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Published: 2022-11-28
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Checklist of water mites from mainland Portugal and its archipelagos

Interdisciplinary Centre of Marine and Environmental Research (CIIMAR), University of Porto, Terminal de Cruzeiros do Porto de Leixões, Avenida General Norton de Matos, s/n, 4450-208, Matosinhos, Porto, Portugal.
Interdisciplinary Centre of Marine and Environmental Research (CIIMAR), University of Porto, Terminal de Cruzeiros do Porto de Leixões, Avenida General Norton de Matos, s/n, 4450-208, Matosinhos, Porto, Portugal. 2Aquamuseu do Rio Minho, Vila Nova de Cerveira, 4920-290, Portugal.
Interdisciplinary Centre of Marine and Environmental Research (CIIMAR), University of Porto, Terminal de Cruzeiros do Porto de Leixões, Avenida General Norton de Matos, s/n, 4450-208, Matosinhos, Porto, Portugal. 2Aquamuseu do Rio Minho, Vila Nova de Cerveira, 4920-290, Portugal.
Interdisciplinary Centre of Marine and Environmental Research (CIIMAR), University of Porto, Terminal de Cruzeiros do Porto de Leixões, Avenida General Norton de Matos, s/n, 4450-208, Matosinhos, Porto, Portugal. Faculdade de Ciências, Universidade do Porto, rua do Campo Alegre s/n, 4169-007 Porto, Portugal
Interdisciplinary Centre of Marine and Environmental Research (CIIMAR), University of Porto, Terminal de Cruzeiros do Porto de Leixões, Avenida General Norton de Matos, s/n, 4450-208, Matosinhos, Porto, Portugal. 2Aquamuseu do Rio Minho, Vila Nova de Cerveira, 4920-290, Portugal.
Acari Taxonomy Portuguese Exclusive Economic Zone Iberian Peninsula North Atlantic Ocean

Abstract

This study focuses on aquatic mites from the Portuguese Exclusive Economic Zone, including the mainland area, and Madeira and Azores archipelagos, aiming at compiling all existing information and generating a list of species and all their data on collecting sites. For the present work, we consider aquatic species, those that inhabit the marine, deep sea, intertidal, freshwater, transition water or brackish environments, with life-cycle occurring entirely at water habitats or their vicinity. Furthermore, breeding should take place in water aquatic lifestyles. The dataset search for this checklist started with the first descriptions of water acarids from the Azores Archipelago performed by Théodore Barrois. Primary sources for occurrence confirmation were Süßwasserfauna von Mitteleuropa book collection, The Global Biodiversity Information Facility (GBIF), complemented with an extensive Portuguese literature review and data biodiversity websites, e.g., European Water Mite Research (Watermite.org). In total, 102 valid species were found belonging to 41 genera, and 19 families, nested in the orders Mesostigmata, Sarcoptiformes and Trombidiformes. This is the first work compiling all the species of aquatic mites found in Portuguese territory, as well as data of collection, serving as the basis for future studies.

 

References

  1. Angelier, E. (1949) Diagnoses sommaires d’Hydracariens phréaticoles nouveaux. Bulletin du Museìum National d’Histoire Naturelle, 21 (Supplement 2), 88–90.
    Angelier, E., Angelier, M.-L. & Lauga, J. (1985) Research on the ecology of water-mites (Hydrachnellae, Acari) in flowing water. Annales de Limnologie, 21, 25–64. https://doi.org/10.1051/limn/1985003
    AQUILA (2022) AQUILA. Senckenberg-Museum Frankfurt. Available from: https://www.senckenberg.de/en/science/research-infrastructure/databases-and-digital-resources/aquila/ (accessed 8 November 2022)
    Barrois, T. (1887) I. Hydrachnides. In: Barrois, T. (Ed.), Matériaux pour servir a l’étude de la faune des eaux douces des Açores. Imprimerie Le Bigot Frères, Lille, pp. 1–16.
    Barrois, T. (1896) Recherches sur la faune des eaux douces des Açores. Mémoires de la Société des sciences, de l’agriculture et des arts de Lille, 5, 1–172. https://doi.org/10.3406/bsef.1896.21760
    Barrois, T. & Moniez, R. (1887) Catalogue des Hydrachnides recueilles dans le nord de la France avec des notes critiques et la description d`espèces nouvelles. Le Bigot Frères, Lille, 36 pp. [http://www.watermite.org/publication/detail.php?id=156]
    Bartsch, I. (1978) Halacaridae (Acari) Aus Der Tiefsee Des Atlantischen Ozeans. Cahiers de Biologie Marine, 19, 47–62.
    Bartsch, I. (1988) Halacaridae (Acari) in the Northem Atlantic Ocean. Description of Halacarus balgimus sp. n. and Agauides cryosi gen. and sp. n. Cahiers de Biologie Marine, 29, 353–359. https://doi.org/10.21411/CBM.A.3E02A4CC
    Bartsch, I. (1994) Halacarid mites (Acari) from hydrothermal deep-sea sites. New records. Cahiers de Biologie Marine, 35, 479–490. https://doi.org/10.21411/CBM.A.72133B3
    Bartsch, I. (1996) Halacarid mites (Acari) from the Mid-Atlantic Ridge. New records. Cahiers de Biologie Marine, 37, 159–167. https://doi.org/10.21411/CBM.A.2616F26B
    Behan-Pelletier, V., Cook, D.R., Harvey, M.S., Keirans, J.E., Krantz, G.W., Lindquist, E.E., Norton, R.A., OConnor, B.M., Smith, I.M. & Walter, D.E. (2009) s.n. In: Krantz, G.W. & Walter, D.E. (Eds.), A manual of acarology. 3rd Edition. Texas Tech University Press, Lubbock, pp. i–viii + 1–807. [https://www.worldcat.org/title/manual-of-acarology/oclc/904205537?referer=di&ht=edition]
    Besch, W. (1964) Systematik und verbreitung der Südamerikanischen rheophilen hydrachnellen. Beitrage zur Neotropischen Fauna, 3, 77–194. https://doi.org/10.1080/01650526409360369
    Besseling, A.J. (1942) Nederlandsche Hydrachnidae. Genus Hygrobates. Entomologische Berichten, 11, 2–6.
    Blott, S.J. & Pye, K. (2001) GRADISTAT: a grain size distribution and statistics package for the analysis of unconsolidated sediments. Earth Surface Processes and Landforms, 26, 1237–1248. https://doi.org/10.1002/esp.261
    Borges, P.A.V., Abreu, C., Aguiar, A.M.F., Carvalho, P., Jardim, R., Melo, I., Oliveira, P., Sérgio, C., Serrano, A.R.M. & Vieira, P. (2008) A list of the terrestrial fungi, flora and fauna of Madeira and Selvagens archipelagos. Direcção Regional do Ambiente da Madeira and Universidade dos Açores, Funchal and Angra do Heroísmo, Ponta Delgada, 440 pp. [https://issuu.com/institutodasflorestaseconservacaoda/docs/listfunffterrmadsel]
    Böttger, K. (1969) Wassermilben als Räuber an Trichopteren-Laich. Naturwissenschaften, 56, 333–333. https://doi.org/10.1007/BF00602188
    Camacho, A.I. & Valdecasas, A.G. (1988) Spatial and biocenotic trends in the water-mite fauna of small ponds. Hydrobiologia, 160, 207–217. https://doi.org/10.1007/BF00007136
    Canestrini, G. & Canestrini, R. (1881) Nuove specie del genere Gamasus. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, 5, 1077–1086.
    Cantallo, H., Gomes, N., Antunes, C., Ribeiro, T., Gomes, M.I. & Costa, D.A. (2021) Catalogue of aquatic mites from the Minho River (NW Iberian Peninsula). Environmental Smoke, 2021, 44–47. https://doi.org/10.32435/envsmoke/xibesymp.7
    Castilho, R.C., Silva, E.S., De Moraes, G.J. & Halliday, B. (2016) Catalogue of the family Ologamasidae Ryke (Acari: Mesostigmata). Zootaxa, 4197 (1), 1–147. https://doi.org/10.11646/zootaxa.4197.1.1
    Coggi, A. (1897) Una nuova specie di Oribatide. In: Canestrini, G. (Ed.), Prospetto Dell’ Acarofauna Italiana. Parte VIII.a. Stabilimento Prosperini, Padova, pp. 916–921.
    Costa-Garcia, R. (2018) Influence of urbanisation on an intertidal ecosystem engineer: Chthamalus barnacles and their associated epifauna. U. Porto. Available from: https://sigarra.up.pt/icbas/pt/pub_geral.show_file?pi_doc_id=182009 (accessed 8 November 2022)
    Dabert, M., Witalinski, W., Kazmierski, A., Olszanowski, Z. & Dabert, J. (2010) Molecular phylogeny of acariform mites (Acari, Arachnida): Strong conflict between phylogenetic signal and long-branch attraction artifacts. Molecular Phylogenetics and Evolution, 56, 222–241. https://doi.org/10.1016/j.ympev.2009.12.020
    Van Dam, M.H., Trautwein, M., Spicer, G.S. & Esposito, L. (2019) Advancing mite phylogenomics: Designing ultraconserved elements for Acari phylogeny. Molecular Ecology Resources, 19, 465–475. https://doi.org/10.1111/1755-0998.12962
    Davids, K., Bartsch, I., Deichsel, R., Di Sabatino, A., Gabrys, G., Gledhill, T., Jäger, P., Makol, J., Smit, H., van der Hammen, H., Weigmann, G., Wohltmann, A. & Wurst, E. (2006) s.n. In: Gerecke, R. (Ed.), Süßwasserfauna von Mitteleuropa. Vol. 7/2-1 Chelicerata: Araneae/Acari I. 1st Edition. Springer, Berlin and Heidelberg, pp. I–X + 1–388. https://doi.org/10.1007/978-3-662-55958-1
    Davids, K., Di Sabatino, A., Gerecke, R., Gledhill, T. & Smit, H. (2005) On the taxonomy of water mites (Acari: Hydrachnidia) described from the Palaearctic, part 1: Hydrachnidae, Limnocharidae and Eylaidae. Zootaxa, 1061 (1), 35–64. https://doi.org/10.11646/zootaxa.1061.1.3
    Domes, K., Althammer, M., Norton, R.A., Scheu, S. & Maraun, M. (2007) The phylogenetic relationship between Astigmata and Oribatida (Acari) as indicated by molecular markers. Experimental and Applied Acarology, 42, 159–171. https://doi.org/10.1007/s10493-007-9088-8
    Dugès, A. (1834) Deuxième mémoire sur l`ordre des Acariens. Recherches sur l`ordre des Acariens: Rémarques sur la famille des Hydracnés. Annales des Sciences Naturelles, 1, 144–174.
    Erman, O., Pešić, V., Esen, Y. & Özkan, M. (2010) A checklist of the water mites of Turkey (Acari: Hydrachnidia) with description of two new species. Zootaxa, 2624 (1), 1. https://doi.org/10.11646/zootaxa.2624.1.1
    Esen, Y., Cihan, Ç. & Erman, O. (2011) A new species of the family Unionicolidae Oudemans, 1909 (Acari, Hydrachnidia) for the Turkish fauna. Turkish Journal of Science & Technology, 6, 111–116.
    García-Jiménez, R., Horreo, J.L. & Valdecasas, A.G. (2017) Minimal barcode distance between two water mite species from Madeira Island: a cautionary tale. Experimental and Applied Acarology, 72, 133–143.
    https://doi.org/10.1007/s10493-017-0147-5
    GBIF (2022) GBIF—Occurrences. Global Biodiversity Information Facility. Available from: https://www.gbif.org/occurrence/search/ (accessed 8 January 2022)
    Gerecke, R. (1991) Taxonomische, faunistische und ökologische Untersuchungen an Wassermilben (Acari, Actinedida) aus Sizilien unter Berücksichtigung anderer aquatischer Invertebraten. Lauterbornia, 7, 1–303.
    Gerecke, R. (2003) Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141–378. https://doi.org/10.1046/j.1096-3642.06-0.00051.x
    Gerecke, R. (2009) Revisional studies on the European species of the water mite genus Lebertia NEUMAN, 1880 (Acari: Hydrachnidia: Lebertiidae). Abhandlungen der Senckenberg Gesellschaft für Naturforschung, 566, 1–144.
    Gerecke, R. (2017) The Water Mites Of The Genus Hydrodroma (Acari, Hydrachnidia, Hydrodromidae) In Europe And Africa. Ecologica Montenegrina, 13, 1–24. https://doi.org/10.37828/em.2017.13.1
    Gerecke, R., Gledhill, T., Pešić, V. & Smit, H. (2016) Süßwasserfauna von Mitteleuropa. Bd. 7/2-3. Chelicerata: Acari III. Band 7/2-3. Springer Spektrum, Heidelberg, Dordrecht, London and New York, pp. i–xi + 1–429. [http://link.springer.com/10.1007/978-3-8274-2689-5]
    Giribet, G. (2018) Current views on chelicerate phylogeny—A tribute to Peter Weygoldt. Zoologischer Anzeiger, 273, 7–13. https://doi.org/10.1016/j.jcz.2018.01.004
    Green, J. & Macquitty, M. (1987) Halacarid mites. In: Kermack, D.M. & Barnes, R.S.K. (Eds.), Synopses of the British Fauna. New Series. No. 36. Field Studies Council, Linnean Society of London, Estuarine and Coastal Sciences Association, Avon, pp. vii+178.
    Halbert, J.N. (1920) The Acarina of the Seashore. Proceedings of the Royal Irish Academy, Section B: Biological, Geological, and Chemical Science, 35, 106–152.
    Van Der Hammen, H. & Smit, H. (1996) The water mites (Acari: Hydrachnidia) of streams in The Netherlands: Distribution and ecological aspects on a regional scale. Netherlands Journal of Aquatic Ecology, 30, 175–185. https://doi.org/10.1007/BF02272237
    Hayward, P.J. & Ryland, J.S. (2017) s.n. In: Hayward, P.J. & Ryland, J.S. (Eds.), Handbook of the marine fauna of North-West Europe. 2nd Edition. Oxford University Press, New York, pp. 1–785. [https://oxford.universitypressscholarship.com/view/10.1093/acprof:oso/9780199549443.001.0001/acprof-9780199549443] https://doi.org/10.1093/acprof:oso/9780199549443.001.0001
    Hermann, J.F. (1804) s.n. In: Hammer, F.-L. (Ed.), Meìmoire Apteìrologique. Levrault, F.G., Strasbourg, pp. 1–144, pl. IX. [http://www.biodiversitylibrary.org/bibliography/137910] https://doi.org/10.5962/bhl.title.137910
    Hughes, S.J., Furse, M.T., Blackburn, J.H. & Langton, P.H. (1998) A checklist of Madeiran freshwater macroinvertebrates. Boletim do Museu Municipal do Funchal, 50, 5–41.
    Koch, C.L. (1837) Deutschlands Crustaceen, Myriapoden und Arachniden: Ein Beitrag zur deutschen Fauna. Band 10. In: Deutschlands Insekten (continuation of Panzer, Faunae Inscetorum Germaniae Initia [= Panzer, 146]). Friedrich Pustet, Regensburg. [unknown pagination]
    Koenike, F. (1886) Zwei neue Hydrachniden aus dem Isergebirge. Zeitschrift für wissenschaftliche Zoologie, 43, 277–284.
    Koenike, F. (1895a) I. Wissenschaftliche Mittheilungen. 1. Über bekannte und neue Wassermilben. Zoologischer Anzeiger, 18, 373–386.
    Koenike, F. (1895b) Neue Sperchon-Arten aus der Schweiz. Revue Suisse de Zoologie 3, 415–427. https://doi.org/10.5962/p.318237
    Koenike, F. (1902) Acht neue Lebertia-Arten, eine Arrenurus- und eine neue Atractides-Art. (Vorläufige Mittheilung). Zoologischer Anzeiger, 25, 610–616.
    Koenike, F. (1903) Vier unbekannte norddeutsche Hydrachniden. Zoologischer Anzeiger, 26, 534–536.
    Koenike, F. (1907) Zwei unbekannte Sperchoniden und eine Curvipes-Spezies. Abhandlungen herausgegeben vom Naturwissenschaftlichen Verein zu Bremen, 19, 133–138. https://doi.org/10.5962/bhl.part.16693
    Koenike, F. (1911) Sechs neue norddeutsche Wassermilben. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen, 20, 236–256.
    Kramer, P. (1879) Neue Acariden. Archiv für Naturgeschichte, 45, 1–18.
    Linnaeus, C. (1758) Systema naturae per regna tria naturae :secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio dec. Impensis Direct. Laurentii Salvii, Holmiæ [Stockholm], 824 pp. [http://www.biodiversitylibrary.org/bibliography/542] https://doi.org/10.5962/bhl.title.542
    Lundblad, O. (1941a) Neue Wassermilben. Vorläufige Mitteilung. Entomologisk Tidskrift, 62, 97–121.
    Lundblad, O. (1941b) Neue Wassermilben aus Madeira: Vorläufige Mitteilung. Entomologisk Tidskrift, 62, 93–96.
    Lundblad, O. (1942) Die Arthropodenfauna von Madeira nach den Ergebnissen der Reise von Prof. Dr. O. Lundblad Juli-August 1935. 31. Hydrachnellae. Arkiv för Zoologi, 5, 1–122.
    Lundblad, O. (1954) Neue südeuropäische Wassermilben. Zoologischer Anzeiger, 153, 154–159.
    Lundblad, O. (1956) Zur Kenntnis süd- und mitteleuropäischer Hydrachnellen. Arkiv för Zoologi 10, 1–306, tafel LXXXIV.
    Lundblad, O. (1962) Wassermilben von den Kanarischen Inseln. Arkiv för Zoologi, 15, 285–300.
    Maglio, C. (1924) Idracarini. In: Monti, R. (Ed.), La Limnologia del Lario in relazione al ripopolamento delle acque ed alla pesca. Luigi Luzzatti, Rome, pp. 75–124.
    Martin, P. (2009) A redescription of Torrenticola (Megapalpis) fagei (E . Angelier, 1949) (Acari, Hydrachnidia). Lauterbornia, 67, 183–188.
    Michael, A.D. (1895) A study of the internal anatomy of Thyas petrophilus, an unrecorded Hydrachnid found in Cornwall. Proceedings of the Zoological Society of London, 63, 174–209. https://doi.org/10.1111/j.1469-7998.1895.tb00003.x
    Migot, A. (1926) Diagnoses sommaires d’Hydracariens nouveaux des Pyrénées. Bulletin de la Société Entomologique de France, 41, 52–56. https://doi.org/10.3406/bsef.1926.27628
    Müller, O.F. (1776) s.n. In: Müller, O.F. (Ed.), Zoologiæ Danicæ Prodromus, seu Animalium Daniæ et Norvegiæ indigenarum characteres, nomina, et synonyma imprimis popularium. Hallageriis, Havniae, pp. i –xxxii + 1–282. [http://www.biodiversitylibrary.org/item/47550] https://doi.org/10.5962/bhl.title.13268
    Norton, R.A. (1994) Evolutionary Aspects of Oribatid Mite Life Histories and Consequences for the Origin of the Astigmata. In: Houck, M.A. (Eds.), Mites, Springer, Boston, Massachusetts, pp. 99–135. https://doi.org/10.1007/978-1-4615-2389-5_5
    Norton, R.A. (1998) Morphological evidence for the evolutionary origin of Astigmata (Acari: Acariformes). Experimental and Applied Acarology, 22, 559–594. https://doi.org/10.1023/A:1006135509248
    NRM (2022) Entomologi samlingsetiketter [Entomological Collections] (NHRS), Naturhistoriska Riksmuseet [Swedish Museum of Natural History] (NRM). Available from: https://naturarv.nrm.se/ (accessed 14 June 2022)
    Peláez, M.L., Horreo, J.L., García-Jiménez, R. & Valdecasas, A.G. (2022) An evaluation of errors in the mitochondrial COI sequences of Hydrachnidia (Acari, Parasitengona) in public databases. Experimental and Applied Acarology, 86, 371–384. https://doi.org/10.1007/s10493-022-00703-0
    Pepato, A.R. & Klimov, P.B. (2015) Origin and higher-level diversification of acariform mites - Evidence from nuclear ribosomal genes, extensive taxon sampling, and secondary structure alignment. BMC Evolutionary Biology, 15, 1–20. https://doi.org/10.1186/s12862-015-0458-2
    Pérez-Íñigo, C. (1997) Acari: Oribatei, Gymnonota I. In: Sánchez, M.A.R., Tercedor, J.A., Ros, X.B., Noguera, J.G., Sierra, A.G., Mayol, E.M., Piera, F.M., Marino, J.S. & González, J.T. (Eds.), Fauna Iberica. Vol. 9. Museo Nacional de Ciencias Naturales, Consejo Superior de Investigaciones Científicas (CSIC), Madrid, pp. 374.
    Pešić, V. (1999) Contribution to the knowledge of Hydracarina (water mites) of Montenegro. Poljoprivreda i Šumarstvo, 45, 83–96.
    Pešić, V., Asadi, M., Cimpean, M., Dabert, M., Esen, Y., Gerecke, R., Martin, P., Savić, A., Smit, H. & Stur, E. (2017) Six species in one: evidence of cryptic speciation in the Hygrobates fluviatilis complex (Acariformes, Hydrachnidia, Hygrobatidae). Systematic and Applied Acarology, 22, 1327. https://doi.org/10.11158/saa.22.9.4
    Pešić, V., Broda, L., Dabert, M., Gerecke, R., Martin, P. & Smit, H. (2019) Re-established after hundred years: Definition of Hygrobates prosiliens Koenike, 1915, based on molecular and morphological evidence, and redescription of H. longipalpis (Hermann, 1804) (Acariformes, Hydrachnidia,. Systematic and Applied Acarology, 24, 1490–1511. https://doi.org/10.11158/saa.24.8.10
    Pešić, V., Gerecke, R. & Cîmpean, M. (2007) Water mites of the genus Neumania Lebert (Acari, Hydrachnidia: Unionicolidae: Pionatacinae) in the Mediterranean area. Annales de Limnologie - International Journal of Limnology, 43, 187–198. https://doi.org/10.1051/limn:2007013
    Pešić, V., Saboori, A. & Asadi, M. (2004) New records of water mite species (Acari, Hydrachnidia) from Iran, with the description of one new species. Systematic and Applied Acarology, 10, 137–147. https://doi.org/10.11158/saa.10.1.13
    Pešić, V. & Smit, H. (2022) Water mites of Corsica: DNA barcode and morphological evidences. International Journal of Acarology 48, 418–428. https://doi.org/10.1080/01647954.2022.2086619
    Piersig, R. (1894) Über Hydrachniden. Zoologischer Anzeiger, 17, 107–111 + 113–118.
    Piersig, R. (1896) Einige neue Hydrachniden-Formen. (Vorläufige Mittheilung.). Zoologischer Anzeiger, 19, 438–441.
    Piersig, R. (1898) 3. Hydrachnidenformen aus den deutschen Mittelgebirgen. Zoologischer Anzeiger, 21, 451–452.
    Piersig, R. (1903) Neues Verzeichnis der bisher im Sächsischen Erzgebirge aufgefundenen Hydrachniden-Formen. Bericht über den Annaberg-Buchholzer Verein für Naturkunde, 11, 34–46.
    Plazi.org (2022) TreatmentBank. Available from: http://plazi.org/treatmentbank/ (accessed 6 July 2022)
    Protz, A. (1896) Beiträge zur Hydrachnidenkunde: Zwei neue Hydrachniden-Arten aus der Umgegend von Berlin. Zoologischer Anzeiger, 19, 23–26.
    Quaglia, M.M. & Conroy, J.C. (1984) New or interesting water mites (Acari, Hydrachnellae) in drift samples from the torch river, Saskatchewan. International Journal of Acarology 10, 89–101. https://doi.org/10.1080/01647958408683358
    Di Sabatino, A., Gerecke, R., Gledhill, T. & Smit, H. (2009) On the taxonomy of water mites (Acari: Hydrachnidia) described from the Palaearctic, part 2: Hydryphantoidea and Lebertioidea. Zootaxa, 2266 (1), 1–34. https://doi.org/10.11646/zootaxa.2266.1.1
    Di Sabatino, A., Gerecke, R., Gledhill, T. & Smit, H. (2010) Süßwasserfauna von Mitteleuropa, Bd. 7/2-2, Chelicerata: Acari II. In: Gerecke, R. (Ed.), Spektrum Akademischer Verlag, Heidelberg, Heidelberg, pp. i–ix + 1–234. [https://link.springer.com/book/9783827418944] https://doi.org/10.1007/978-3-8274-2266-8_1
    Schatz, H. & Behan-Pelletier, V. (2008) Global diversity of oribatids (Oribatida: Acari: Arachnida). Hydrobiologia, 595, 323–328. https://doi.org/10.1007/s10750-007-9027-z
    Schatz, H., Behan-Pelletier, V.M., OConnor, B.M. & Norton, R.A. (2011) Suborder Oribatida van der Hammen, 1968. Zootaxa, 3148 (1), 141–148. https://doi.org/10.11646/zootaxa.3148.1.26
    Schaub, R.V. (1888) Über die Anatomie von Hydrodroma (C. L. Koch): Ein Beitrag zur Kenntniss der Hydrachniden. Sitzungsberichte Abt. 1 Mineralogie, Krystallographie, Botanik, Physiologie der Pflanzen, Zoologie, Paläontologie, Geologie, physische Geographie und Reisen, 97, 98–151.
    Schrank, F.P. (1781) Enumeratio insectorum Austriae indigenorum. Apvd vidvam Eberhardi Klett et Franck, Augusta Vindelicorum [actual Augsburg], 548 pp. [https://www.biodiversitylibrary.org/bibliography/15091]
    Schwoerbel, J. (1957) Zur Kenntnis der Wassermilbenfauna des südlichen Schwarzwaldes (Hydrachnellae, Acari). 3. Beitrag. Mitteilungen des Badischen Landesvereins für Naturkunde und Naturschutz, Neue Folge, 7, 41–52.
    Shultz, J.W. (2007) A phylogenetic analysis of the arachnid orders based on morphological characters. Zoological Journal of the Linnean Society, 150, 221–265. https://doi.org/10.1111/j.1096-3642.2007.00284.x
    Smit, H. & Gerecke, R. (2010) A checklist of the water mites of France (Acari: Hydrachnidia). Acarologia, 50, 21–91. https://doi.org/10.1051/acarologia/20101952
    Smit, H. & van der Hammen, H. (2000) Nederlandse Faunistische Mededelingen I3 Atlas van de Nederlandse watermijten (Acari: Hydrachnidia). Naturalis Institutional Repository, Amsterdam, 266 pp. [https://repository.naturalis.nl/document/45797]
    Soar, C.D. (1902) Ecpolus papillosus, n. sp. An unrecorded hydrachnid found in Britain. Journal of the Quekett Microscopical Club, 8, 251–252.
    Ström, H. (1768) Beskrivelse over Norske Insecter. Andet Stykke. Det Kongelige Norske Videnskabers Selskabs Skrifter, 4, 313–371.
    Stryjecki, R., Bańkowska, A. & Szenejko, M. (2018) A faunistic and ecological characterization of the water mites (Acari: Hydrachnidia) of the Bukowa River (central-eastern Poland). Acta Biologica, 25, 77–94. https://doi.org/10.18276/ab.2018.25-07
    Subías, L.S. (2018) Systematic, synonimical and biogeographical check-list of the world’s oribatid mites (Acariformes, Oribatida) (Except fossils) (13rd update) (Originally published in Graellsia, 60 (special issue): 3–305 (2004). Updated in June 2006, April 2007, May 2008. Graellsia, 60, 3–305. https://doi.org/10.3989/graellsia.2004.v60.iExtra.218
    Subías, L.S. (2021) Adiciones al listado mundial de Ácaros Oribátidos (Acari, Oribatida) (16a Actualización). Revista Ibérica de Aracnología, 38, 107–108.
    Subías, L.S. & Arillo, A. (2001) Acari, Oribatei, Gymnonota II. In: Sánchez, M.A.R., Tercedor, J.A., Ros, X.B., Noguera, J.G., Sierra, A.G., Mayol, E.M., Piera, F.M., Marino, J.S. & González, J.T. (Eds.), Fauna Ibérica. Vol. 15. Museo Nacional de Ciencias Naturales, Consejo Superior de Investigaciones Científicas (CSIC), Madrid, pp. 289.
    Subías, L.S. & Shtanchaeva, U.Y. (2015) The oribatid mites (Acari, Oribatida) from central Portugal and species cheklist of Portugal. Boletín de la Real Sociedad Española de Historia Natural, Sección Biológica, 109, 91–101.
    Szalay, L. (1927) Eine neue Wassermilbe aus dem Balatonsee. Zoologischer Anzeiger, 71, 279–281.
    Szalay, L. (1970) Verzeichnis der aus dem Karpatenbecken bisher bekannt gewordenen Wassermilben (Hydracarina, Acari) [The watermite fauna of the Carapthia basin (Hydracarina, Acari)]. Acarologia, 12, 136–159.
    Thon, K. (1899) Ein neues Hydrachnidengenus aus Böhmen, nebst einigen Bemerkungen über böhmische Hydryphantes-Formen. (Vorläufige Mitteilung). Zoologischer Anzeiger, 22, 100–102.
    Thor, S. (1898a) Ljania, en ny hydrachnide-slegt fra omegnen af Kristiania, Norge. Forelobig meddelelse. Archiv for mathematik og naturvidenskab, 20, 1–4.
    Thor, S. (1898b) Nye Hydrachnideformer. Fundne I Norge Sommeren. Archiv for mathematik og naturvidenskab, 20, 1–10.
    Thor, S. (1899) En ny Hydrachnide-slegt og andre nye arter fundne i Norge sommeren 1899. Forelobig meddelelse. Kristiania, 1899, 1–5.
    Thor, S. (1900) Hydrachnologische Notizen I-III. Nyt Magazin for Naturvidenskaberne, 38, 267–279.
    Tuzovskij, P.V. & Semenchenko, K.A. (2015) Water mites of the genus Unionicola Haldeman, 1842 (Acari, Hydrachnidia, Unionicolidae) in Russia. Zootaxa, 3919 (3), 401–456. https://doi.org/10.11646/zootaxa.3919.3.1
    Valdecasas, A.G. (1988) Lista sinonímica y bibliográfica de las Hidracnelas (Acari, Hydrachnellae) de la Península Ibérica, Islas Baleares e Islas Canarias. Lista de la flora y fauna de las aguas continentales de la Peninsula Iberica. Publicacion no 5. Asociacion Española de Limnologia, Museo Nacional de Ciencias Naturales (CSIC), Madrid, 81 pp. [http://www.limnetica.net/documentos/listas/Lista_5_Hidracnelas.pdf]
    Viets, K. (1918) Eine neue Limnesia-Species. Zoologischer Anzeiger, 50, 111–112.
    Viets, K. (1930) Zur Kenntnis der Hydracarinen-Fauna von Spanien. Archiv für Hydrobiologie, 21, 175–240 + 359–446.
    Walter, B. (1928) Hydracariens de l’Algérie et de la Tunisie (Collections de M. H. Gauthier) (Deuxième liste). Bulletin de la Société d’histoire naturelle de l’Afrique du nord, 19, 280–336.
    Walter, C. (1925) Hydracariens de l’Algérie et de la Tunisie (Collections de M. H. Gauthier), Première liste. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord, 16, 189–238.
    Walter, C. & Motas, C. (1927) Hydracariens nouveaux ou peu connus du Sud-Est de la France. Travaux du Laboratoire d’hydrobiologie et de pisciculture de l’Université de Grenoble, 11, 65–163.
    Weigmann, G. (2008) Oribatid mites (Acari: Oribatida) from the coastal region of Portugal. I. Peloptulus sacculiferus n. sp., an aberrant species of Phenopelopidae compared with similar European species of the genus. Soil Organisms, 80, 145–151.
    Weigmann, G. (2009a) Oribatid mites (Acari: Oribatida) from the coastal region of Portugal. II. The genera Zachvatkinibates and Punctoribates (Mycobatidae). Soil Organisms, 81, 85–105.
    Weigmann, G. (2009b) Oribatid mites (Acari: Oribatida) from the coastal region of Portugal. III. New species of Scutoverticidae and Scheloribatidae. Soil Organisms, 81, 107–127. https://doi.org/10.1007/0-306-48162-6_3
    Weigmann, G. (2010) Oribatid mites (Acari: Oribatida) from the coastal region of Portugal. IV. The genera Coronoquadroppia, Scheloribates, Haplozetes and Pilobates. Soil Organisms, 82, 383–406.
    Weigmann, G. (2011) Oribatid mites (Acari: Oribatida) from the coastal region of Portugal. V. Xenillus, Oribatella, Galumna, Eupelops and Lucoppia. Soil Organisms, 83, 287–306.
    Weigmann, G. (2012) Oribatid mites (Acari: Oribatida) from the coastal region of Portugal. VI. Chamobates, Protozetomimus, Protoribates, Oribatula. Soil Organisms, 83, 529–550.
    Weigmann, G. (2013) Ecology and biogeography of oribatid mites (Acari: Oribatida) from the coastal region of Portugal. Soil Organisms, 85, 147–160.
    Więcek, M., Martin, P. & Lipinski, A. (2013) Water mites as potential long-term bioindicators in formerly drained and rewetted raised bogs. Ecological Indicators, 34, 332–335. https://doi.org/10.1016/j.ecolind.2013.05.019
    Więcek, M., Szydło, W., Dabert, J. & Proctor, H. (2020) Delimiting species of water mites of the genus Hydrodroma (Acari: Hydrachnidiae: Hydrodromidae) from North America and Europe: Integrative evidence of species status from COI sequences and morphology. Zoologischer Anzeiger, 284, 16–29. https://doi.org/10.1016/j.jcz.2019.11.004
    WoRMS Editorial Board (2022) World Register of Marine Species. Available from: http://www.marinespecies.org/ (accessed 21 April 2022)