/10.5962/bhl.title.50533 
    Smith, F. (1873) Descriptions of aculeate Hymenoptera of Japan, collected by Mr. George Lewis at Nagasaki and Hiogo. Transactions of the Entomological Society of London, 1873 (2), 181–206. https://doi.org/10.1111/j.1365-2311.1873.tb00641.x 
    Suárez, F.J. (1959) Especies nuevas o poco conocidas de la fauna Mediterránea. II nota. Eos, Revista Española de Entomología, Madrid, 35 (3), 283–300.
    Suárez, F.J. (1975) Especies nuevas o poco conocidas de la fauna Mediterránea. IV. Tres nuevas Smicromyrme de la zona oriental (Hymenoptera, Mutillidae). Archivos del Instituto de Aclimatación, 20, 155–171.
    Thomson, C.G. (1870) Öfversigt af Sveriges rofsteklar. Opuscula Entomologica, 1 (2), 202–251.
    Turner, R.E. (1911) New Hymenoptera from Ceylon. Mutillidae and Scoliidae. Spolia Zeylanica, 7 (27), 141–154.
    Villers, C. de (1789) Caroli Linnaei entomologia, faunae Suecicae descriptionibus aucta; etc. Tomus tertius. s.n., Lugduni [Lyon], xxiv + 657 pp., pls. 7–10.
    Wesmael, M. (1851) Revue critique des Hyménoptères fouisseurs de Belgique. Bulletin de l’Académie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 18 (2), 362–384, 451–493.
    Williams, K.A., Lelej, A.S., Okayasu, J., Borkent, C.J., Malee, R., Thoawan, K. & Thaochan, N. (2019) The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia. Zootaxa, 4602 (1), 1–69. https://doi.org/10.11646/zootaxa.4602.1.1