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Published: 2022-09-21
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The plagiatus species group of the genus Geodromicus Redtenbacher, 1857 (Insecta: Coleoptera: Staphylinidae: Omaliinae), and additional data on some species from Japan

Daugavpils University, Institute of Life Sciences and Technologies, Coleopterological Research Centre, Vienibas 13, Daugavpils, LV-5401, Latvia.
Coleoptera Palaearctic Europe Caucasus Middle Asia Siberia Japan Geodromicus taxonomy species groups synonymy lectotype designation distribution new records

Abstract

The plagiatus group of the genus Geodromicus Redtenbacher, 1857 includes eight Palaearctic species: the widespread G. plagiatus (Fabricius, 1798), the European G. corsicus Fagel, 1958, G. nigrita (P.W.J. Müller, 1821), G. pedemontanus Fagel, 1958, and G. suturalis (Lacordaire, 1835), and the East Palaearctic G. magnus Shavrin, 2020, G. paramerus Shavrin 2000, and G. shii Cheng & Peng, 2020 (in Cheng et al. 2020). The species group is reviewed, defined, and briefly discussed. The species are illustrated and a key is provided. Two additional species groups are established for four species from Japan with a strongly sclerotized flagellum: the aokii group, containing G. aokii (Watanabe, 1972) and G. sprius (Watanabe, 1990), and the sinuatus group containing G. sinuatus (Sharp, 1889) and G. suensoni Bernhauer, 1936. Both groups are defined, briefly discussed, and a key to each group is given. Several new synonymies are established: G. plagiatus = G. (s.str.) klimai A. Bordoni, 1984 syn. n., = G. (s.str.) schoenmanni A. Bordoni, 1984 syn. n.; G. nigrita = G. nigrita lohsei Jarrige, 1963 syn. n., = G. niveus Bernhauer, 1941 syn. n.; G. sinuatus = G. watanabei Hayashi, 2018 syn. n.; G. suensoni = G. takabai Hayashi, 2018 syn. n. Lectotypes of Staphylinus plagiatus Fabricius, 1798 and Anthophagus sinuatus Sharp, 1889 are designated. Distribution maps are provided for G. plagiatus, G. sinuatus, and G. suensoni. Geodromicus plagiatus is recorded from Cataluña province of Spain, Moldova, Georgia, Uzbekistan, and several regions of Russia (Pskov Area, Stavropol Area, Karachay-Cherkess and Chechen republics, Sverdlovsk Area, Khanty-Mansi Autonomous Region, Krasnoyarsk Territory, Evenkia and Yakutia), G. nigrita from Andorra and G. suturalis from Slovenia for the first time.

 

References

  1. Bernhauer, M. (1902) Neue Staphyliniden der paläarktischen Fauna nebst Bemerkungen. Deutsche Entomologische Zeitschrift, 1901, 241–251.
    Bernhauer, M. (1936) Neuheiten der palearktischen Staphylinidenfauna II. Pùbblicazioni del Museo Entomologico “Pietro Rossi” Duino, 1, 303–325.
    Bernhauer, M. (1941) Neuheiten der palaearktischen Staphylinidenfauna (Staphylinid.). Entomologische Blätter, 37, 209–211.
    Bernhauer, M. & Schubert, K. (1910) Staphylinidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. 5 (19). Junk, Berlin, pp. 1–86. https://doi.org/10.1007/978-94-011-9697-0_1
    Bordoni, A. (1984) Appunti per una revisione dei Geodromicus Redt. della regione Paleartica occidentale (Col., Staphylinidae). Redia, 67, 19–59.
    Brisout de Barneville, C. (1863) [New species]. In: Grenier, A. (Ed.), Catalogue des Coléoptères de France, par M. le Dr. A. Grenier, et matériaux pour servir a la faune des Coléoptères Français. A. Grenier, Paris, pp. i–iv + 3–79 + 1–135.
    Burakowski, B., Mroczkowski, M. & Stefańska, J. (1979) Katalog fauny Polski [Catalogus faunae Poloniae]. Część XXIII, tom 6. Chrząszcze [Coleoptera]. Kusakowate—Staphylinidae, część 1. Polska Akademia Nauk, Instytut Zoologii, Warsawa, 310 pp.
    Casey, T.L. (1893) Coleopterological notices. V. Annals of the New York Academy of Sciences, 7, 281–606. https://doi.org/10.1111/j.1749-6632.1893.tb55411.x
    Cheng, Z.-F., Shavrin, A.V. & Peng, Z. (2020) New species and records of Geodromicus Redtenbacher, 1857 from China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 4789 (1), 132–170. https://doi.org/10.11646/zootaxa.4789.1.4
    Duff, A.G. (2012) Family Staphylinidae Latreille, 1802. In: Duff, A.G. (Ed.), Checklist of Beetles of the British Isles. 2nd edition. Pemberly Books, Iver, pp. 33–55
    Eichler, W. (1930) Chrząszcze okolic Tyflisu—Coleoptera der Umgebung von Tiflis. Polskie pismo entomologiczne—Bulletin entomologique de la Pologne, 9 (3–4), 213–258.
    Eppelsheim, E. (1878) Staphylinidae. In: Schneider, O. & Leder, H., Beiträge zur Kenntniss der Kaukasischen Käferfauna. Verhandlungen des Naturforschenden Vereines in Brunn, 16 , 90–131.[1877]
    Eppelsheim, E. (1887) Synonymische Bemerkungen über europäische Staphylinen. Deutsche Entomologische Zeitschrift, 31, 430–432.
    Erichson, W.F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F.H. Morin, Berlin, pp. 401–954. https://doi.org/10.5962/bhl.title.59644
    Fabricius, J.C. (1798) Supplementum entomologiae systematicae. C.G. Proft & Storch, Hafniae, 2 + 572 pp.
    Fabricius, J.C. (1801) Systema eleutheratorum secundum ordines, genera, species: adiectis synonimis, locis, observationibus, descriptionibus. Tomus II. Bibliopolii Academici Novi, Kiliae, 687 pp.
    Fagel, G. (1958) Contribution à la connaissance des Staphylinidae. L. Sur quelques Geodromicus européens. Bulletin et Annales de la Société Royale d`Entomologie de Belgique, 94, 216–225.
    Fairmaire, L. & Laboulbène, A. (1856) s.n. In: Faune entomologique française ou description des insectes qui se trouvent en France. Vol. 1. Coléopteres. Deyrolle, Paris, pp. 371–665.
    Fauvel, A. (1865) Énumération des insectes recueillis en Savoie et en Dauphine (1861–1863) et descriptions d’espèces nouvelles. Bulletin de la Société Linnéenne de Normandie, 9, 253–321.
    Fauvel, A. (1871) Faune Gallo-Rhénane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhénanes et la Valais avec tableaux synoptiques et planches gravées. Bulletin de la Société Linnéenne de Normandie, Series 2, 5 (1869–1870), 27–192.
    Fauvel, A. (1872) Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhénane, la Nassau et la Valais avec tableaux synoptiques et planches gravées. Vol. 3. liv. 3. Le Branc-Hardel, Caen, 214 pp.
    Fauvel, A. (1878) Les Staphylinides de l’Amerique du nord. Bulletin de la Société Linnéenne de Normandie, 3 (2), 167–269.
    Fowler, W.W. (1888) The Coleoptera of the British Islands. A descriptive account of the families, genera, and species indigenous to Great Britain and Ireland, with notes as to localities, habitats, etc. Vol. II. Staphylinidae. L. Reeve & Co., London, 444 pp. https://doi.org/10.5962/bhl.title.29517
    Gamarra, P. & Outerelo, R. (2008) Catálogo iberobalear de los Omaliinae (Coleoptera: Staphylinidae). Boletin Sociedad Entomológica Aragonesa, 43, 211–231.
    Ganglbauer, L. (1895) Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des französischen und italienischen Alpengebietes. Zweiter Band. Familienreihe Staphylinoidea. 1.Theil: Staphylinidae, Pselaphidae. Carl Gerold’s Sohn, Wien, vi + 881 pp. https://doi.org/10.1002/mmnd.48018950245
    Gemminger, M. & Harold, B. de (1868) s.n. In: Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. II. Dytiscidae, Gyrinidae, Hydrophilidae, Staphylinidae, Pselaphidae, Gnostidae, Paussidae, Scydmaenidae, Silphidae, Trichopterygidae, Scaphidiidae. E.H. Gummi, Monachii, pp. 425–752 + [6].
    Germar, E.F. (1827) Fauna insectorum Europae. Fascicle 13. [Kuemmelii, Halae, 25 pls., [26] pp.
    Gravenhorst, J.L.C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. Carolus Reichard, Brunsuigae, lxvi + 206 pp.
    Gravenhorst, J.L.C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, xvi + 236 + [12] pp. https://doi.org/10.5962/bhl.title.67769
    Gyllenhal, L. (1810) Insecta Suecica descripta. Classis I. Coleoptera sive Eleuterata. 1 (2), i–xx + 1–660. [F. J. Leverentz, Scaris]
    Hansen, V. (1951) Biller XV. Rovbiller, 1. Danmarks Fauna 57. G.E.C. Gads Forlag, København, 274 pp.
    Hayashi, Y. (2018) Two new species of the genus Geodromicus Redtenbacher (Coleoptera, Staphylinidae, Omalinae [sic] from Japan). Elytra, Tokyo, New Series, 8 (1), 201–205.
    Heer, O. (1839) s.n. In: Fauna Coleopterorum Helvetica. 1 (2). Orelii, Fuesslini et Sociorum, Turici, pp. 145–360.
    Herman, L.H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1–7), i–vi + 1–4218 https://doi.org/10.1206/0003-0090.265.1.1
    Hochhuth, J.H. (1863) Beitraege zur naeheren Kenntnis der Staphylinen Russlands. Bulletin de la Société Impériale des Naturalistes de Moscou, 35 (3), 1–113.
    Iablokov-Khnzorian, S.M. (1989 [1988]) Dva novykh Kavkazskich predstavitelya iz roda Geodromicus Redt. (Coleoptera, Staphylinidae) [Two new Caucasian congeners of the genus Geodromicus Redt. (Coleoptera, Staphylinidae)]. Doklady Akademii Armianskoi SSR, 87 (3), 134–139. [in Russian, with Abstract in Armenian]
    Jarrige, J. (1963) Nouveaux brachyélytres pyrénéens. Bulletin de la Société Entomologique de France, 67 , 164–168.[1962]
    Johansen, J.P. (1914) Danmarks rovbiller eller Billefam. Staphylinidae’s danske Slaegter og Arter. Bianco Lunos Bogtrykkeri, København, [3] + 660 pp.
    Kashcheev, V.A. (1999) Novye vidy Oxytelinae (Coleoptera, Staphylinidae) fauny Kazakhstana [New species of Oxytelinae (Coleoptera, Staphylinidae) of the fauna of Kazakhstan]. Selevinia, 1996, 12–16. [in Russian]
    Kittel, G. (1873–1884) Systematische Übersicht der Käfer, welche in Baiern und der nächsten Umgebung vorkommen. Correspondenzblatt des zoologisch-mineralogischen Vereines in Regensburg, 27–38, 1–639.
    Khnzorian, S.M. (1962) Novye vidy zhestkokrylykh iz Zakavkazya. New species of Coleoptera from Transcaucasus (Insecta-Coleoptera). Zoologichesky Sbornik Zoologichesky Institut, Akademii Nauk Armyanskoy SSR, 12, 99–124. [in Russian, with Abstract in Armenian]
    Kraatz, G. (1857) s.n. In: Naturgeschichte der Insecten Deutschlands. Erste Abtheilung Coleoptera. Zweiter Band, Staphylinii. Lieferung 3–6. Nicolai, Berlin, pp. 377–1080.
    Lacordaire, J.T. (1835) s.n. In: Boisduval, J.B.A.D. & Lacordaire, J.T. (Eds.), Faune entomologique des environs de Paris; ou species général des insectes qui se trouvent dans un rayon de quinze à vingt lieues aux alentours de Paris. Tome premier. Méquignon-Marvis, Paris, pp. 1–17 + 1–696.
    Latreille, P.A. (1804) Histoire naturelle, générale et particulière, des crustacés et des insectes. 9. F. Dufart, Paris, 416 pp. https://doi.org/10.5962/bhl.title.15764
    LeConte, J.L. (1850) General Remarks upon the Coleoptera of Lake Superior. In: Agassiz, L. & Cabot, J.E. (Eds.), Lake Superior: Its physical character, vegetation, and animals, compared with those of other and similar regions. Gould, Kendall and Lincoln, Boston, Massachusetts, pp. 201–242.
    Li, J. (1992) The Coleoptera fauna of northeast China. Jilin, Jilin Education Publishing House, 205 pp. [in Chinese]
    Li, J. (1993) The rove beetles of northeast China. In: Li, J. & Chen, P. (Eds.), Studies on fauna and ecogeography of soil animal [sic!]. Press of Northeast Normal University, Changchun, pp. 1–63. [in Chinese]
    Li, J.-K., Zhang, L.-M. & Zhang, X.-P. (2015) [Primary color illustration of soil beetles in Northeast China. Staphylinidae and Tenebrionoidea]. Harbin Map Press, Harbin, 199 pp. [in Chinese]
    Lohse, G.A. (1958) Neuheiten der deutschen Käferfauna IV. Entomologische Blätter für Biologie und Systematik der Käfer, 54, 51–58.
    Lohse, G.A. (1964) Staphylinidae I. (Micropeplinae bis Tachyporinae). In: Freude, H., Harde, K. & Lohse, G. (Eds.), Die Käfer Mitteleuropas. Band 4. Goecke & Evers, Krefeld, pp. 1–264.
    Luze, G. (1903) Revision der paläarktischen Arten der Staphyliniden-Gattung Geodromicus Redtenb. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 103–117.
    Mannerheim, C.G. von (1830) Précis d’un nouvel arrangement de la famille des brachélytres de l’ordre des insectes coléoptères. s.n., St. Petersbourg, 87 pp.
    Mannerheim, C.G. von (1831) Précis d’un nouvel arrangement de la famille des brachélytres de l’ordre des insectes coléoptères. Mémoires présentés a L’Académie Impériale des Sciences de St. Pétersbourg, 1, 415–501.
    Motschulsky, V. de. (1860) Énumération des nouvelles especes de coléopteres rapportées de ses voyages. 3-ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539–588.
    Müller, P.W.J. (1821) III. Neue Insekten. Magazin der Entomologie, 4, 184–230. [Halle]
    Mulsant, E. & Rey, C. (1880) Tribu des brévipennes. Onzième famille: Omaliens. Douxième famille: Pholidiens. Annales de la Société Linnéenne de Lyon, Series 2, 27, 1–430. https://doi.org/10.3406/linly.1880.3986
    Neumann, K. (1907) Einiges über Staphylinen (Col.). Die Typen und Originale aus der Sammlung des †Pfarrers D. Scriba. Deutsche Entomologische Zeitschrift, 1907, 327–330.
    Newton, A.F. (2022) StaphBase: Staphyliniformia world catalog database (version October 2021): Staphylinoidea, Hydrophiloidea, Synteliidae, Sphaeritidae, Cretohisteridae. In: Bánki, O. & Roskov, Y. (Eds.), Catalogue of Life Checklist (October 2021). Species 2000: Naturalis, Leiden. Available from: http://www.catalogueoflife.org/ (accessed 18 June 2022)
    Notman, H. (1920) Coleoptera collected at Windsor, Broome Co., N.Y., 26 May to 5 June, 1918, with notes and descriptions. Journal of the New York Entomological Society, 28 (2), 178–194.
    Palm, T. (1948) Svensk Insekfauna. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Unterfam. Micropeplinae, Phloeocharinae, Olisthaerinae, Proteininae, Omaliinae. Almquist & Wiksells, Uppsala, 133 pp.
    Palm, T. (1961) Die schwedischen Arten der Gattung Geodromicus Redtb. mit Beschreibung einer neuen Art (Col. Staphylinidae). Opuscula Entomologica, 26, 153–157.
    Paykull, G. de (1800) Fauna Suecica. Insecta. Tomus III. J.F. Edman, Upsaliae, 459 pp.
    Pope, R.D. (1977) Coleoptera and Strepsiptera. In: Kloet, G.S. & Hincks, W.D. (Eds.), A checklist of British Insects. Second edition. Handbooks for the identification of British Insects. Royal Entomological Society of London, London, pp. 1–105.
    Poppius, B.R. (1907 [1905]) Beiträge zur Kenntnis der Coleopteren-Fauna des nordöstlichen Europäischen Russlands. I. Carnivora, Brachelytra, Silphidae und Corylophidae. Annuaire du Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 10, 302–315.
    Porta, A. (1926) Fauna Coleopterorum Italica. Vol. II. Stabilimento Tipografico Piacentino, Piacenza, 405 pp.
    Portevin, G. (1929) Histoire naturelle des coléoptères de France. Tome I. Adephaga. Polyphaga: Staphylinoidea. Encyclopédie Entomologique. Série A. Tome XII. P. Lechevalier, Paris, 649 pp.
    Redtenbacher, L. (1849) Bogen. In: Fauna Austriaca. Die Käfer. Nach der analytischen Methode bearbeitet. C. Gerold, Wien, pp. 41–58. https://doi.org/10.5962/bhl.title.37851
    Redtenbacher, L. (1857) Fauna Austriaca. Die Käfer. Nach der analytischen Methode bearbeitet. Zweite, gänzlich umgearbeitete, mit mehreren Hunderten von Arten und mit der Charakteristik sämmtlicher europäischen Käfergattungen vermehrte Auflage. C. Gerold’s Sohn, Wien, pp. 129–976. [1856–1858] https://doi.org/10.5962/bhl.title.9570
    Redtenbacher L. (1874) Fauna Austriaca. Die Käfer. Nach der analytischen Methode bearbeitet. Dritte, gänzlich umgearbeitete und beteutend vermehrte Auflage. C. Gerolds Sohn, Wien, 564 + 571 + viii + cliii pp., 2 pls.
    Reitter, E. (1870) Uebersicht der Käfer-Fauna von Mähren und Schlesien. Verhandlungen des Naturforschenden Vereines in Brünn, Abhandlungen, 8 (2), iii–viii + 1–195.
    Reitter, E. (1909) Fauna Germanica. Die Käfer des Deutschen Reiche. Nach der analytischen Methode bearbeitet. II Band. Schriften des Deutschen Lehrervereins für Naturkunde 24. K.G. Lutz, Stuttgart, 392 pp., pls. 41–80.
    Ryabukhin, A.S. (1999) A catalogue of rove beetles (Coleoptera: Staphylinidae exclusive of Aleocharinae) of the northeast of Asia. Pensoft Series Faunistica No. 16. Pensoft, Sofia and Moscow, 140 pp.
    Sahlberg, C.R. (1827) s.n. In: Insecta Fennica, dissertationibus academicis, A. 1817–1834 editis. Pars I: A. Helsingforsiae, Frenckelliana, pp. 185–280.
    Sahlberg, J. (1876) Enumeratio coleopterorum brachypterorum Fenniae. Systematisk förteckning öfver de inom Finlands naturalhistoriska område hittils funna Coleoptera Brachelytra jemte uppgift om arternas utbredning och beskrifningar af nya och mindre kända species. I. Staphylinidae. Acta Societatis pro Fauna Flora Fennica, 1, 1–248.
    Sainte-Claire Deville, J. (1907) Faune des coleopteres du Bassin de la Seine. II. Staphylinoidea. Société Entomologique de France, Paris, 92 pp.
    Say, T. (1823) Descriptions of coleopterous insects collected in the late expedition to the Rocky Mountains, performed by order of Mr. Calhoun, Secretary of War, under the command of Major Long. Journal of the Academy of Natural Sciences of Philadelphia, 3, 139–216.
    Say, T. (1831) [untitled continuation of Say, T. (1830)]. In: Say, T. (Ed.), Description of new species of North American insects, and observations on some already described, pp. 42–49. [New Harmony, Indiana]
    Say, T. (1834) Descriptions of new North American insects, and observations on some already described. Transactions of the American Philosophical Society, New Series, 4, 409–470.
    Scheerpeltz, O. von (1933) Staphylinidae VII: Supplementum I (pars 129). In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 989–1500.
    Schülke, M. & Smetana, A. (2015) Staphylinidae. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea—Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304–1134.
    Scriba, W. (1863) Die Käfer im Großherzogthum Hessen und seiner nächsten Umgebung. Berichte der Oberhessischen Gesellschaft für Natur- und Heilkunde, 10 (1863), 1–61.
    Sharp, D. (1889) The Staphylinidae of Japan [concluded]. The Annals and Magazine of Natural History, Serie 6, 3 (18), 463–476. https://doi.org/10.1080/00222938909460371
    Smetana, A. (1959) Die tschechoslowakischen Arten der Gattung Geodromicus Redt. (Col., Staphylinidae). Časopis Československé Společnosti Entomologické, 56, 355−363.
    Steel, W.O. (1970) The larvae of the genera of the Omaliinae (Coleoptera: Staphylinidae) with particular reference to the British fauna. Transactions of the Royal Entomological Society of London, 122 (1), 1–47. https://doi.org/10.1111/j.1365-2311.1970.tb00524.x
    Stein, J.P.E.F. (1868) Catalogus Coleopterorum Europae. F. Nicolai, Berlin, iv + 149 pp.
    Stephens, J.F. (1834) Illustrations of British entomology; or, a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, 5, 305–368. [Baldwin and Cradock, London]
    Stephens, J.F. (1839) A manual of British Coleoptera, or beetles; containing a brief description of all the species of beetles hitherto ascertained to inhabit Great Britain and Ireland; together with a notice of their chief localities, times and places of appearances, etc. Longman, Orme Brown, Green, and Longmans, London, xii + 443 pp. https://doi.org/10.5962/bhl.title.59227
    Szujecki, A. (2008) Klucze do oznaczania owadów Polski. Cz. XIX. Chrząszcze—Coleoptera. Zeszyt. 24a. Kusakowate—Staphylinidae Wstęp oraz podrodziny: Micropeplinae, Piestinae, Osoriinae, Pseudopsiinae [sic], Phloeocharinae, Olisthaerinae, Proteininae, Omaliinae, Oxytelinae, Oxyporinae. Polskie Towarzystwo Entomologiczne, Toruń, 229 pp.
    Thomson, C.G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol 1. Berlingska Boktryckeriet, Lund, 290 pp. https://doi.org/10.5962/bhl.title.138677
    Thomson, C.G. (1861) Skandinaviens Coleoptera, synoptiskt bearbetade, 3, 1–278. [Berlingska Boktryckeriet, Lund]
    Tóth, L. (1982) Holyvák II.—Staphylinidae II. Magyarország Állatvilága—Fauna Hungariae VII. Kötet, Coleoptera II. 6. Füzet. Fauna Hung. 151. Akadémiai Kiadó, Budapest, 110 + [4] pp.
    Tottenham, C.E. (1954) Coleoptera, Staphylinidae. Section (a) Piestinae to Euaesthetinae. Handbooks for the Identification of British Insects, 4 (8), 1–79.
    Watanabe, Y. (1972) Some staphylinid beetles from the Hidaka Mountains in Hokkaido, Japan. Memoirs of the National Science Museum, 5, 111–121.
    Watanabe, Y. (1985) Omaliinae. In: Uéno, S.-L., Kurosawa, Y. & Satô, M. (Eds.), The Coleoptera of Japan in Color, 2, pp. 264–268, pls. 46–47. [Hoikusha, Osaka, in Japanese]
    Watanabe, Y. (1986) Staphylinid beetles found along the water edges of mountain streams in Oku-Nikko. In: Editorial Board of Fauna and Flora of Nikko (Ed.), Fauna and Flora of Nikko, 1986, pp. 541–547. [Kanuma, Tochinoha-shob.ANt., in Japanese]
    Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59–391.
    Waterhouse, G.R. (1858) s.n. In: Catalog of British Coleoptera. Taylor and Francis, London, pp. i–iv + 3–117.
    Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. Fauna d’Italia, Calderini, Bologna, 25, i–xii + 1–472.
    Zanetti, A. (1995) Genera 1−46. Ciceroni, A., Puthz, V. & Zanetti, A. Fascicolo 48, Coleoptera Polyphaga III (Staphylinidae). In: Minelli, A., Ruffo, S. & La Posta, S. (Eds.), Checklist delle Specie della Fauna Italiana. Edizioni Calderini, Bologna, pp. 1–65.
    Zanetti, A. (2011) Unterfamilie Omaliinae. In: Assing, V. & Schülke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer—Die Käfer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49−117.
    Zanetti, A., Pavićević, D. & Zerche, L. (2008) Contribution to the knowledge of Omaliinae of the Balkans (Coleoptera, Staphylinidae). In: Pavićević, D. & Perreau, M. (Eds.), Advances in the studies of the Fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia Monograph, 22, pp. 397–416.
    Zerche, L. (2003) Arten der Hygrogeus-Gruppe aus dem Fernen Osten Russlands, aus Japan und China (Coleoptera, Staphylinidae, Omaliinae, Anthophagini). Beiträge zur Entomologie, 53 (2), 277–295.
    Zetterstedt, J.W. (1828) Fauna insectorum Lapponica. Pars I. Libraria Schulziana, Hammone, XX + 563 pp.
    Zetterstedt, J.W. (1838) Columns 7–868. In: Insecta Lapponica. Lipsiae: Leopoldi Voss, vi pp., 1140 columns. [issued in parts: 1838–1840]

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