Abstract
Studies of ticks in Malaysia from past articles were reviewed, resulting in a list of 47 species belonging to seven genera (Argas, Ornithodoros, Amblyomma, Dermacentor, Haemaphysalis, Ixodes, and Rhipicephalus). The most prevalent genus in Malaysia is Haemaphysalis (20 species), followed by Amblyomma (eight species) and Dermacentor (seven species). Out of 47 species, only 28 have bitten humans, mostly belonging to Haemaphysalis. The most researched tick-borne pathogens in Malaysia are Rickettsia and Anaplasma, and most research was focused on the tropical-lineage brown dog ticks, R. sanguineus sensu lato, and the cattle ticks, Haemaphysalis bispinosa and Rhipicephalus microplus. 18 species were excluded from the list due to lack of definite records or dubious findings: Ornithodoros mimon, O. turicata, Amblyomma breviscutatum, A. clypeolatum, A. integrum, A. maculatum, Dermacentor marginatum, D. taiwanensis, Haemaphysalis birmaniae, H. flava, H. humerosa, H. longicornis, H. punctata, H. sulcata, Ixodes holocyclus, Rhipicephalus appendiculatus, R. annulatus and R. bursa. This paper presents the first complete and updated list for Dermacentor and Ixodes tick species in Malaysia since Kohls (1957).
References
Ajithkumar, K.G., Ravindran, R. & Ghosh, S. (2012) Dermacentor auratus Supino, 1897 (Acarina, Ixodidae) reported from Wayanad, Kerala. The Indian Journal of Medical Research, 135, 435–436.
Ali, A., Zahid, H., Zeb, I., Tufail, M., Khan, S., Haroon, M., Tufail, M., Bilal, M., Hussain, M., Alouffi, A.S., Muñoz-Leal, S. & Labruna, M.B. (2021) Risk factors associated with tick infestations on equids in Khyber Pakhtunkhwa, Pakistan, with notes on Rickettsia massiliae detection. Parasites & Vectors, 14, 363. https://doi.org/10.1186/s13071-021-04836-w
Alsarhan, Q.T.S. (2017) Prevalence, risk factors and genetic diversity of equine piroplasmosis in Kelantan, Malaysia. Ph.D dissertation. Universiti Malaysia Kelantan, Kelantan, 366 pp.
Álvarez, V.C., Hernández, V.F. & Hernández, J.G. (2005) Catálogo de garrapatas suaves (Acari: Argasidae) y duras (Acari: Ixodidae) de Costa Rica. Brenesia, 2005, 63–64 + 81–88.
Amarga, A.K.S., Supsup, C.E., Tseng, H.Y., Kwak, M.L. & Lin, S.M. (2022) The Asian turtle tick Amblyomma geoemydae Cantor, 1847 (Acari: Ixodidae) in the Philippines: first confirmed local host and locality with a complete host index. Ticks and Tick-borne Diseases, 13, 101958. https://doi.org/10.1016/j.ttbdis.2022.101958
Amin-Babjee, S.M., Lee, C.C. & Krishnasamy, M. (1985) A preliminary survey of parasites of Malaysian red jungle fowl (Gallus gallus spadiceus). Kajian Veterinaire, 17, 141–146.
Amin-Babjee, S.M., Lee, C.C. & Mahmood, A.A. (1997) Prevalence of ectoparasite infestation in different age groups of village chicken. Jurnal Veterinar Malaysia, 9, 55–59.
Anastos, G. (1950) The scutate ticks, or Ixodidae, of Indonesia. Entomologica Americana, 30, 1–144.
Andrew, R., Bonnin J.M. & Williams, S. (1946) Tick typhus in North Queensland. The Medical Journal of Australia, 2, 253–258. https://doi.org/10.5694/j.1326-5377.1946.tb34304.x
Aoki, C. (1993) Five human cases of ixodid tick-bite in Oita Prefecture, Japan. Medical Entomology and Zoology, 44, 405–406. https://doi.org/10.7601/mez.44.405
Apanaskevich, D.A. (2016) First description of the nymph and larva of Dermacentor compactus (Acari: Ixodidae), parasites of squirrels in Southeast Asia. Systematic Parasitology, 93, 355–365. https://doi.org/10.1007/s11230-016-9633-0
Apanaskevich, D.A. & Apanaskevich, M.A. (2016) Description of two new species of Dermacentor Koch, 1844 (Acari: Ixodidae) from Oriental Asia. Systematic Parasitology, 93, 159–171. https://doi.org/10.1007/s11230-015-9614-8
Apanaskevich, D.A., Apanaskevich, M.A, Nooma, W., Ahantarig, A. & Trinachartvanit, W. (2021) Reinstatement of Dermacentor tricuspis (Schulze, 1933) n. comb., n. stat. (Acari: Ixodidae) as a valid species, synonymization of D. atrosignatus Neumann, 1906 and description of a new species from Indonesia, Malaysia and Thailand. Systematic Parasitology, 98, 207–230. https://doi.org/10.1007/s11230-021-09972-6
Apanaskevich, M.A. & Apanaskevich, D.A. (2015a) Description of new Dermacentor (Acari: Ixodidae) species from Malaysia and Vietnam. Journal of Medical Entomology, 52, 156–162. https://doi.org/10.1093/jme/tjv001
Apanaskevich, M.A. & Apanaskevich, D.A. (2015b) Reinstatement of Dermacentor bellulus (Acari: Ixodidae) as a valid species previously confused with D. taiwanensis and comparison of all parasitic stages. Journal of Medical Entomology, 52, 573–595. https://doi.org/10.1093/jme/tjv034
Apanaskevich, D.A., Bandarayanaka, K.O, Apanaskevich, M.A. & Rajakaruna, S. (2016) Redescription of Amblyomma integrum adults and immature stages. Medical and Veterinary Entomology, 30, 330–341. https://doi.org/10.1111/mve.12178
Aragão, H.B. (1911) Notas sobre ixódidas brazileiros. Memorias do Instituto Oswaldo Cruz, 3, 145–195. https://doi.org/10.1590/S0074-02761911000200001
Ariyarathne, S., Apanaskevich, D.A, Amarasinghe, P.H. & Rajakaruna, R.S. (2016) Diversity and distribution of tick species (Acari: Ixodidae) associated with human otoacariasis and socio-ecological risk factors of tick infestations in Sri Lanka. Experimental and Applied Acarology, 70, 90–123. https://doi.org/10.1007/s10493-016-0056-z
Asyikha, R., Sulaiman, N. & Mohd-Taib, F.S. (2020) Detection of Bartonella sp. in ticks and their small mammal hosts in mangrove forests of Peninsular Malaysia. Tropical Biomedicine, 37, 919–931. https://doi.org/10.47665/tb.37.4.919
Audy, J.R., Nadchatram, N. & Boo-Liat, L. (1960) Malaysian parasites XLIX. Host distribution of Malayan ticks (Ixodoidea). Studies from the Institute for Medical Research, Federated Malay States, 29, 225–246.
Barker, S.C. & Walker, A.R. (2014) Ticks of Australia. The species that infest domestic animals and humans. Zootaxa, 3816 (1), 1–144. https://doi.org/10.11646/zootaxa.3816.1.1
Barker, S.C., Walker, A.R. & Campelo, D. (2014) A list of the 70 species of Australian ticks; diagnostic guides to and species accounts of Ixodes holocyclus (paralysis tick), Ixodes cornuatus (southern paralysis tick) and Rhipicephalus australis (Australian cattle tick); and consideration of the place of Australia in the evolution of ticks with comments on four controversial ideas. International Journal for Parasitology, 44, 941–953. https://doi.org/10.1016/j.ijpara.2014.08.008
Barros-Battesti, D.M., Landulfo, G.A., Onofrio, V.C., Faccini, J.L.H., Marcili, A., Nieri-Bastos, F.A., Venzal, J.M. & Labruna, M.B. (2011) Carios mimon (Acari: Argasidae): description of adults and redescription of larva. Experimental and Applied Acarology, 54, 93–104. https://doi.org/10.1007/s10493-010-9416-2
Beck, A.J. (1971) A survey of bat ectoparasites in West Malaysia. Journal of Medical Entomology, 8, 147–152. https://doi.org/10.1093/jmedent/8.2.147
Bochkov, A.V., Abramov, A.V., Durden, L.A., Apanaskevich, D.A., Stekolnikov, A.A., Stanyukovich, M.K., Gnophanxay, S. & Tikhonov, A.N. (2011) Arthropod symbiotes of Laonastes aenigmamus (Rodentia: Diatomyidae). Journal of Parasitology, 97, 352–353. https://doi.org/10.1645/GE-2508.1
Boero, J.J. (1944) Notas ixodológicas. II) Nueva lista de los ixodideos argentinos y sus huéspedes. Revista de la Asociación Médica Argentina, 58, 354–355.
Boero, J.J. (1955) Los ixodideos de la República Argentina y sus huéspedes. Revista de la Facultad de Agronomia y Veterinaria, 12, 505–514.
Borsoi, A.B.P. & Serra-Freire, N.M. (2012) Relações parasitárias entre humanos e carrapatos no município de Volta Redonda, Estado do Rio de Janeiro. Revista UNIABEAU, 5, 306–317.
Burridge, M.J. (2011) Non-native and invasive ticks. threats to human and animal health in the United States. University of Florida Press, Gainesville, Florida, 292 pp.
Bursali, A., Tekin, S., Orhan, M., Keskin, A. & Ozkan, M. (2010) Ixodid ticks (Acari: Ixodidae) infesting humans in Tokat Province of Turkey: species diversity and seasonal activity. Journal of Vector Ecology, 35, 180–186. https://doi.org/10.1111/j.1948-7134.2010.00075.x
Camicas, J.L., Hervy, J.P., Adam, F. & Morel, P.C. (1998) Les tiques du monde (Acarida, Ixodida). Nomenclature, stades décrits, hôtes, réparition. ORSTOM, Paris, 233 pp.
Chang, P.H. (1958a) A study of Amblyomma cyprium Neumann, 1899, collected from human body. Acta Ecologica Sinica, 8, 290–292.
Chang, P.H. (1958b) A case of tick infestation in the anus of the human body. National Medical Journal of China, 44, 304–305.
Chao, L.L., Hsieh, C.K. & Shih, C.M. (2012) First report of Amblyomma helvolum (Acari: Ixodidae) from the Taiwan stink snake, Elaphe carinata (Reptilia: Colubridae), collected in southern Taiwan. Ticks and Tick-borne Diseases, 4, 246–250. https://doi.org/10.1016/j.ttbdis.2012.11.002
Chao, L.L., Lu, C.W., Lin, Y.F. & Shih, C.M. (2017) Molecular and morphological identification of a human biting tick, Amblyomma testudinarium (Acari: Ixodidae), in Taiwan. Experimental and Applied Acarology, 71, 401–414. https://doi.org/10.1007/s10493-017-0119-9
Chao, L.L., Chen, T.H., Lien, W.C., Erazo, E. & Shih, C.M. (2022) Molecular and morphological identification of a reptile-associated tick, Amblyomma geoemydae (Acari: Ixodidae), infesting wild yellow-margined box turtles (Cuora flavomarginata) in northern Taiwan. Ticks and Tick-borne Diseases, 13, 101901. https://doi.org/10.1016/j.ttbdis.2022.101901
Che-Lah, F.C., Mariana, A., Haron, M.S. & Ho, T.M. (2013) Blood meal identification of field collected on-host ticks surrounding two human settlements in Malaysia. The Experiment, 17, 1177–1183.
Che-Lah, E.F., Yaakop, S., Ahamad, M. & Md-Nor, S. (2015) Molecular identification of blood meal sources of ticks (Acari, Ixodidae) using cytochrome b gene as a genetic marker. Zookeys, 478, 27–43. https://doi.org/10.3897/zookeys.478.8037
Che-Lah, E.F., Yaakop, S., Ahamad, M., George, E. & Md-Nor, S. (2014) Molecular phylogeny of a tick, Ixodes granulatus (Acari: Ixodidae) based on cytochrome oxidase subunit I (COI) marker. AIP Conference Proceedings, 1614, 719. https://doi.org/10.1063/1.4895290
Che-Lah, E.F., Yaakop, S., Ahamad, M., George, E. & Md-Nor, S. (2016) Precise identification of different stages of a tick, Ixodes granulatus Supino, 1897 (Acari: Ixodidae). Asian Pacific Journal of Tropical Biomedicine, 6, 597–604. https://doi.org/10.1016/j.apjtb.2016.05.003
Chien, N.T.H., Linh, B.K., Tho, N.V., Hieu, D.D. & Lan, N.T. (2016) Status of cattle ticks infection in yellow and dairy cows in Ba Vi District. In: Proceedings of the international conference on agriculture development in the context of international integration: opportunities and challenges, Hanoi, Vietnam, 7–8 December 2016. [unknown pagination]
Chong, S.T., Kim, H.C., Park, J.G., Choi, C.Y., Park, C.U., Klein, T.A. & Robbins, R.G. (2018) Ticks surveillance of migratory birds during 2010-2011 on Hong and Heuksan Islands, Jeollanam Province, Republic of South Korea. Systematic and Applied Acarology, 23, 2214–2223. https://doi.org/10.11158/saa.23.11.13
Clavijo, J.J., Vásquez, C.L. & Vlaera, N.C. (2009) Garrapatas duras de los géneros Amblyomma Koch y Rhipicephalus Koch (Acari: Ixodidae) presents en la colección de zoología agrícola, decanato de agronomía, Universidad Centro Occidental Lisandro Alvarado, Venezuela. Entomotrópica, 24, 41–50.
Colas-Belcour, J. & Millot, J. (1948) Contributions à l’étude des ixodidés de Madagascar. Sur une variété nouvelle de Haemaphysalis hoodi. Parasitisme humain par un Boophilus. Bulletin de la Société de Pathologie Exotique, 41, 384–388.
Department of Wildlife and National Parks of Peninsular Malaysia (2010) Red list of mammals for Peninsular Malaysia. Department of Wildlife and National Parks of Peninsular Malaysia, Kuala Lumpur, 159 pp.
Department of Wildlife and National Parks of Peninsular Malaysia (2017) Red list of mammals for Peninsular Malaysia Version 2.0. Department of Wildlife and National Parks of Peninsular Malaysia, Kuala Lumpur, 216 pp.
Dhanda, V. & Rao, T.R. (1964) A report on a collection of ixodid ticks made in the North East Frontier Agency, India. Indian Journal of Medical Research, 52, 1139–1153.
Díaz-Ungría, C. (1957) Nota sobre las especies de Acarina de Venezuela. Revista de Sanidad y Asistencia Social, 22, 457–467.
Doeve, W. (1923) The tick in Celebes. Nederland. Indisch. BI. Diergeneesk. Dierent., 34, 17–23.
Domrow, R. & Nadchatram, M. (1963) Two field collections of Malayan mites and ticks. Malayan Nature Journal, 17, 145–164.
Donaldson, T.G., Pèrez-de-León, A.A., Li, A.I., Castro-Arellano, I., Wozniak, E., Boyle, W.K., Hargrove, R., Wilder, H.K., Kim, H.J., Teel, P.D. & Lopez, J.E. (2016) Assessment of the geographic distribution of Ornithodoros turicata (Argasidae): climate variation and host diversity. PLoS Neglected Tropical Diseases, 10, e0004383. https://doi.org/10.1371/journal.pntd.0004383
Doss, M.A., Farr, M.M., Roach, K.F. & Anastos, G. (1974) Index-Catalogue of Medical and Veterinary Zoology. Department of Agriculture, Washington, D.C., 663 pp.
Durden, L.A., Merker, S. & Beati, L. (2008) The tick fauna of Sulawesi, Indonesia (Acari: Ixodoidea: Argasidae and Ixodidae). Experimental and Applied Acarology, 45, 85–110. https://doi.org/10.1007/s10493-008-9144-z
Edussuriya, B.D.P. & Weilgama, D.J. (2003) Case reports: intra-aural tick infestations in humans in Sri Lanka. Transactions of the Royal Society of Tropical Medicine and Hygiene, 97, 412–413. https://doi.org/10.1016/S0035-9203(03)90072-1
Ellis, S. & Talukdar, B. (2020) Rhinoceros sondaicus. The IUCN Red List of Threatened Species. https://doi.org/10.2305/IUCN.UK.2020-2.RLTS.T19495A18493900.en
Ernieenor, F.C.L., Ernna, G. & Mariana, A. (2017) Phenotypic and genotypic identification of hard ticks of the genus Haemaphysalis (Acari: Ixodidae) in Peninsular Malaysia. Experimental and Applied Acarology, 71, 387–400. https://doi.org/10.1007/s10493-017-0120-3
Ernieenor, F.C.L., Apanaskevich, D.A., Ernna, G. & Mariana, A. (2020) Description and characterization of questing hard tick, Dermacentor steini (Acari: Ixodidae) in Malaysia based on phenotypic and genotypic traits. Experimental and Applied Acarology, 80, 137–149. https://doi.org/10.1007/s10493-019-00439-4
Ernieenor, F.C.L., Ernna, G., Apanaskevich, D.A., Mariana, A. & Yaakop, S. (2022) First record of the tortoise tick, Amblyomma geoemydae (Cantor, 1847) (Acari: Ixodidae) parasitizing a tree shrew, Tupaia glis (Scandentia: Tupaiidae) in West Malaysia. Journal of Medical Entomology, 59, 1473–1478. https://doi.org/10.1093/jme/tjac042
Ernieenor, F.C.L., Apanaskevich, D.A., Ernna, G., Ellyncia, B.B., Md-Zain, B.M., Mariana, A. & Yaakop, S. (2021) Morphological and molecular identification of medically important questing Dermacentor species collected from some recreational areas of Peninsular Malaysia. Systematic Parasitology, 98, 731–751. https://doi.org/10.1007/s11230-021-10008-2
Estrada-Peña, A., Mihalca, A.D. & Petney, T.N. (2017) Ticks of Europe and North Africa: a guide to species identification. Springer Nature, Cham, 368 pp. https://doi.org/10.1007/978-3-319-63760-0
Estrada-Peña, A., Venzal, J.M., Nava, S., Mangold, A., Guglielmone, A.A., Labruna, M.B. & de la Fuente, J. (2012) Reinstatement of Rhipicephalus (Boophilus) australis (Acari: Ixodidae) with redescription of the adult and larval stages. Journal of Medical Entomology, 49, 794–802. https://doi.org/10.1603/ME11223
Fatimah, C.T.N.I., Hassuzana, K., Amin-Babjee, S.M., Saharee, A.A. & Sharifah, S.H. (1994) Skin conditions in sheep around Serdang, Malaysia. Jurnal Veterinar Malaysia, 6, 37-41.
Filippova, N.A. & Chunikhin, S.P. (1969) The tick Argas (Chiropterargas) boueti Roubaud and Coulas-Belcour, 1933 (Parasitiformes, Argasidas). A new species for the USSR fauna. Zoologicheskii Zhurnal, 48, 1407–1409.
Gargili, A., Certi, Ç., Alp, H.G., Kar, S., Sönmez, G., Vatansever, Z., Yilmazer, N. & Şahin, F. (2010) Evaluation of ticks biting humans in Thrace Province, Turkey. Kafkas Universitesi Veteriner Fakultesi Dergisi, 16 (Supplement A), S141–S146.
Gould, D.J., Clifford, C.M., Lakshana, P., Emerson, E.C., Hoogstraal, H., Marshall, M.T., Stark, H.E., Nadchatram, M., Davis, W.E., Inlao, I., Maneechai, N., Klaimanee, N. & Punthusiri, P. (1970) Distribution and ecology of ectoparasites of vertebrates in Southeast Asia. Annual Program Report SEATO Medical Research Laboratory, United States Army Medical Component, Bangkok, Thailand, April 1969–March 1970. [unknown pagination]
Grassman, L.I., Sarataphan, N., Tewes, M.E., Silvy, N.J. & Nakanakrat, T. (2004) Ticks (Acari: Ixodidae) parasitizing wild carnivores in Phu Khieo wildlife sanctuary. Journal of Parasitology, 90, 657–659. https://doi.org/10.1645/GE-3327RN
Green, P.E. (1971) An unusual host of Boophilus microplus. Australian Veterinary Journal, 47, 179–180. https://doi.org/10.1111/j.1751-0813.1971.tb02139.x
Grokhovskaya, I.M. & Nguyen, H.H. (1968) Contribution to the study of ixodid ticks (Ixodidae) in Vietnam. Meditsinskaia Parazitologiia, 37, 710–715.
Guerrero-Sanchez, S. (2020) Ecology and health of the Asian water monitor lizard (Varanus salvator) in the fragmented landscape of the Kinabatangan floodplain, Sabah, Malaysia. Ph.D. dissertation, Cardiff University, Cardiff, Wales, 153 pp.
Guglielmone, A.A. & Nava, S. (2014) Names for Ixodidae (Acari: Ixodoidea): valid, synonyms, incertae sedis, nomina dubia, nomina nuda, lapsus, incorrect and suppressed names—with notes on confusions and misidentifications. Zootaxa, 3767 (1), 1–256. https://doi.org/10.11646/zootaxa.3767.1.1
Guglielmone A.A. & Robbins, R.G. (2018) Hard ticks (Acari: Ixodida: Ixodidae) parasitizing humans. A global overview. Springer, Cham, 326 pp. https://doi.org/10.1007/978-3-319-95552-0
Guglielmone, A.A., Nava, S. & Robbins, R.G. (2021) Neotropical hard ticks (Acari: Ixodida: Ixodidae). A critical analysis of their taxonomy, distribution, and host relationships. Springer Nature, Cham, 486 pp. https://doi.org/10.1007/978-3-030-72353-8
Guglielmone, A.A., Mangold, A.J. & Viñabal, A.E. (1991) Ticks (Ixodidae) parasitizing humans in four provinces of northwestern Argentina. Annals of Tropical Medicine and Parasitology, 85, 539–542. https://doi.org/10.1080/00034983.1991.11812606
Guglielmone, A.A., Robbins, R.G., Apanaskevich, D.A., Petney, T.N., Estrada-Peña, A. & Horak, I.G. (2014) The hard ticks of the world (Acari: Ixodida: Ixodidae). Springer, Dordrecht, 751 pp. https://doi.org/10.1007/978-94-007-7497-1
Guglielmone, A.A., Sánchez, M.E., Franco, L.G., Nava, S., Rueda, L.M. & Robbins, R.G. (2015) Available from: http://rafaela.inta.gob.ar/nombresgarrapatas/Hard_ticks_query.aspx
Gunes, T., Poyraz, O. & Vatansever, Z. (2011) Crimean-Congo hemorrhagic fever virus in ticks collected from humans, livestock, and picnic sites in the hyperendemic region of Turkey. Vector-Borne and Zoonotic Diseases, 11, 1411–1416. https://doi.org/10.1089/vbz.2011.0651
Hara, N., Hou, S., Miyazaki, T., Yamakage, A., Yamazaki, S., Kawai, S., Chigusa, Y. & Matsuda, H. (1999) A case of Amblyomma testudinarium bite. Japanese Journal of Tropical Medicine and Hygiene, 27, 141.
Hasegawa, T., Takahasi, T., Okano, T., Kanoh, H., Fujihiro, M., Maeda, M., Shikano, Y., Kachi, H., Watanabe, A., Kadosaka, T. & Seishima, M. (2016) A clinical study of 94 patients with tick bite in 8 medical institutions in Gifu Prefecture. The Japanese Journal of Dermatology, 126, 2095–2102.
Hashizume, H., Fujiyama, T., Umayahara, T., Kageyama, R., Walls, A.F. & Satoh, T. (2018) Repeated Amblyomma testudinarium tick bites are associated with elevated anti-galactose-ɑ-1,3-galactose carbohydrate IgE antibody levels: a retrospective cohort study in a single institute. Journal of the American Academy of Dermatology, 78, 1135–1141. https://doi.org/10.1016/j.jaad.2017.12.028
Hassan, M., Sulaiman, M.H. & Lian, C.J. (2013) The prevalence and intensity of Amblyomma javanense infestation on Malayan Pangolins (Manis javanica Desmarest) from Peninsular Malaysia. Acta Tropica, 126, 142–145. https://doi.org/10.1016/j.actatropica.2013.02.001
Heisch, R.B. (1951) Argas boueti Roubaud and Colas-Belcour 1933, in the northern province of Kenya. East African Medical Journal, 28, 354.
Hirst, S. (1929) Fauna of Batu Caves, Selangor. XI. Arachnida: Acarina (Ixodidae). Journal of the Federated Malay States Museums, 14, 365.
Hirst, S. & Hirst, L.F. (1910) Description of five new species of ticks (Ixodidae). Annals and Magazine of Natural History, including Zoology, Botany, and Geology, 8, 299–308. https://doi.org/10.1080/00222931008692856
Ho, T.M. & Krishansamy, M. (1990) Ectoparasitic acari and endoparasitic of small mammals in Taman Negara. Journal of Wildlife and Parks, 10, 54–61.
Hoogstraal, H. (1955) Bat ticks the of genus Argas (Ixodoidea, Argasidae). 1. The subgenus Chiropterargas. Fieldiana. Zoology, 37, 579–600. https://doi.org/10.5962/bhl.title.2895
Hoogstraal, H. (1956) African Ixodoidea 1. Ticks of the Sudan (with special reference to Equatoria Province and with preliminary reviews of Boophilus, Margaropus and Hyalomma). Department of the Navy, Bureau of Medicine and Surgery, Washington, D.C., 1105 pp.
Hoogstraal, H. (1964) Studies on Southeast Asian Haemaphysalis ticks (Ixodoidea, Ixodidae). Redescription, hosts, and distribution of H. traguli Oudemans. The larva and nymph of H. vidua W. and N. Identity of H. papuana toxopei Warburton (new combination). Journal of Parasitology, 50, 765–782. https://doi.org/10.2307/3276200
Hoogstraal, H. (1966) Studies on Southeast Asian Haemaphysalis ticks (Ixodoidea, Ixodidae). H. (H.) capricornis sp. n., the large malayan serow haemaphysalid from Southwestern Thailand. Journal of Parasitology, 52, 783–786. https://doi.org/10.2307/3276456
Hoogstraal, H. (1970) Haemaphysalis (H.) birmaniae Supino and H. (H.) goral sp. n. (Ixodoidea: Ixodidae), Asian parasites of artiodactyl mammals. Journal of Parasitology, 56, 1227–1238. https://doi.org/10.2307/3277573
Hoogstraal, H. (1971) Haemaphysalis (Kaiseriana) borneata sp. n. (Ixodoidea: Ixodidae), a tick of the H. (K.) aculeata group parasitizing the sambar deer in Borneo. Journal of Parasitology, 57, 1096–1098. https://doi.org/10.2307/3277869
Hoogstraal, H. (1973) Viruses and ticks. In: Gibbs, A.J. (Ed.), Viruses and invertebrates. North Holland Publishing Company, Amsterdam, pp. 351–390.
Hoogstraal, H. (1985) Argasid and nuttallielid ticks as parasites and vectors. Advances in Parasitology, 24, 136–220. https://doi.org/10.1016/S0065-308X(08)60563-1
Hoogstraal, H. & El-Kammah, K.M. (1971) Studies on Southeast Asian Haemaphysalis ticks (Ixodoidea: Ixodidae). H. (H.) traubi Kohls, redescription of male, description of female, and new artiodactyl host and Malayan distribution records. Journal of Parasitology, 57, 426–431. https://doi.org/10.2307/3278054
Hoogstraal, H. & Kim, K.C. (1985) Tick and mammal coevolution, with emphasis on Haemaphysalis. In: Kim, K.C. (Ed.), Coevolution of parasitic arthropods and mammals. John Wiley & Sons, New York, pp. 1–800.
Hoogstraal, H. & Trapido, H. (1966) Studies on Southeast Asian Haemaphysalis ticks (Ixodoidea, Ixodidae). Species described by Supino in 1897 from Burma, with special reference to H. (Rhipistoma) asiaticus (= H. dentipalpis Warburton and Nuttall). Journal of Parasitology, 52, 1172–1187. https://doi.org/10.2307/3276365
Hoogstraal, H. & Wassef, H.Y. (1973) The Haemaphysalis ticks (Ixodoidea: Ixodidae) of birds. 3. H. (Ornithophysalis) subgen. n.: definition, species, hosts, and distribution in the Oriental, Palearctic, Malagasy, and Ethiopian faunal regions. Journal of Parasitology, 59, 1099–1117. https://doi.org/10.2307/3278650
Hoogstraal, H. & Wassef, H.Y. (1982) Haemaphysalis (Garnhamphysalis) mjoebergi: identity, structural variation and biosystematic implications, deer hosts, and distribution in Borneo and Sumatra (Ixodoidea: Ixodidae). Journal of Parasitology, 68, 138–144. https://doi.org/10.2307/3281337
Hoogstraal, H. & Wassef, H.Y. (1984) Dermacentor (Indocentor) compactus (Acari: Ixodoidea: Ixodidae): wild pigs and other hosts and distribution in Malaysia, Indonesia, and Borneo. Journal of Medical Entomology, 21, 174–178. https://doi.org/10.1093/jmedent/21.2.174
Hoogstraal, H. & Wassef, H.Y. (1985) Dermacentor (Indocentor) auratus (Acari: Ixodoidea: Ixodidae): hosts, distribution, and medical importance in tropical Asia. Journal of Medical Entomology, 22, 170–177. https://doi.org/10.1093/jmedent/22.2.170
Hoogstraal, H., Lim, B.L. & Anastos, G. (1969) Haemaphysalis (Kaiseriana) bispinosa Neumann (Ixodoidea: Ixodidae): evidence for consideration as an introduced species in the Malay Peninsula and Borneo. Journal of Parasitology, 55, 1075–1077. https://doi.org/10.2307/3277178
Hoogstraal, H., Santana, F.J. & Van Peenen, P.F.D. (1968a) Ticks (Ixodoidea) of Mt. Sontra, Danang, Republic of Vietnam. Annals of the Entomological Society of America, 61, 722–729. https://doi.org/10.1093/aesa/61.3.722
Hoogstraal, H., Trapido, H. & Kohls, G.M. (1965a). Studies on Southeast Asian Haemaphysalis ticks (Ixodoidea, Ixodidae). H. atherurus sp. n. and redescription of type material of H. birmaniae Supino, 1897. Journal of Parasitology, 51, 114–125. https://doi.org/10.2307/3275657
Hoogstraal, H., Trapido, H. & Kohls, G.M. (1965b) Studies on Southeast Asian Haemaphysalis ticks (Ixodoidea, Ixodidae). The identity, distribution, and hosts of H. (Kaiseriana) hystricis Supino. Journal of Parasitology, 51, 467–480. https://doi.org/10.2307/3275974
Hoogstraal, H., Trapido, H. & Kohls, G.M. (1965c) Southeast Asian Haemaphysalis ticks (Ixodoidea, Ixodidae). H. (Kaiseriana) papuana nadchatrami ssp. n. and redescription of H. (K.) semermis Neumann. Journal of Parasitology, 51, 433–451. https://doi.org/10.2307/3275971
Hoogstraal, H., Clifford, C.M., Keirans, J.E. & Wassef, H.Y. (1979) Recent developments in biomedical knowledge of Argas ticks (Ixodoidea: Argasidae). In: Rodriguez, J.G. (Ed.), Recent advances in acarology. Vol. II. Academic Press, New York, pp. 269–278. https://doi.org/10.1016/B978-0-12-592202-9.50040-9
Hoogstraal, H., El Kammah, K.M., Santana, F.J. & Van Peenen, P.F.D. (1973) Studies on Southeast Asian Haemaphysalis ticks (Ixodoidea: Ixodidae). H. (Kaiseriana) lagrangei Larrouse: identity, distribution, and hosts. Journal of Parasitology, 59, 1118–1129. https://doi.org/10.2307/3278651
Hoogstraal, H., Lim, L.B., Nadchatram, M. & Anastos, G. (1972) The Gunong Benom Expedition 1967. 8. Ticks (Ixodidae) of Gunong Benom and their altitudinal distribution, hosts and medical relationships. Bulletin of the British Museum (Natural History), Zoology, 23, 167–186. https://doi.org/10.5962/p.314200
Hoogstraal, H., Roberts, F.H.S., Kohls, G.M. & Tipton, V.J. (1968b) Review of Haemaphysalis (Kaiseriana) longicornis Neumann (resurrected) of Australia, New Zealand, New Caledonia, Fiji, Japan, Korea, and northeastern China and USSR, and its parthenogenetic and bisexual populations (Ixodoidea, Ixodidae). Journal of Parasitology, 54, 1197–1213. https://doi.org/10.2307/3276992
Hoogstraal, H., Saito, Y., Dhanda, V. & Bhat, H.R. (1971) Haemaphysalis (H.) obesa Larrousse (Ixodoidea: Ixodidae) from Northeast India and Southeast Asia: description of immature stages and biological observations. Journal of Parasitology, 57, 177–184. https://doi.org/10.2307/3277776
Horak, I.G., Heyne, H., Williams, R., Gallivan, G.J., Spickett, A., Bezuidenhout, J.D. & Estrada-Peña, A. (2018) The ixodid ticks (Acari: Ixodidae) of Southern Africa. Springer Nature, Cham, 676 pp. https://doi.org/10.1007/978-3-319-70642-9
Hussain, N.S., Arshad, S.S., Amin-Babjee, S.M. & Sharma, R. (2010) Parasite and virus infracommunity of Malayan water monitor lizard (Varanus salvator). 5th Proceedings of the Seminar in Veterinary Sciences, 5–8 January 2010. [unknown pagination]
Hutcheson, H.J., Dergousoff, S.J. & Lindsay, L.R. (2019) Haemaphysalis longicornis: a tick of considerable veterinary importance, now established in North America. The Canadian Veterinary Journal, 60, 27–28. https://doi.org/10.17269/s41997-018-0152-4
Indudharan, R., Dharap, A.S. & Htun, Y.N. (1995) An unusual differential diagnosis of myringitis bullosa haemorrhagica. Tropical and Geographical Medicine, 47, 227–228.
Indudharan, R., Ahamad, M., Ho, T.M., Salim, R. & Htun, Y.N. (1999) Human otoacariasis. Annals of Tropical Medicine and Parasitology, 93, 163–167. https://doi.org/10.1080/00034983.1999.11813406
Ishak, S.N., Shiang, L.F., Khoo, J.J. & Mohd-Taib, F.S. (2019) Screening of Rickettsia sp. In ticks (Acari: Ixodidae) collected from small mammals in three recreational forests in Selangor, Malaysia. Serangga, 24, 116–125.
Ishak, S.N., Yusof, M., Md-Nor, S., Mohd-Sah, S., Lim, F., Khoo, J.J. & Mohd-Taib, F. (2018) Prevalence of on-host ticks (Acari: Ixodidae) in small mammals collected from forest near to human vicinity in Selangor, Malaysia. Systematic and Applied Acarology, 23, 1531–1544. https://doi.org/10.11158/saa.23.8.4
Ishak, S.N., Shiang, L.F., Mohd-Taib, F.S., Khoo, J.J., Md-Nor, S., Yusof, M.A., Sah, S.A.M., Sitam, F.T. & Japning, J.R.R. (2018) Molecular identification of hard ticks (Ixodes sp.) infesting rodents in Selangor, Malaysia. AIP Conference Proceedings, 1940, 020054. https://doi.org/10.1063/1.5027969
Ishibashi, O., Niizuma, J., Miura, A., Iizuka, S., Fujita, H., Ogura, G., Sakashita, M., Ganeko, H., Sunagawa, K. & Nakada, T. (2009) Survey of parasitic ixodid ticks on small Asian mongoose on Okinawajima Island, Japan. Japanese Journal of Zoo and Wildlife Medicine, 14, 51–57. https://doi.org/10.5686/jjzwm.14.51
Ishida, M., Shiori, Y., Ishizaka, T., Iwasaki, H., Fujita, H. & Takada, N. (2004) A case of multi-infestation with more than 200 larval Amblyomma testudinarium in human. Hifu no Kagaku, 3, 55–61.
Isohisa, T., Nakai, N., Okuzawa, Y., Yamada, M., Arizono, N. & Katoh, N. (2011) Case of tick bite with infestation of an extraordinary number of larval Amblyomma testudinarium ticks. Journal of Dermatology, 38, 1110–1112. https://doi.org/10.1111/j.1346-8138.2010.01180.x
Ivancovich, J.C. & Luciani, C.A. (1992) Las garrapatas de Argentina. Monografía de la Asociación Argentina de Parasitología Veterinaria. Asociación Argentina de Parasitología Veterinaria, Buenos Aires, 95 pp.
Jiang, Z.J. (1983) Biology of Haemaphysalis hystricis Supino. Acta Entomologica Sinica, 11, 413–418.
Joo, H.J., Kim, M.R., Cho, B.K., Park, H.J., Lee, I.Y. & Kim, T.H. (2016) Tick bite by male Amblyomma testudinarium in a Korean woman. Korean Journal of Dermatology, 54, 125–128.
Kambayashi, C., Hemmi, K. & Kurabayashi, A. (2021) First record of Amblyomma testudinarium Koch, 1844 (Acari: Ixodidae) parasitising Reeves’ Pond Turtle, Mauremys reevesii (Gray, 1831) (Testudines: Geoemydidae), in Japan. Herpetology Notes, 14, 613–615.
Kamio, T., Rajamanickam, C., Kawazu, S. & Fujisaki, K (1990) Epidemiology and pathogenicity of bovine theileriosis in Malaysia. Japan Agricultural Research Quarterly, 24, 231–234.
Kawashima, K., Kamo, H. & Miyazaki, I. (1960) A case of human infestation with a hard tick Amblyomma testudinarium Koch, 1844 in Japan (Acarina: Ixodidae). Kyushu Journal of Medical Science, 11, 77–80.
Kazim, A.R., Houssaini, J., Ehlers, J., Tappe, D. & Heo, C.C. (2021) Soft ticks (Acari: Argasidae) in the island nations of Southeast Asia: A review on their distribution, associated hosts and potential pathogens. Acta Tropica, 223, 106085. https://doi.org/10.1016/j.actatropica.2021.106085
Kazim, A.R., Low, V.L., Houssaini, J., Tappe, D. & Heo, C.C. (2022a) Morphological abnormalities and multiple mitochondrial clades of Rhipicephalus haemaphysaloides (Ixodida: Ixodidae). Experimental and Apllied Acarology, 87, 133–141. https://doi.org/10.1007/s10493-022-00731-w
Kazim, A.R., Low, V.L., Tappe, D., Houssaini, J. & Heo, C.C. (2022b) Rhipicephalus annulatus, R. australis or R. microplus? Discordance between morphological and genetic data among three cattle tick species. Experimental and Applied Acarology, 87, 119–131. https://doi.org/10.1007/s10493-022-00726-7
Keegan, H.L. & Toshioka, S. (1957) Ixodid Ticks of Japan, Korea and the Ryukyu Islands. Medical General Laboratory, Camp Zama, Japan, 37 pp.
Keirans, J.E. (1985) George Henry Falkiner Nuttall and the Nuttall tick catalogue. U.S. Dept. of Agriculture, Agricultural Research Service, Washington, D.C., 1785 pp. https://doi.org/10.5962/bhl.title.65326
Keirans, J.E. & Durden, L.A. (2001) Invasion: exotic ticks (Acari: Argasidae, Ixodidae) imported into the United States. A review and new records. Journal of Medical Entomology, 38, 850–861. https://doi.org/10.1603/0022-2585-38.6.850
Khattak, R.M., Ali, S., Jahangir, M., Khan, M.N., Rasul, A. & Iqbal, F. (2012) Prevalence of ectoparasites in wild and domesticated grey (Francolinus pondicerianus) and black partridge (Francolinus francolinus) from Khyber Pakhtoonkhwa Province of Pakistan. Pakistan Journal of Zoology, 44, 1239–1244.
Kho, K.L., Tan, P.E. & Tay, S.T. (2019) Diversity of Rickettsiae in feeding and questing ticks collected from a Malaysian forest reserve area. Journal of Medical Entomology, 56, 547–552. https://doi.org/10.1093/jme/tjy168
Kho, K.L., Koh, F.X., Hasan, L.I.M., Wong, L.P., Kisomi, M.G., Bulgiba, A., Nizam, Q.N.H. & Tay, S.T. (2017) Rickettsial seropositivity in the indigenous community and animal farm workers, and vector surveillance in Peninsular Malaysia. Emerging Microbes & Infections, 6, 1–9. https://doi.org/10.1038/emi.2017.4
Kho, K.L., Koh, F.X., Jaafar, T., Nizam, Q.N.H. & Tay, S.T. (2015a) Prevalence and molecular heterogeneity of Bartonella bovis in cattle and Haemaphysalis bispinosa ticks in Peninsular Malaysia. BMC Veterinary Research, 11, 153. https://doi.org/10.1186/s12917-015-0470-1
Kho, K.L., Koh, F.X. & Tay, S.T. (2015b) Molecular evidence of potential novel spotted fever group rickettsiae, Anaplasma and Ehrlichia species in Amblyomma ticks parasitizing wild snakes. Parasites & Vectors, 8, 112. https://doi.org/10.1186/s13071-015-0719-3
Khoo, J.J., Chen, F., Kho, K.L., Shanizza, A.I.A., Lim, F.S., Tan, K.K. & Abu-Bakar, S. (2016a) Bacterial community in Haemaphysalis ticks of domesticated animals from the Orang Asli communities in Malaysia. Ticks and Tick-borne Diseases, 7, 929–937. https://doi.org/10.1016/j.ttbdis.2016.04.013
Khoo, J.J., Husin, N.A., Lim, F.S., Oslan, S.N.H., Mohd Azami, S.N.I., To, S.W., Abd Majid, M.A., Lee, H.Y., Loong, S.K., Khor, C.S. & Abu-Bakar, S. (2020) Molecular detection of pathogens from ectoparasites recovered from peri-domestic animals, and the first description of a Candidatus Midichloria sp. From Haemaphysalis wellingtoni from rural communities in Malaysia. Parasitology International, 80, 102202. https://doi.org/10.1016/j.parint.2020.102202
Khoo, J.J., Ishak, S.N., Lim, F.S., Mohd-Taib, F.S., Khor, C.S., Loong, S.K. & Abu-Bakar, S. (2018) Detection of a Borrelia sp. from Ixodes granulatus ticks collected from rodents in Malaysia. Journal of Medical Entomology, 55, 1642–1647. https://doi.org/10.1093/jme/tjy122
Khoo, J.J., Lim, F.S., Chen, F., Phoon, W.H., Khor, C.S., Pike, B.L. & Abu-Bakar, S. (2016b) Coxiella detection in ticks from wildlife and livestock in Malaysia. Vector-borne and Zoonotic Diseases, 16, 744–751. https://doi.org/10.1089/vbz.2016.1959
Khoo, J.J., Lim, F.S., Tan, K.K., Chen, F.S., Phoon, W.H., Khor, C.S. & Abu-Bakar, S. (2017) Detection in Malaysia of a Borrelia sp. from Haemaphysalis hystricis (Ixodida: Ixodidae). Journal of Medical Entomology, 54, 1444–1448. https://doi.org/10.1093/jme/tjx131
Khoo, J.J., Husin, N.A., Lim, F.S., Oslan, S.N.H., Mohd-Azami, S.N.I., To, S.W., Abd-Majid, M.A., Lee, H.Y., Loong, S.K., Khor, C.S. & Abu-Bakar, S. (2021) Molecular detection of pathogens from ectoparasites recovered from peridomestic animals, and the first description of a Candidatus Midichloria sp. from Haemaphysalis wellingtoni from rural communities in Malaysia. Parasitology International, 80, 102202. https://doi.org/10.1016/j.parint.2020.102202
Kim, J., Joo, H.S., Moon, H.J. & Lee, Y.J. (2010) A case of Amblyomma testudinarium tick bite in a Korean woman. The Korean Journal of Parasitology, 48, 313–317. https://doi.org/10.3347/kjp.2010.48.4.313
Kim, J., Kang, H.A., Kim, S.S., Joo, H.S. & Chong, W.S. (2014) Perianal tick-bite lesion caused by a fully engorged female Amblyomma testudinarium. The Korean Journal of Parasitology, 52, 685–690. https://doi.org/10.3347/kjp.2014.52.6.685
Kim, Y.H., Lee, J.H., Park, Y.M. & Lee, J.Y. (2016) Tick bite by nymphal Amblyomma testudinarium. Annals of Dermatology, 28, 762–764. https://doi.org/10.5021/ad.2016.28.6.762
Koh, F.X. (2018) A molecular survey of tick-borne pathogens (Anaplasma and Ehrlichia spp.) in animal and tick samples in Malaysia. Master dissertation, University of Malaya, Kuala Lumpur, 198 pp.
Koh, F.X., Kho, K.L., Kisomi, M.G., Wong, L.P., Bulgiba, A., Tan, P.E. & Tay, S.T. (2017) Ehrlichia and Anaplasma infections: serological evidence and tick surveillance in peninsular Malaysia. Journal of Medical Entomology, 55, 269–276. https://doi.org/10.1093/jme/tjx204
Koh, F.X., Kho K.L., Panchadcharam, C., Sitam, F.T. & Tay, S.T. (2016) Molecular detection of Anaplasma spp. in pangolins (Manis javanica) and wild boars (Sus scrofa) in Peninsular Malaysia. Veterinary Parasitology, 227, 73–76. https://doi.org/10.1016/j.vetpar.2016.05.025
Koh, F.X., Panchadcharam, C. & Tay, S.T. (2015) Vector-borne diseases in stray dogs in Peninsular Malaysia and molecular detection of Anaplasma and Ehrlichia spp. from Rhipicephalus sanguineus (Acari: Ixodidae) ticks. Journal of Medical Entomology, 53, 183–187. https://doi.org/10.1093/jme/tjv153
Kohls, G.M. (1950) Ticks (Ixodoidea) of the Philippines. National Institutes of Health, Washington, D.C., 34 pp.
Kohls, G.M. (1957) Malaysian parasites. 18. Ticks (Ixodoidea) of Borneo and Malaya. Studies from the Institute of Medical Research, Federation of Malaya, 28, 65–94.
Kohls, G.M. (1962) A new species of Ixodes from Malaya (Acarina, Ixodidae). Proceedings of the Entomological Society of Washington, 64, 103–105.
Kohls, G.M. & Clifford, C.M. (1968) Ixodes (Lepidixodes) kopsteini Oudemans, 1925, valid name for I. (L.) paradoxus Kohls and Clifford, 1961, with notes on distribution and hosts (Acarina: Ixodidae). Journal of Medical Entomology, 5, 503–506. https://doi.org/10.1093/jmedent/5.4.503
Kohls, G.M., Clifford, C.M. & Jones, E.K. (1969) The systematics of the subfamily Ornithodorinae (Acarina: Argasidae). IV. Eight new species of Ornithodoros from the Western Hemisphere. Annals of the Entomological Society of America, 62, 1035–1043. https://doi.org/10.1093/aesa/62.5.1035
Kolonin, G.V. (1995) Review of the ixodid tick fauna (Acari: Ixodidae) of Vietnam. Journal of Medical Entomology, 32, 276–282. https://doi.org/10.1093/jmedent/32.3.276
Kolonin, G.V. (2009) Fauna of ixodid ticks of the world. Available from: http://www.kolonin.org (accessed 13 September 2022)
Konto, M., Fufa, G.I., Zakaria, A., Tukur, S.M., Watanabe, M., Ola-Fadunsin, S.D., Khan, M.S., Shettima, Y.M. & Babjee, S.M.A. (2015) Tick fauna of Malaysian red jungle fowl (Gallus gallus) in Bangi, Malaysia. Veterinary World, 8, 1167–1171. https://doi.org/10.14202/vetworld.2015.1167-1171
Krige, A.S., Loh, S.M. & Oskam, C.L. (2017) New host records for ticks (Acari : Ixodidae) from the echidna (Tachyglossus aculeatus) revealed in Australian museum survey. Australian Journal of Zoology, 65, 379–382. https://doi.org/10.1071/ZO18018
Krijgsman, B.J. & Ponto, S.A.S. (1932) De Teken van den Oost-Indischen Archipel. Veeartsenijkundige Mededeelingen, 79, 1–62.
Kumar, K.G.A., Ravindran, R., Johns, J., Chandy, G., Rajagopal, K., Chandrasekhar, L., George, A.J. & Ghosh, S. (2018) Ixodid tick vectors of wild mammals and reptiles of southern India. Journal of Arthropod-borne Diseases, 12, 276–285.
Kuo, C.C., Lin, Y.F., Yao, C.T., Shih, H.C., Chung, L.H., Liao, H.C., Hsu, Y.C. & Wang, H.C. (2017) Tick-borne pathogens in ticks collected from birds in Taiwan. Parasites & Vectors, 10, 587. https://doi.org/10.1186/s13071-017-2535-4
Kwak, M.L. (2017) Keys for the morphological identification of the Australian paralysis ticks (Acari: Ixodidae), with scanning electron micrographs. Experimental and Applied Acarology, 72, 93–101. https://doi.org/10.1007/s10493-017-0133-y
Kwak, M.L. (2020) A checklist and key to the tick fauna (Acari: Ixodidae, Argasidae) of Pulau Tioman, Malaysia. Experimental and Applied Acarology, 81, 51–58. https://doi.org/10.1007/s10493-020-00487-1
Kwak, M.L., Neves, E.S., Borthwick, S.A., Smith, G.J.D., Meier, R. & Mendenhall, I.H. (2021) Habitat impacts the abundance and network structure within tick (Acari: Ixodidae) communities on tropical small mammals. Ticks and Tick-borne Diseases, 12, 101654. https://doi.org/10.1016/j.ttbdis.2021.101654
Kwak, M. L., Foo, M., Pocklington, K., Hsu, C.D., Cheong, W., How, C.B., Shunari, M. & Tahir, M.G. (2019) Tick-crocodilian interactions: a review, with the first record of tick (Acari: Ixodidae) infestation in the saltwater crocodile (Crocodylus porosus), and a concise host-parasite index. Experimental and Applied Acarology, 78, 127–132. https://doi.org/10.1007/s10493-019-00378-0
Labruna, M.B., Camargo, L.M.A, Terrassini, F.A, Ferreira, F., Schumaker, T.T.S. & Camargo, E.P. (2005) Ticks (Acari: Ixodidae) from the State of Rondônia, western Amazon, Brazil. Systematic and Applied Acarology, 10, 17–32. https://doi.org/10.11158/saa.10.1.4
Lamattina, D. & Nava, S. (2016) Ticks infesting humans in northern Misiones, Argentina. Medicina, Buenos Aires, 76, 89–92.
Lamattina, D., Venzal, J.M., Costa, S.A., Arrabal, J.P., Flores, S., Berrozpe, P.E., González-Acuña, D., Guglielmone, A.A. & Nava, S. (2018) Ecological characterization of a tick community across a landscape gradient exhibiting differential anthropogenic disturbance in the Atlantic Forest ecoregion in Argentina. Medical and Veterinary Entomology, 32, 271–281. https://doi.org/10.1111/mve.12295
Lancaster, W.E. (1939) Report of the Veterinary Department of the Federated Malay States. Kuala Lumpur: the Federated Malay States, 49–52.
Latreille, P.A. (1806) Genera crustaceorum et insectorum secundum ordinem naturalem in familia disposita, iconibus exemplisque plurimis explicata. Vol. 1. A. Koenig, Parisiis, 302 pp. https://doi.org/10.5962/bhl.title.65741
Lau, A.C.C., Qui, Y., Mohamed-Abdallah, M.M., Nakao, R., Shimozuru, M., Onuma, M., Mohd-Azlam, J. & Tsubota, T. (2020) Detection of Borrelia burgdorferi sensu lato and relapsing fever Borrelia in feeding Ixodes ticks and rodents in Sarawak, Malaysia: New geographical records of Borrelia yangtzensis and Borrelia miyamotoi. Pathogens, 9, 1–17. https://doi.org/10.3390/pathogens9100846
Lee, C.C. & Amin-Babjee, S.M. (1993) New host records of parasites in the Malayan red jungle fowl, Gallus gallus spadiceus. Pertanika Journal of Tropical Agricultural Science, 16, 107–110.
Li, L.H., Zhang, Y., Wang, J.Z., Li, X.S., Yin, S.Q., Zhu, D., Xue, J.B. & Li, S.G. (2018) High genetic diversity in hard ticks from a China-Myanmar border county. Parasites & Vectors, 11, 469. https://doi.org/10.1186/s13071-018-3048-5
Lim, F.S., Khoo, J.J., Akhavanrezeai, M., Loong, S.K. Khor, C.S. & Abu-Bakar, S. Detection of Theileria luwenshuni (Piroplasmida: Theileriidae) from ticks infesting goats in peninsular Malaysia. (2019) Systematic and Applied Acarology, 24, 1971–1974. https://doi.org/10.11158/saa.24.10.14
Lim, F.S, Khoo, J.J, Chen, F., Bell-Sakyi, L., Khor, C.S., Chang, L.Y. & Abu-Bakar, S. (2017) Initiation of primary cell cultures from embryonic Haemaphysalis bispinosa ticks. Systematic and Applied Acarology, 22, 323–332. https://doi.org/10.11158/saa.22.3.1
Lim, F.S., Khoo, J.J., Tan, K.K., Zainal, N., Loong, S.K., Khor, C.S. & Abu-Bakar, S. (2020) Bacterial communities in Haemaphysalis, Dermacentor and Amblyomma ticks collected from wild boar of an Orang Asli Community in Malaysia. Ticks and Tick-borne Diseases, 11, 101352. https://doi.org/10.1016/j.ttbdis.2019.101352
Lim, F.S., Loong, S.K., Khoo, J.J., Tan, K.K., Zainal, N., Abdullah, M.F. & Abu-Bakar, S. (2018) Identification and characterization of Corynebacterium lactis isolated from Amblyomma testudinarium of Sus scrofa in Malaysia. Systematic and Applied Acarology, 23, 1838–1844. https://doi.org/10.11158/saa.23.9.10
Liyanaarachchi, D., Rajakaruna, R.S., Dikkumbura, A.W. & Rajapakse, R.V.P.J. (2015a) Ticks infesting wild and domestic animals and humans of Sri Lanka with new host records. Acta Tropica, 142, 64–70. https://doi.org/10.1016/j.actatropica.2014.11.001
Liyanaarachchi, D.R., Rajakaruna, R.S., Dikkumbura, A.W, De Silva, A. & Rajapakse, R.P.V.J. (2015b) Ticks (Acarina: Ixodida) infesting five reptile species in Sri Lanka with sixteen new host records. Zootaxa, 3964 (1), 146–148. https://doi.org/10.11646/zootaxa.3964.1.11
Loong, S.K., Ishak, S.N., Lim, F.S., Khoo, J.J., Tan, S.N., Freddy-Jalin, E.J., Mohd-Taib, F.S. & Abu-Bakar, S. (2018) Paenibacillus lautus, an opportunistic bacterial pathogen, isolated from Ixodes granulatus Supino (Acari: Ixodidae) collected from a Müller’s giant Sunda rat (Sundamys muelleri). Systematic and Applied Acarology, 23, 597–602. https://doi.org/10.11158/saa.23.4.2
Loong, S.K., Lim, F.S., Khoo, J.J., Lee, H.Y., Suntharalingam, C., Ishak, S.N., Mohd-Taib, F.S. & Abu-Bakar, S. (2020) Culturable pathogenic bacteria in ticks parasitizing farm animals and rodents in Malaysia. Tropical Biomedicine, 37, 803–811. https://doi.org/10.47665/tb.37.3.803
Low, V.L., Prakash, B.K., Lim, Y.A.L., Tan, T.K., Vinnie-Siow, Y.M., Sofian-Azirun, M. & Abu-Bakar, S. (2018) Detection of Anaplasmataceae agents and co-infection with other tick-borne protozoa in dogs and Rhipicephalus sanguineus sensu lato ticks. Experimental and Applied Acarology, 75, 429–435. https://doi.org/10.1007/s10493-018-0280-9
Ma, M., Li, S.C. & Fan, Q.H. (2015) Mites and ticks (Acari) in Shanxi Province, China: an annotated checklist. Zootaxa, 4006 (1), 1–39. https://doi.org/10.11646/zootaxa.4006.1.1
Maa, T.C. & Kuo, J.S. (1966) Catalogue and bibliography of ticks and mites parasitic on vertebrates in Taiwan. Q. Journal of the National Taiwan Museum, 19, 373–413.
Macadam, I., Gudan, D., Timbs, D.V., Urquhart, H.R. & Sewell, M.M.H. (1984) Metazoan parasites of dogs in Sabah, Malaysia. Tropical Animal Health and Production, 16, 34–38. https://doi.org/10.1007/BF02248927
Madinah, A., Mariana, A. & Abdullah, M.T. (2014) Detection of Rickettsiae in engorged ticks from small mammals in Malaysia. Borneo Journal of Resource Science and Technology, 4, 34–41. https://doi.org/10.33736/bjrst.242.2014
Madinah, A., Fatimah, A., Mariana, A. & Abdullah, M.T. (2011) Ectoparasites of small mammals in four localities of wildlife reserves in Peninsular Malaysia. The Southeast Asian Journal of Tropical Medicine and Public Health, 42, 803–813.
Madinah, A., Mariana, A., Fatimah, A. & Abdullah, M.T. (2013) A preliminary field survey of ectoparasites of rodents in urban park, Sarawak, Malaysian Borneo. Tropical Biomedicine, 30, 547–551.
Mahara, F. (1997) Japanese spotted fever: report of 31 cases and review of the literature. Emerging Infectious Diseases, 3, 105–111. https://doi.org/10.3201/eid0302.970203
Mahara, F. (2006) Rickettsiosis in Japan and the Far East. Annals of the New York Academy of Sciences, 1078, 60–73. https://doi.org/10.1196/annals.1374.007
Malaysia Biodiversity Information System. (2016) Geographical. Available from: https://www.mybis.gov.my/art/143 (accessed 13 September 2022)
Mao, J.J., Norval, G., Robbins, R.G. & Wong, S.T. (2019) An instance of Boiga dendrophila dendrophila (Boie, 1827) (Reptilia: Colubridae) being parasitized by Amblyomma helvolum Koch, 1844 (Acari: Ixodidae), with comments about the attachment sites of this tick species. Acarologia, 59, 115–119. https://doi.org/10.24349/acarologia/20194316
Mariana, A., Mohd Kulaimi, B., Halimaton, I., Suhaili, Z., Shahrul-Anuar, M., Nor Zalipah, M. & Ho, T. (2011) Acarine ectoparasites of Panti Forest Reserve in Johore, Malaysia. Asian Pacific Journal of Tropical Biomedicine, 1, 1–5. https://doi.org/10.1016/S2221-1691(11)60057-9
Mariana, A., Ibrahim, H., Bujang, M.K., Mohd Sah, S.A., Mohamad, N., Nor, S.M. & Ho, T.M. (2013) A survey of acarine ectoparasites of bats (Chiroptera) in Malaysia. Journal of Medical Entomology, 50, 140–146. https://doi.org/10.1603/ME11240
Mariana, A., Zuraidawati, Z., Ho, T.M., Mohd-Kulaimi, B., Saleh, I., Shukor, M.N. & Shahrul-Anuar, M.S. (2005) A survey of ectoparasites in Gunung Stong Forest Reserve, Kelantan, Malaysia. The Southeast Asian Journal of Tropical Medicine and Public Health, 36, 1125–1131.
Mariana, A., Zuraidawati, Z., Ho, T.M., Mohd0Kulaimi, B., Saleh, I., Shukor, M.N. & Shahrul-Anuar, M.S. (2008a) Ticks (Ixodidae) and other ectoparasites in Ulu Muda Forest Reserve, Kedah, Malaysia. The Southeast Asian Journal of Tropical Medicine and Public Health, 36, 496–506.
Mariana, A., Srinovianti, N., Ho, T.M., Halimaton, I., Hatikah, A., Shaharudin, M.H., Rosmaliza, I., Wan-Islah, L. & Sathananthar, K.S. (2008b) Intra-aural ticks (Acari: Metastigmata: Ixodidae) from human otoacariasis cases in Pahang, Malaysia. Asian Pacific Journal of Tropical Medicine, 1, 20–24.
Martins, T.F., Teixeira, R.H., Souza, J.C., Luz, H.R., Montenegro, M.M., Jerusalinsky, L., Bueno, M.G., Onofrio, V.C., Amorim, M., Gazêta, G.S., Silva, P.D.J., Bitencourth, K., Borsoi, A.B.P., Marques, S., Mattos, M.O., Hernandes, L.S.I., Scofild, A., Vieira, R.F.C., Pacheco, R.C., Horta, M.C., Silva, V.P., Silva, P.W., Igayara, C.A., Sanches, T.C., Nardi, M.S., Lugarini, C., Maia, N.L., Siqueira, C.L.M., Ferreira, J.M., Soares, J.F. & Labruna, M.B. (2021) Ticks (Parasitiformes: Ixodida) on new world wild primates in Brazil. International Journal of Acarology, 47, 95–106. https://doi.org/10.1080/01647954.2020.1870554
Matsuda, M., Oiso, N., Yano, Y. & Kawada, A. (2011) Dermoscopy for tick bite: reconfirmation of the usefulness for the initial diagnosis. Case Reports in Dermatology, 3, 94–97. https://doi.org/10.1159/000328181
Merten, H.A. & Durden, L.A. (20000 A state-by-state survey of ticks recorded from humans in the United States. Journal of Vector Ecology, 25, 102–113.
Michalik, J., Wodecka, B., Liberska, J., Dabert, M., Postawa, T., Piksa, K. & Stańczak, J. (2020) Diversity of Borrelia burgdorferi sensu lato species in Ixodes ticks (Acari: Ixodidae) associated with cave-dwelling bats from Poland and Romania. Ticks and Tick-borne Diseases, 11, 101300. https://doi.org/10.1016/j.ttbdis.2019.101300
Mihalca, A.D., Gherman, C.M. & Cozma, V. (2011) Coendangered hard-ticks: threatened or threatening? Parasites & Vectors, 4, 71. https://doi.org/10.1186/1756-3305-4-71
Ministry of Health. (2017) Case definitions for infectious diseases in Malaysia. Ministry of Health, 124 pp.
Mitchell, R.M. & Dick, J.A. (1978) Ectoparasites from Nepal birds. Journal of the Bombay Natural History Society, 74, 264–274.
Mitchell, C.J., Hoogstraal, H., Schaller, G.B. & Spillett, J. (1966) Ectoparasites from mammals in Kanha National Park, Madhya Pradesh, India, and their potential disease relationships. Journal of Medical Entomology, 3, 113–124. https://doi.org/10.1093/jmedent/3.2.113
Mohd-Hasan, L.I., Kho, K.L., Koh, F.X., Nizam, Q.N.H. & Tay, S.T. (2017) Molecular evidence of hemoplasmas in Malaysian cattle and ticks. Tropical Biomedicine, 34, 668–674.
Mohd-Taib, F.S., Asyikha, R. & Md-Nor, S. (2021) Small mammal assemblages d their ectoparasite prevalence (Acarina) in mangrove forests of Peninsular Malaysia. Tropical Zoology, 34, 24–43. https://doi.org/10.4081/tz.2021.78
Mohd-Zain, S.N., Sahimin, N., Pal, P. & Lewis, J.W. (2013) Macroparasite communities in stray cat populations from urban cities in Peninsular Malaysia. Veterinary Parasitology, 196, 469–477. https://doi.org/10.1016/j.vetpar.2013.03.030
Moritz, M. & Fischer, S.C. (1981) Die typen der arachniden-sammlung des Zoologischen Museums Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin, 57, 341–346. https://doi.org/10.1002/mmnz.19810570205
Munaf, H.B. (1978) Tick fauna of Baluran Wildlife Reserve, Indonesia. Hemera Zoa, 70, 37–44.
Nadchatram, M. (2006). A review of endoparasitic acarines of Malaysia with special reference to novel endoparasitism of mites in amphibious sea snakes and supplementary notes on ecology of chiggers. Tropical Biomedicine, 23, 1–22.
Nadchatram, M. (2008) The beneficial rain forest ecosystem, with environmental effects on zoonoses involving ticks and mites (Acari), a Malaysian perspective and review. Tropical Biomedicine, 25 (Supplement), 3–92.
Nadchatram, M., Domrow, R. & Ng, C.K. (1966) Parasitic Acarina of the mammals. Bulletin of the Singapore National Museum, 34, 129–140.
Nadchatram, M. & Kohls, G.M. (1963) Discovery of the tick Argas boueti Roubaud and Colas-Belcour (Acarina: Argasidae) in Malaya. Journal of Parasitology, 49, 350. https://doi.org/10.2307/3276013
Nakamura-Uchiyama, F., Komuro, Y., Yoshii, A. & Nawa, Y. (2000) Amblyomma testudinarium tick bite: one case of engorged adult and a case of extraordinary number of larval tick infestation. Journal of Dermatology, 27, 774–777. https://doi.org/10.1111/j.1346-8138.2000.tb02280.x
Nakao, Y., Tanigawa, T. & Shibata, R. (2017) Human otoacariasis caused by Amblyomma testudinarium: diagnosis and management: case report. Medicine(Baltimore), 96, 7394. https://doi.org/10.1097/MD.0000000000007394
Nakao, R., Shinjo, K., Sakiyama, T., Ogata, S., Kusakisako, K., Kinoshita, G., Naguib, D., Chatanga, E., Mohamed, W.M.A., Moustafa, M.A.M., Matsuno, K., Ito, T., Nonaka, N., Sashika, M., Tsubota, T. & Shimozuru, M. (2021) Amblyomma testudinarium infestation on a brown bear (Ursus arctos yesoensis) captured in Hokkaido, a northern island of Japan. Parasitology International, 80, 102209. https://doi.org/10.1016/j.parint.2020.102209
Nagahana, M. & Matsuo, K. (1962) Two tick species parasitic on the human body. Medical Biology, 62, 119–121.
Natsuaki, M., Takada, N., Kawabata, H., Sato, K. & Takano, A. (2013) Erythema migrans associated with Amblyomma testudinarium tick bite: tick-associated rash illness. Medical and Entomology Zoology, 64, 47–49. https://doi.org/10.7601/mez.64.47
Nava, S., Beati, L., Venzal, J.M., Labruna, M.B., Szabo, M.P.J., Petney, T., Saracho-Bottero, M.N., Tarragona, E.L., Santos Silva, M.M., Mangold, A.J., Guglielmone, A.A. & Estrada-Pena, A. (2018) Rhipicephalus sanguineus (Latreille, 1806): neotype designation, morphological re-description of all parasitic stages and molecular characterization. Ticks and Tick-borne Diseases, 9, 1573–1585. https://doi.org/10.1016/j.ttbdis.2018.08.001
Nava, S., Lareschi, M., Rebollo, C., Benítex Usher, L., Beati, L., Robbins, R.G., Durden, L.A., Mangold, A.J. & Guglielmone, A.A. (2007) The ticks (Acari: Ixodida: Argasidae, Ixodidae) of Paraguay. Annals of Tropical Medicine & Parasitology, 101, 255–270. https://doi.org/10.1179/136485907X176319
Navarrete-Abarca, L.R., Rodríguez-Romero, E.A., Valle-Martínez, C.A., Vargas-Artiga, M.J. & Pérez-Romero, L.E. (2014) Principales Especies de Garrapatas (Ixodidae) en El Salvador. Universidad de El Salvador, Facultad de Ciencias Agrarias, Departamento de MedicinaVeterinaria del Ministerio de Agricultura y Ganadería, San Salvador, 66 pp.
Need, J.T., Dale, W.E., Keirans, J.M. & Dasch, G.A. (1991) Annotated list of ticks (Acari: Ixodidae: Argasidae) reported in Peru: distribution, hosts, and bibliography. Journal of Medical Entomology, 28, 590–597. https://doi.org/10.1093/jmedent/28.5.590
Neumann, L.G. (1899) Révision de la famille des ixodidés (3e mémoire). Mémoires de la Société zoologique de France, 12, 107–294.
Neumann, L.G. (1911) Ixodidae. Das Tierreich, 26, 1–169. https://doi.org/10.5962/bhl.title.22339
Ng, Y.L., Hamdan, N.E.S., Tuen, A.A., Mohd-Azlan, J. & Chong, Y.L. (2017) Co-infections of ectoparasite species in synanthropic rodents of western Sarawak, Malaysian Borneo. Tropical Biomedicine, 34, 1–9.
Noh, Y.T. (1965) A taxonomic study on ticks in Korea (I) The genus Ixodes. Korean Journal of Zoology, 7, 73–76.
Norhidayu, S., Mohd-Zain, S.N., Jeffery, J. & Lewis, J.W. (2012) The dog louse Heterodoxus spiniger from stray cats in Penang, Malaysia. Tropical Biomedicine, 29, 301–303.
Nurkunasegran, M.D. (2017) Molecular survey of Coxiella burnetii in veterinary sample and ticks in Malaysia. Ph.D. dissertation. University of Malaya, Kuala Lumpur, 108 pp.
Nurkunasegran, M., Kho, K.L., Koh, F.X., Tan, P.E., Nizam, Q.N., Ong, B.L., Panchadcharam, C., Mat Amin, M., Abdul Majid, N., Ramli, R. & Tay, S.T. (2017) Molecular detection of Coxiella burnetii from farm animals and ticks in Malaysia. Tropical Biomedicine, 34, 675–680.
Nursyazana, M.T., Mohdzain, S.N. & Jeffery, J. (2013) Biodiversity and macroparasitic distribution of the wild rat population of Carey Island, Klang. Tropical Biomedicine, 30, 199–210.
Nuttall, G.H.F. (1916) Notes on ticks. IV. Relating to the genus Ixodes and including description of three new species and two varieties. Parasitology, 8, 294–337. https://doi.org/10.1017/S0031182000010623
Nuttall, G.H.F. & Warburton, C. (1911) Ticks, A Monograph of the Ixodoidea. Part II. Cambridge University Press, London, pp. 105–348.
Nuttall, G.H.F. & Warburton, C. (1915) Ticks, A Monograph of the Ixodoidea. Part III. Cambridge University Press, London, pp. 349–550.
Oiso, N., Nakano, A., Yano, Y. & Kawada, A. (2010) The diagnostic usefulness of dermoscopy for identifying six-legged larval tick. Ticks and Tick-borne Diseases, 1, 197–198. https://doi.org/10.1016/j.ttbdis.2010.09.005
Okino, T., Ushirogawa, H., Matoba, K. & Hatsushika, R. (2010) Bibliographical studies on human cases of hard tick (Acarina: Ixodidae) bites in Japan: (6) Cases of infestation with rare tick species. Kawasaki Medical Journal, 36, 121–126.
Okino, T., Ushirogawa, H., Matoba, K. & Hatsushika, R. (2012) Bibliographical studies on human cases of hard tick (Acarina: Ixodidae) bites in Japan: (8) Whole aspect of tick bite cases. Kawasaki Medical Journal, 38, 143–150.
Paddock, C.D. & Goddard, J. (2015) The evolving medical and veterinary importance of the Gulf Coast tick (Acari: Ixodidae). Journal of Medical Entomology, 52, 230–252. https://doi.org/10.1093/jme/tju022
Paramasvaran, S., Sani, R.A., Hassan, L., Krishnasamy, M., Jeffery, J., Oothuman, P., Salleh, I., Lim, K.H., Sumarni, M.G. & Santhana, R.L. (2009) Ectoparasite fauna of rodents and shrews from four habitats in Kuala Lumpur and the states of Selangor and Negeri Sembilan, Malaysia and its public health significance. Tropical Biomedicine, 26, 303–311.
Parrish, D. (1971) The occurrence, geographical distribution and wild vertebrate host relationships of ticks (Ixodoidea) on Luzon Island, Philippines, with descriptions of three new species Ph.D Dissertation, Oklahoma State University. Available from: https://core.ac.uk/download/pdf/215252358.pdf (accessed 14 September 2022)
Péter, A., Barti, L., Corduneanu, A., Hornok, S., Mihalca, A.D. & Sándor, A.D. (2021) First record of Ixodes simplex found on a human host, with a review of cases of human infestation by bat tick species occurring in Europe. Ticks and Tick-borne Diseases, 12, 101722. https://doi.org/10.1016/j.ttbdis.2021.101722
Petney, T.N. & Keirans, J.E. (1995) Ticks of the genera Amblyomma and Hyalomma (Acari: Ixodidae) in South-east Asia. Tropical Biomedicine, 12, 45–56.
Petney, T., Saijuntha, W., Boulanger, N., Chitimia-Dobler, I., Pfeffer, M., Eamudomkarn, C., Andrews, R.H., Ahamad, M., Putthasorn, N., Muders, S.V., Petney, D.A. & Robbins, R.G. (2019) Ticks (Argasidae, Ixodidae) and Tick-borne diseases of continental Southeast Asia. Zootaxa, 4558 (1), 1–89. https://doi.org/10.11646/zootaxa.4558.1.1
Phan, T.D., Nijhawan, S., Li, S. & Xiao, L. (2020) Capricornis sumatraensis. The IUCN Red List of Threatened Species. Available from: https://doi.org/10.2305/IUCN.UK.2020-2.RLTS.T162916735A162916910.en (accessed 13 September 2022)
Phan Trong, C. (1977) Ve bet va con trung ky sinh o Viet Nam. Tap 1 (Ixodoidea). Mo ta va phan loai, Hanoi, Vietnam, pp. 469–478.
Prakash, B.K. (2019) Molecular characterisation of Rhipicephalus sanguineus (Acari: Ixodidae) and occurrence of pathogens in ticks and host dogs. Ph.D. dissertion. Univeristy of Malaya, Kuala Lumpur, 95 pp.
Prakash, B.K., Low, V.L., Vinnie-Siow, W.Y., Tan, T.K., Lim, Y.A.L., Morvarid, A.R. & Sofian-Azirun, M. (2018) Detection of Babesia spp. in dogs and their ticks from Peninsular Malaysia: Emphasis on Babesia gibsoni and Babesia vogeli infections in Rhipicephalus sanguineus sensu lato (Acari: Ixodidae). Journal of Medical Entomology, 55, 1337–1340. https://doi.org/10.1093/jme/tjy072
Rageau, J. & Vervent, G. (1959) Les tiques (Acariens Ixodoidea) des ïles françaises du Pacifique. Bulletin de la Société de Pathologie Exotique, 52, 819–835.
Rahim, S., Ling, L.C., Singh, A.S.M. & Yunus, M.R.M. (2013) Facial nerve palsy from intra-aural tick (Ixodes holocyclus) infestation. Brunei International Medical Journal, 9, 382–384.
Rajagopalan, P.K. (1972) Ixodid ticks (Acarina: Ixodidae) parasitizing wild birds in the Kyasanur Forest disease area of Shimoga District, Mysore State, India. Journal of the Bombay Natural History Society, 69, 55–78.
Rajagopalan, P.K., Patil, A.P. & Boshell, M.J. (1968) Ixodid ticks on their mammalian hosts in the Kyasanur Forest disease area of Mysore State, India, 1961–64. Indian Journal of Medical Research, 56 (Supplement), 510–525.
Rajamanickam, C. (1968) Some notes on the ticks of domestic animals in West Malaysia and their distribution. A preliminary report. Kajian Veterinar, 1, 168–177.
Ramzan, M., Naeem-Ullah, U., Bokhari, S.H.M., Saba, S., Khan, K.A. & Saeed, S. (2020) Checklist of the tick (Acari: Argasidae, Ixodidae) species of Pakistan. Veterinaria Italiana, 56, 221–236.
Razali, N.B., Shamsudin, N., Rahaniza, A.M.J., Yaakop, S., Khoo, J.J. & Mohd-Taib, F.S. (2018) Ectoparasites (ticks and mites) prevalence on small to medium-sized mammals associated with habitat condition in Kemasul, Pahang. Serangga, 23, 72–88.
Rebello, M.J. & Reuben, R. (1967) A report on ticks collected from birds and small mammals in North Arcot and Chittoor Districts, South India. Journal of the Bombay Natural History Society, 68, 283–289.
Robbins, R.G., Phong, B.D., McCormack, T., Behler, J.L., Zwartepoorte, H.A., Hendrie, D.B. & Calle, P.P. (2006) Four new host records for Amblyomma geoemydae (Cantor) (Acari: Ixodida: Ixodidae) from captive tortoises and freshwater turtles (Reptilia: Testudines) in the Turtle Conservation Center, Cuc Phuong National Park, Vietnam. Proceedings of the Entomological Society of Washington, 108, 726–729.
Robbins, R.G. & Platt, S.G. (2000) First report of Amblyomma clypeolatum Neumann (Acari: Ixodida: Ixodidae) from the Union of Myanmar, with two new records from tortoises. Proceedings of the Entomological Society of Washington, 102, 225–226.
Rodríguez-Vivas, R.I., Apanaskevich, D.A., Ojeda-Chi, M.M., Trinidad-Martínez, I., Reyes-Novelo, E., Esteve-Gassent, M.D. & Pérez de León, A.A. (2016) Ticks collected from humans, domestic animals, and wildlife in Yucatán. Mexico. Veterinary Parasitology, 215, 106–113. https://doi.org/10.1016/j.vetpar.2015.11.010
Rohr, C.J. (1909) Estudos sobre Ixódidas do Brasil. Gómes Irmao, Rio de Janeiro, 220 pp.
Sahara, A., Nugraheni, Y.R., Patra, G., Prastowo, J. & Priyowidodo, D. (2019) Ticks (Acari: Ixodidae) infestation on cattle in various regions in Indonesia. Veterinary World, 12, 1755. https://doi.org/10.14202/vetworld.2019.1755-1759
Sajiuntha, W., Petney, T.N., Andrews, R.H. & Robbins, R.G. (2021) Ticks: a largely unexplored factor in disease transmission. In: Petney, T.N., Sajiuntha, W. & Mehlhorn, H. (Eds.), Biodiversity of southeast Asian parasites and vectors causing human disease. Springer Nature, Cham, pp. 165–182. https://doi.org/10.1007/978-3-030-71161-0_8
Sándor, A.D., Mihalca, A.D., Domsa, C., Péter, A. & Hornok, S. (2021) Argasid ticks of Palearctic bats: distribution, host selection, and zoonotic importance. Frontiers in Veterinary Science, 8, 684737. https://doi.org/10.3389/fvets.2021.684737
Santos Dias, J.A.T. (1959) Notas ixodológicas. VII. Sobre uma pequena colecção remetida pelo Dr. K. Lindberg. Memórias e Estudos do Museu Zoológico da Universidade de Coimbra, 261, 1–16.
Schulze, P. (1939) Zur Zeckenfauna Burmas. Zeitschrift für Parasitenkunde, 10, 722–728. https://doi.org/10.1007/BF02122034
Seddon, H.R. (1947) Influence of movement of animals of the spread of disease in Australia. Australian Veterinary Journal, 23, 200–212. https://doi.org/10.1111/j.1751-0813.1947.tb04564.x
Seneviratna, P. (1965) The Ixodoidea (ticks) of Ceylon. Parts II and III. Sri Lanka Veterinary Journal, 13, 28–54.
Seo, H.J., Noh, J., Kim, H.C., Chong, S.T., Klein, T.A., Park, C.U., Choi, C.Y., Kwon, Y.S., Kim, M., Park, Y., Yoo, M.S. & Cho, Y.S. (2021) Molecular detection and phylogenetic analysis of Anaplasma and Borrelia species in ticks collected from migratory birds at Heuksan, Hong, and Nan Islands, Republic of Korea. Vector-Borne and Zoonotic Diseases, 21, 20–31. https://doi.org/10.1089/vbz.2020.2629
Serra-Freire, N.M. (2014) Joined species of hard ticks on Amerindians, Quilombolas and domestic animals in the State of Paraiba. Revista UNIABEU, 7, 61–68.
Shabrina, M.S. (1990) Ectoparasites of small mammals trapped at the Ulu Gombak Forests, Selangor Darul Ehsan. Journal of Wildlife and Parks, 9, 9–17.
Sharifah, N., Heo, C.C., Ehlers, J., Houssaini, J. & Tappe, D. (2020) Ticks and tick-borne pathogens in animals and humans in the island nations of Southeast Asia: a review. Acta Tropica, 209, 105527. https://doi.org/10.1016/j.actatropica.2020.105527
Shariff, S.M. (1990) Ectoparasites of small mammals trapped at the Ulu Gombak Forest, Slenagor Darul Ehsan. The Journal of Wildlife and Parks, 9, 9–17.
Sharma, B.D. (1993) Medical and veterinary arthropod-disease ecology. Ashish Publishing House, New Delhi, 335 pp.
Shiang, L.F. (2019) Molecular detection of bacterial microbiome of ticks parasitizing wild boars (Sus scrofa) in an Orang Asli community. Master dissertation. Univesiti Malaya, Kuala Lumpur, 89 pp.
Singh, I., Kapila, C.C., Basu, S.M., Varma, R.N., Narsimhan, D., Rao, K.N.A., Sardana, D.N., Chopra, S.K. & Karani, M.P.D. (1965) Haemorrhagic disease following tick bites. Suspected toxoplasmosis. Lancet, 7390, 834–838. https://doi.org/10.1016/S0140-6736(65)91374-7
Silva, H. (1960) Prospecção Parasitológica Em Timor: Subsídios para o Estudo Da Fauna Parasitológica Dos Seus Animais Domésticos. Junta de Investigação do Ultramar. Estudos, ensaios e documentos, No. 76, 1–117.
Šlapeta, J., Chandra, S. & Halliday, B. (2021) The “tropical lineage” of the brown dog tick Rhipicephalus sanguineus sensu lato identified as Rhipicephalus linnaei (Audouin, 1826). International Journal for Parasitology, 51, 431–436. https://doi.org/10.1016/j.ijpara.2021.02.001
Šlapeta, J., Halliday, B., Chandra, S., Alanazi, A.D. & Abdel-Shafy, S. (2022) Rhipicephalus linnaei (Audouin, 1826) recognised as the “tropical lineage” of the brown dog tick Rhipicephalus sanguineus sensu lato: neotype designation, redescription, and establishment of morphological and molecular reference. Ticks and Tick-borne Diseases, 13, 102024. https://doi.org/10.1016/j.ttbdis.2022.102024
Smith, C.E.G. (1956) A virus resembling Russian Spring-Summer Encephalitis Virus from an Ixodid tick in Malaya. Nature, 178, 581–582. https://doi.org/10.1038/178581a0
Soundararajan, C., Nagarajan, K. & Prakash, M.A. (2018) Tick infestation in human beings in the Nilgiris and Kancheepuram District of Tamil Nadu, India. Journal of Parasitic Diseases, 42, 50–54. https://doi.org/10.1007/s12639-017-0963-x
Sreenivasan, M.A., Bhat, H.R. & Naik, S.V. (1979) Experimental transmission of Kyasanur Forest disease virus by Dermacentor auratus Supino. Indian Journal of Medical Research, 69, 701–707.
Strickland, C. & Roy, D.N. (1939) Ticks found on man. Indian Journal of Medical Research, 27, 251–252.
Sugimoto, M. (1935) On the Ixodoidea of Formosa. Taiwan Hakubutsu Gakkai Kaiho, 3, 1–7.
Sun, Y. & Xu, R. (2013) The genus Dermacentor and the subgenus Indocentor (Acari: Ixodidae) from China. Oriental Insects, 47, 155–168. https://doi.org/10.1080/00305316.2013.811019
Supino, F. (1897) Nuovi Ixodes della Birmaina (nota preventiva). Atti della Società veneto-trentina di scienze naturali, 2, 230–238. https://doi.org/10.5962/bhl.title.83719
Suzuki, N., Yamaguti, N., Saruta, T. & Hashiguchi, Y. (1990) Twelve cases of tick-bite among humans in Kochi Prefecture. Japanese Journal of Sanitary Zoology, 41, 71–74. https://doi.org/10.7601/mez.41.71
Takada, N., Fujita, H., Yano, Y., Oikawa, Y. & Mahara, F. (1992) Vectors of Japanese spotted fever. The Journal of the Japanese Association for Infectious Diseases, 66, 1218–1225. https://doi.org/10.11150/kansenshogakuzasshi1970.66.1218
Takahashi, M., Tamura, H., Baba, Y. & Misumi, H. (2022) First records of the hard tick Amblyomma testudinarium Koch, 1844 (Acari: Ixodiae) and larval trombiculid mite Eutrombicula wichmanni (Oudemans, 1905) (Acari: Trombiculidae) parasitizing sword-tail newt Cynops ensicauda (Hallowell, 1860) (Urodela: Salamandridae) in Japan. International Journal of Acarology, 48, 84–86. https://doi.org/10.1080/01647954.2021.2018042
Takhampunya, R., Thaloengsok, S., Tippayachai, B., Promsathaporn, S., Leepitakrat, S., Gross, K. & Davidson, S.A. (2021) Retrospective survey of Borrelia spp. from rodents and ticks in Thailand. Journal of Medical Entomology, 58, 1331–1344. https://doi.org/10.1093/jme/tjaa279
Tan, D.S.K., Smith, C.E.G., McMahon, D.A. & Bowen, E.T.W. (1967) Lanjan virus, a new agent isolated from Dermacentor auratus in Malaya. Nature, 214, 1154–1155. https://doi.org/10.1038/2141154a0
Tan, L.P., Choong, S.S., Samsuddin, A.S. & Lee, S.H. (2019) Amblyomma cordiferum Neumann, 1899 (Acari: Ixodidae) parasitizing reticulated pythons, Malayopython reticulatus (Schneider, 1801) (Reptilia: Pythonidae) in Peninsular Malaysia. Ticks and Tick-borne Diseases, 10, 101285. https://doi.org/10.1016/j.ttbdis.2019.101285
Tanskul, P.L. & Inlao, I. (1989) Keys to the adult ticks of Haemaphysalis Koch, 1844, in Thailand with notes on changes in taxonomy (Acari: Ixodoidea: Ixodidae). Journal of Medical Entomology, 26, 573–600. https://doi.org/10.1093/jmedent/26.6.573
Tanskul, P.L., Stark, H.E. & Inlao, I. (1983) A checklist of ticks of Thailand (Acari: Metastigmata: Ixodoidea) Journal of Medical Entomology, 20, 330–341. https://doi.org/10.1093/jmedent/20.3.330
Tappe, D., Gross, Y., Ngui, R., Rauch, J., Tay, S.T. & Lim, Y.A.L. (2019) High seroprevalence against typhus group and spotted fever group Rickettsiae in rural indigenous populations of Peninsular Malaysia. Vector-Borne and Zoonotic Diseases, 19, 323–327. https://doi.org/10.1089/vbz.2018.2391
Tay, S.T., Koh, F.X., Kho, K.L. & Ong, B.L. (2014) Molecular survey and sequence analysis of Anaplasma spp. in cattle and ticks in a Malaysian farm. Tropical Biomedicine, 31, 769–776.
Teng, K.F. & Jiang, Z.J. (1991) Economic insect fauna of China. Fascicule 39. Acari: Ixodidae. Science Press, Beijing, 355 pp.
Timmins, R., Kawanishi, K., Giman, B., Lynam, A., Chan, B., Steinmetz, R., Sagar Baral, H. & Samba Kumar, N. (2015) Sambar. Rusa unicolor. https://doi.org/10.2305/IUCN.UK.2015-2.RLTS.T41790A22156247.en
Tijjani, M. (2019) Identification of endoparasitic and ectoparasitic agents in rodents found in university colleges. Master thesis, Universiti Putra Malaysia, Serdang, Selangor. [unknown pagination]
Toft, II, J.D. & Eberhand, M.L. (1998) Parasitic diseases. In: Bennett, B.T., Abee, C.R. & Henrickson, R. (Eds.), Nonhuman primates in biomedical research: diseases. Academic Press, San Diego, pp. 1–852. https://doi.org/10.1016/B978-012088665-4/50005-1
Tominaga, C., Natsuaki, M. & Yamanishi, K. (2014) Study of tick bites in Hyogo Prefecture. Hifu no Hagaku, 13, 20–25.
Toumanoff, C. (1944) Les tiques (Ixodoidea) de l’Indochine. Institut Pasteur de l’Indochine. S.I.L.I., Ho Chi Minh City, 220 pp.
Trapido, H. (1965) Notes on critical Haemaphysalis specimens in European museum collections, with designations of lectotypes and a neotype. Proceedings of the Entomological Society of Washington, 67, 152–165.
Trapido, H., Hoogstraal, H. & Varma, M.G. (1964) Status and descriptions of Haemaphysalis p. papuana Thorell (n. comb.) and of H. papuana kinneari Warburton (n. comb.) (Ixodoidea, Ixodidae) of southern Asia and New Guinea. Journal of Parasitology, 50, 172–188. https://doi.org/10.2307/3276058
Tsunoda, T. (2004) Tick bite in researchers studying deer in Boso Peninsula, central Japan. Medical Entomology and Zoology, 55, 243–245. https://doi.org/10.7601/mez.55.243
Uchikawa, K. & Kobayashi, T. (1978) A contribution to the ectoparasite fauna of bats in Thailand. II. Blood-sucking Acari (Argasidae, Spinturnicidae and Macronyssidae). Contributions from the Biological Laboratory, Kyoto University, 25, 249–255.
Valente, J.D., Kakimori, M.T., Silva, P.W., Arzua, M., Barros-Battesti, D.M., Saldanha, A., Martini, R., Lange, D.R., Martins, T.F., Vieira, T.S.W.J., Labruna, M.B. & Vieira, R.F. (2022) Retrospective and new records of hard ticks (Acari: Ixodidae) on wild animals from Paraná State, southern of Brazil. Systematic and Applied Acarology, 27, 460–472. https://doi.org/10.11158/saa.27.3.5
van Peenan, P.F.D, Carney, W.P., Sudomo, M. & Sulianti-Saroso, J. (1974) Parasites of mammals in Gumbasa Valley, Central Sulawesi, Indonesia. Tropical and Geographical Medicine, 26, 352–358.
Varma, R.N. & Mahedevan, B. (1970) Ixodid ticks collected in the Eastern Himalayas and their potential disease relationships. Indian Journal of Medical Research, 58, 693–706.
Venzal, J.M., Autino, A.G., Nava, S. & Guglielmone, A.A. (2004) Ornithodoros mimon Kohls, Clifford & Jones, 1969 (Acari: Argasidae) on Argentinean bats, and new records from Uruguay. Systematic and Applied Acarology, 9, 37–39. https://doi.org/10.11158/saa.9.1.7
Voltzit, O.V. & Keirans, J.E. (2002) A review of Asian Amblyomma species (Acari, Ixodida, Ixodidae). Acarina, 10, 95–136.
Vongphayloth, K., Brey, P.T., Robbins, R.G. & Sutherland, I.W. (2016) First survey of the hard tick (Acari: Ixodidae) fauna of Nakai District, Khammouane Province, Laos, and an updated checklist of the ticks of Laos. Systematic and Applied Acarology, 21, 166–180. https://doi.org/10.11158/saa.21.2.2
Walker, J.B., Keirans, J.E. & Horak, I.G. (2000) The genus Rhipicephalus (Acari, Ixodidae): a guide to the brown ticks of the world. Cambridge University Press, Cambridge, 656 pp. https://doi.org/10.1017/CBO9780511661754
Wassef, H.Y. & Hoogstraal, H. (1983) Dermacentor (Indocentor) compactus (Acari: Ixodoidea: Ixodidae): identity of male and female. Journal of Medical Entomology, 20, 648–652. https://doi.org/10.1093/jmedent/20.6.648
Wassef, H.Y. & Hoogstraal, H. (1984) Dermacentor (Indocentor) auratus (Acari: Ixodoidea: Ixodidae): identity of male and female. Journal of Medical Entomology, 21, 169–173. https://doi.org/10.1093/jmedent/21.2.169
Wassef, H.Y. & Hoogstraal, H. (1986) Dermacentor (Indocentor) steini (Acari: Ixodoidea: Ixodidae): identity of male and female. Journal of Medical Entomology, 23, 532–537. https://doi.org/10.1093/jmedent/23.5.532
Wassef, H.Y. & Hoogstraal, H. (1988) Dermacentor (Indocentor) steini (Acari: Ixodoidea: Ixodidae): hosts, distribution in the Malay Peninsula, Indonesia, Borneo, Thailand, the Philippines, and New Guinea. Journal of Medical Entomology, 25, 315–320. https://doi.org/10.1093/jmedent/25.5.315
Watanabe, M., Nakao, R., Amin-Babjee, S.M., Maizatul, A.M., Youn, J.H., Qiu, Y., Sugimoto, C. & Watanabe, M. (2015) Molecular screening for Rickettsia, Anaplasmataceae and Coxiella burnetii in Rhipicephalus sanguineus ticks from Malaysia. Tropical Biomedicine, 32, 390–398.
Wells, K., Beaucournu, J.C., Durden, L.A., Petney, T.N., Lakim, M.B. & O’Hara, R.B. (2012) Ectoparasite infestation patterns of domestic dogs in suburban and rural areas in Borneo. Parasitology Research, 111, 909–919. https://doi.org/10.1007/s00436-012-2917-7
Wells, K., O’Hara, R.B., Pfeiffer, M., Lakim, M.B., Petney, T.N. & Durden, L.A. (2013) Inferring host specificity and network formation through agent-based models: tick–mammal interactions in Borneo. Oecologia, 172, 307–316. https://doi.org/10.1007/s00442-012-2511-9
Yamada, Y., Dekio, S., Jidoi, J., Isobe, A., Shiwaku, K. & Yamane, Y. (1996) A case of tick bite from Amblyomma testudinarium on the glans penis. Journal of Dermatology, 23, 136–138. https://doi.org/10.1111/j.1346-8138.1996.tb03986.x
Yamamoto, S., Fukuzaki, M. & Matsushita, T. (1997) A human case of simultaneous tick bites by Ixodes acutitarsus and Ixodes monospinosus in Kyushu, Japan. Medical Entomology and Zoology, 48, 69–71. https://doi.org/10.7601/mez.48.69
Yamaguti, N. (1989) Human tick bite: variety of tick species and increase of cases. Saishin-Igaku, 44, 903–908.
Yamaguti, N. & Takada, N. (1981) Thirty nine case reports of human infestation with ixodid ticks. Japanese Journal of Sanitary Zoology, 32, 86–89. https://doi.org/10.7601/mez.32.86
Yamaguti, N., Tipton, V.J., Keegan, H.L. & Toshioka, S. (1971) Ticks of Japan, Korea, and the Ryukyu Islands. Brigham Young University Science Bulletin, Biological Series, 15, 1–185. https://doi.org/10.5962/bhl.part.25691
Yamauchi, T. (2001) A bibliographical survey of host-parasite relationships between birds and ticks from Japan. Hoshizaki Gurîn Zaidan kenkyű hôkoku, 5, 271–308.
Yamauchi, T., Fukui, Y., Watanabe, M., Nakagawa, H. & Kamimura, K. (2010a) Forty cases of human infestations with hard ticks (Acari: Ixodidae) in Toyama Prefecture, Japan. Medical Entomology and Zoology, 61, 133–143. https://doi.org/10.7601/mez.61.133
Yamauchi, T. & Nakatani, T. (2016) Additional 10 cases of human infestation with hard ticks in Toyama Prefecture, Japan, with two cases of Amblyomma testudinarium (Acari: Ixodidae), in Japan. Medical Entomology and Zoology, 67, 239–242. https://doi.org/10.7601/mez.67.239
Yamauchi, T., Shimazu, Y. & Mizuta, H. (2009) A case of human tick bite by a nymphal tick, Haemaphysalis hystricis (Acari: Ixodidae), in Japan. Medical Entomology and Zoology, 60, 135–137. https://doi.org/10.7601/mez.60.135
Yamauchi, T., Takano, A., Maruyama, M. & Kawabata, H. (2012) Human infestation by Amblyomma testudinarium (Acari: Ixodidae) in Malay Peninsula, Malaysia. Journal of the Acarological Society of Japan, 21, 143–148. https://doi.org/10.2300/acari.21.143
Yamauchi, T., Takano, A., Sakata, A., Baba, S., Okushima, Y. & Kawabata, H. (2010b) Five cases of human infestations by Amblyomma testudinarium (Acari: Ixodidae) in Japan. Journal of the Acarological Society of Japan, 19, 15–21. https://doi.org/10.2300/acari.19.15
Yamauchi, T., Yano, S., Yamamoto, E. & Miyamoto, T. (2013) Ticks (Acari: Ixodidae) from medium-sized to large mammals in Ehima Prefecture, Japan. Experimental and Applied Acarology, 60, 263–270. https://doi.org/10.1007/s10493-012-9643-9
Yano, Y., Takada, N., Ootsuki, N., Sakai, H., Imamura, Y. & Masuzawa, T. (1998) Epidemiological discussion on multi-infestation with larval tick in human. Journal of the Acarological Society of Japan, 7, 145–148. https://doi.org/10.2300/acari.7.145
Yoneda, Y., Gouhara, S., Mitarai, Y. & Fukuma, T. (1997) Seventeen human cases infested with ixodid ticks in Kyushu, Japan. Medical Entomology and Zoology, 48, 269–274. https://doi.org/10.7601/mez.48.269
Yun, S.M., Lee, W.G., Ryou, J., Yang, S.C., Park, S.W., Roh, J.Y., Lee, Y.J., Park, C. & Han, M.G. (2014) Severe fever with thrombocytopenia syndrome virus in ticks collected from humans, South Korea. Emerging Infectious Diseases, 20, 1358–1361. https://doi.org/10.3201/eid2008.131857
Zaini, Z.H., Abu, J. & Amin-Babjee, S.M. (2016) Comparison of parasite species and burden in apparently healthy and clinically ill red junglefowls (Gallus gallus). 11th Proceedings of the Seminar on Veterinary Sciences, 29 February–2 March 2016. [unknown pagination]
Zhang, Y.K., Zhang, X.Y. & Liu, J.Z. (2019) Ticks (Acari: Ixodoidea) in China: Geographical distribution, host diversity, and specificity. Archives of Insect Biochemistry and Physiology, 102, e21544. https://doi.org/10.1002/arch.21544