Abstract
A new species of Filispirifer is described from the Lower Devonian Assa Formation of the Dra Valley (Morocco). Filispirifer hamadae n. sp. differs from the other taxa of Filispirifer in having a smaller shell, fewer costae, a lower ventral interarea, a smaller ventral process, and a less conspicuous dorsal adductor field. Due to their micro-ornamentation, the capillate Rhenish delthyridoid genera, Filispirifer and Mauispirifer, are excluded from the fimbriate family Acrospiriferidae and put together in the new family Filispiriferidae. The evolutionary lineage of Filispirifer and its palaeobiogeographic occurrence are briefly described.References
Allan, R.S. (1947) A revision of the Brachiopoda of the Lower Devonian of Reefton, New Zealand. Journal of Paleontology, 21 (5), 436–452.
Becker, R.T., Jansen, U., Plodowski, G., Schindler, E., Aboussalam, Z.S. & Weddige, K. (2004) Devonian litho- and biostratigraphy of the Dra Valley area: An overview. In: El Hassani, A. (Ed.), Devonian neritic-pelagic correlation and events in the Dra Valley (Western Anti-Atlas, Morocco). International Meeting on Stratigraphy, Rabat, March 1–10, 2004. Documents de l’Institut Scientifique, 19, 3–18.
Béclard, F. (1887) Les fossiles coblenziens de St. Michel près de Saint Hubert. Bulletin de la Société Belge de Géologie de Paléontologie et d'Hydrologie, 1, 60–97.
Boucot, A.J. (1962) The Eospiriferidae. Paleontology, 5 (4), 97–104.
Bultynck, P. & Walliser, O. (2000) Devonian Boundaries in the Moroccan Anti-Atlas. In: Bultynck, P. (Ed.), Subcommission on Devonian Stratigraphy – Recognition of Devonian Series and stage boundaries in geological areas. Courier Forschungsinstitut Senckenberg, 225, 211–226.
Carls, P. (1987) Ein Vorschlag zur biostratigraphischen Redefinition der Grenze Gedinnium/Siegenium und benachbarter Unter-Stufen. 1. Teil: Stratigraphische Argumente und Korrelation. Courier Forschungsinstitut Senckenberg, 92, 77–121.
Carls, P. & Valenzuela-Ríos, J.I. (1998) The ancestry of the Rhenish Middle Siegenian brachiopod fauna in the Iberian Chains and its palaeozoogeography (Early Devonian). Revista Española de Paleontología, no extraordinario, Homenaje al Prof. Gonzalo Vidal, 123–142.
Carter, J.L., Johnson, J.G., Gourvennec, R. & Hou Hongfei (1994) A revised classification of the spiriferid brachiopods. Annals of the Carnegie Museum, 63 (4), 327–374.
Conrad, T.A. (1840) Third annual report on the Paleontological Department of the Survey. New York Geological Survey, Annual Report 4, 199–207.
Drot, J. (1964) Rhynchonelloidea et Spiriferoidea siluro-dévoniens du Maroc pré-saharien. Notes et Mémoires du Service Géologique du Maroc, 178, 1–288.
Giebel, C.G. (1858) Die Silurische Fauna des Unterharzes nach Herrn C. Bischoff’s Sammlung. Abhandlungen des Naturwissenschaftlichen Vereines der Provinzen Sachsen und Thüringen, 1, 25–70.
Gourvennec, R. (1989) Brachiopodes Spiriferida du Dévonien inférieur du Massif Armoricain. Systématique, paléobiologie, évolution, biostratigraphie. Biostratigraphie du Paléozoïque, 9, 1–281.
Helmbrecht, W. & Wedekind, R. (1923) Versuch einer biostratigraphischen Gliederung der Siegener Schichten auf Grund von Rensselaerien und Spiriferen. Glückauf, 41, 949–953.
Hollard, H. (1963) Un tableau stratigraphique du Dévonien du Sud de l’Anti-Atlas. Notes du Service Géologique du Maroc, 23, 105–109.
Hollard, H. (1981a) Principaux charactères des formations dévoniennes de l’Anti-Atlas. Notes du Service Géologique du Maroc, 42, 15–22.
Hollard, H. (1981b) Tableaux de corrélations du Silurien et du Dévonien de l’Anti-Atlas. Notes du Service Géologique du Maroc, 42, 23.
Jahnke, H. (1971) Fauna und Alter der Erbslochgrauwacke (Brachiopoden und Trilobiten, Unter-Devon, Rheinisches Schiefergebirge und Harz). Göttinger Arbeiten zur Geologie und Paläontologie, 9, 1–105.
Jansen, U. (1999) Brachiopod fauna and age of the Assa Formation (Early Devonian, Dra Plains, southern Anti-Atlas, Moroccan Pre-Sahara). Senckenbergiana lethaea, 79 (1), 191–207.
Jansen, U. (2000) The Early Devonian genus Acrospirifer. A taxonomic odyssey. The Millenium Brachiopod Congress, Abstracts; London.
Jansen, U. (2001a) Morphologie, Taxonomie und Phylogenie unter-devonischer Brachiopoden aus der Dra-Ebene (Marokko) und dem Rheinischen Schiefergebirge (Deutschland). Abhandlungen der senckenbergischen naturforschenden Gesellschaft, 554, 1–389.
Jansen, U. (2001b) On the genus Acrospirifer Helmbrecht et Wedekind, 1923 (Brachiopoda, Lower Devonian). Contributions to Lower Devonian brachiopods from the Rheinisches Schiefergebirge and adjacent areas, 1. In J. Fryda, R.B. Blodgett & M. Mergl (Eds.), Havlíček Volume. Journal of the Czech Geological Society, 46 (3–4), 131–144.
Jansen, U., Becker, G., Plodowski, G., Schindler, E., Vogel, O. & Weddige, K. (2004a) The Emsian to Eifelian near Foum Zguid (NE Dra Valley, Morocco). In El Hassani, A. (Ed.), Devonian neritic-pelagic correlation and events in the Dra Valley (Western Anti-Atlas, Morocco). International Meeting on Stratigraphy, Rabat, March 1–10, 2004. Documents de l’Institut Scientifique, 19, 19–28.
Jansen, U., Plodowski, G., Schindler, E. & Weddige, K. (2004b) The Pragian at Assa. In El Hassani, A., (Ed.), Devonian neritic-pelagic correlation and events in the Dra Valley (Western Anti-Atlas, Morocco). International Meeting on Stratigraphy, Rabat, March 1–10, 2004. Documents de l’Institut Scientifique, 19, 64–68.
Jansen, U., Lazreq, N., Plodowski, G., Schemm-Gregory, M., Schindler, E. & Weddige, K. (2007) Neritic-pelagic correlation in the Lower and basal Middle Devonian of the Dra Valley (S Anti-Atlas, Moroccan Pre-Sahara). In Becker, R.T. & Kirchgasser, W. (Eds.), Devonian Events and Correlations. Geological Society, Special Publications, 278, 9–37.
Johnson, J.G. & Hou Hongfei (2006) Delthyridoidea. In Kaesler, R.L. (Ed.), Treatise on Invertebrate Paleontology, Part H, Brachiopoda, 5 (revised), 1825–1847, Lawrence, Kansas (Geological Society of America and University of Kansas).
Phillips, J. (1841) Figures and description of the Palaeozoic fossils of Cornwall, Devon and West Somerset. Longman & Co London, xii+231 pp.
Racheboeuf, P.R. (1976) Chonetacea (brachiopodes) du Dévonien inférieur du Bassin de Laval (Massif Armoricain). Palaeontographica Abteilung A, 152 (1–3), 14–89.
Schemm-Gregory, M. (2007) The Lower Devonian brachiopod genus Acrospirifer Helmbrecht and Wedekind, 1923 ― globally distributed or highly endemic? Subcommission on Devonian Stratigraphy and IGCP 499 Devonian Land Sea Interaction: Devonian Global Change: compelling changes in the Devonian world, highlighting new findings in the terrestrial and marine biomes: fish, invertebrates, plants, terrestrial vertebrates, global warming, mass extinction, bolide strikes, and global correlation. Program and Abstracts; Eureka, Nevada, 9–17 September 2007, 78–79.
Schemm-Gregory, M. (in press) A new terebratulid brachiopod species from the Siegenian (middle Lower Devonian) of the Dra Valley, Morocco. Palaeontology.
Schemm-Gregory, M. (in review) A new delthyridoid spiriferid genus from the Dra Valley, Morocco, and its phylogenetic affinities (Brachiopoda, Lower Devonian). Paläontologische Zeitschrift.
Steininger, J. (1853) Geognostische Beschreibung der Eifel. Lintz’sche Buchhandlung, Trier, 144 pp.
Termier, H. & Termier, G. (1949) Essai sur l’évolution des Spiriféridés: Maroc. Service Géologique. Division de Mines et Géologique, Notes et Mémoires, 74, 85–1123.
Termier, G. & Termier, H. (1950) Paléontologie marocaine. Tome II: Invertébrés de l’ère primaire, Bryozoaires et brachiopodes. Actualités Scientifiques et Industrielles, 1093, fascicule II, 1–253.
Vandercammen, A. (1963) Spiriferidae du Dévonien de la Belgique. Mémoires Institut Royal des Sciences Naturelles de Belgique, 150, 1–170.
de Verneuil, E. (1850) Tableau des fossiles du terrain dévonien du département de la Sarthe. Bulletin de la Société Géologique de France, 7 (série 2), 778–787.
Waagen, W.H. (1883) Salt Range fossils, Part 4 (2) Brachiopoda. Palaeontologica Indica, Memoires, 13, 329–390.