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Issue: Vol. 5241 No. 1: 14 Feb. 2023
Type: Monograph
Published: 2023-02-14
DOI: 10.11646/zootaxa.5241.1.1
Page range: 1-115
Abstract views: 348
PDF downloaded: 59

A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)

Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.
SKLLQG, Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97, Yanta District, Xi’an 710061 P.R. China.
Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.
Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Box 92, No. 1, Beichen West Road, Chaoyang District, Beijing, 100101, P.R. China.
Chafers revision keys new species new combinations southeast Asia Coleoptera

Abstract

The current paper gives an overview of the species of continental Asia so far assigned to the genus Microserica Brenske, 1894. Moronoserica Ahrens, Lukic & Liu, new genus, and Gastroserica (Helioserica Ahrens, Lukic & Liu, new subgenus), and 33 new species are described: Gastroserica bannok Ahrens, Lukic & Liu, new species, G. fumaria Ahrens, Lukic & Liu, new species, G. ivoi Ahrens, Lukic & Liu, new species, G. loei Ahrens, Lukic & Liu, new species, G. lucidomarginalis Ahrens, Lukic & Liu, new species, G. phukradung Ahrens, Lukic & Liu, new species, G. piceocoerulea Ahrens, Lukic & Liu, new species, G. rubropicea Ahrens, Lukic & Liu, new species, G. simaoensis Ahrens, Lukic & Liu, new species, G. yuebaensis Ahrens, Lukic & Liu, new species, Microserica allovarians Ahrens, Lukic & Liu, new species, Mic. banlaoana Ahrens, Lukic & Liu, new species, Mic. caiyangheana Ahrens, Lukic & Liu, new species, Mic. chaiyaphum Ahrens, Lukic & Liu, new species, Mic. cucphuong Ahrens, Lukic & Liu, new species, Mic. ferestictica Ahrens, Lukic & Liu, new species, Mic. hieroglyphica Ahrens, Lukic & Liu, new species, Mic. multipunctata Ahrens, Lukic & Liu, new species, Mic. namnao Ahrens, Lukic & Liu, new species, Mic. neosimplex Ahrens, Lukic & Liu, new species, Mic. parasimplex Ahrens, Lukic & Liu, new species, Mic. paravicula Ahrens, Lukic & Liu, new species, Mic. planiforceps Ahrens, Lukic & Liu, new species, Mic. viengvai Ahrens, Lukic & Liu, new species, Mic. vinden Ahrens, Lukic & Liu, new species, Mic. vipinglangensis Ahrens, Lukic & Liu, new species, Moronoserica banmethout Ahrens, Lukic & Liu, new species, Mor. renong Ahrens, Lukic & Liu, new species, Mor. squamulatoides Ahrens, Lukic & Liu, new species, Mor. songbae Ahrens, Lukic & Liu, new species, Mor. thungyai Ahrens, Lukic & Liu, new species, Oxyserica bifascipennis Ahrens, Lukic & Liu, new species, O. serena Ahrens, Lukic & Liu, new species. The revision results in 39 new combinations: Gastroserica bisignata (Nomura, 1974) new combination, G. cognata (Frey, 1972) new combination, G. dohertyi (Ahrens & Fabrizi, 2009) new combination, G. hiulca (Brenske, 1897) new combination, G. lucens (Ahrens & Fabrizi, 2009) new combination, G. nitidipyga (Nomura, 1974) new combination, G. quateorum (Frey, 1972) new combination, G. roingensis (Fabrizi & Ahrens, 2016) new combination, G. varians (Moser, 1915) new combination, Microserica avicula (Arrow, 1946) new combination, Mic. costisquamosa (Ahrens, Fabrizi & Liu, 2019) new combination, Mic. septemfoliata (Frey, 1972) new combination, Moronoserica banvaneue (Bohacz & Ahrens, 2020) new combination, Mor. crenatostriata (Ahrens, 2004) new combination, Mor. geberbauer (Ahrens, 2004) new combination, Mor. ginae (Bohacz & Ahrens, 2020) new combination, Mor. helferi (Bohacz & Ahrens, 2020) new combination, Mor. lineata (Moser, 1915) new combination, Mor. squamulata (Moser, 1915) new combination, Mor. tenasserimensis (Bohacz & Ahrens, 2020) new combination, Mor. ventrosa (Bohacz & Ahrens, 2020) new combination, Oxyserica arunensis (Ahrens, 1998) new combination, O. bhutanensis (Frey, 1975) new combination, O. cechovskyi (Ahrens, 1999) new combination, O. diversicornis (Moser, 1915) new combination, O. elegans (Frey, 1975) new combination, O. gandakiensis (Ahrens, 1998) new combination, O. hispidula (Frey, 1975) new combination, O. interrogator (Arrow, 1946) new combination, O. marginata (Brenske, 1896) new combination, O. martensi (Ahrens, 1998) new combination, O. myagdiana (Ahrens, 1998) new combination, O. nigropicta (Fairmaire, 1891) new combination, O. pedongensis (Ahrens, 1998) new combination, O. pruinosa (Hope, 1831) new combination, O. schawalleri (Ahrens, 1998) new combination, O. schulzei (Ahrens, 1998) new combination, O. steelei (Ahrens, 2004) new combination, O. truncata (Brenske, 1898) new combination. Lectotypes were designated of Microserica hiulca Brenske, 1897 and Serica nigropicta Fairmaire, 1891. A checklist and key to genera, species groups, and species is given, the diagnostic characters of the newly revised taxa and their distribution is illustrated.

References

  1. Ahrens, D. (1995) Taxonomische Studien an Sericinae. Die Gattungen Deroserica Moser, 1915, Microsericaria Nikolaev, 1979, Pachyderoserica Moser, 1920 und Microserica Brenske, 1894. (Ins., Col., Melolonth.). Entomologische Abhandlungen, 57 (2), 37–56.

  2. Ahrens, D. (1996) Revision der Sericini des Himalaya und angrenzender Gebiete. Die Gattungen Lasioserica Brenske, 1896 und Gynaecoserica Brenske, 1896. Coleoptera- Schwanfelder Coleopterologische Mitteilungen, 16, 1–48.

  3. Ahrens, D. (1998) Revision der Sericinae des Himalaya-Gebiets. Neue Arten der Microserica marginata (Brsk.)-Gruppe (Coleoptera, Scarabaeoidea). Entomologische Abhandlungen, 58 (1), 29–51.

  4. Ahrens, D. (1999) Eine neue Art der Microserica marginata-Gruppe aus Ost-Nepal. (Coleoptera: Melonthidae: Sericini). Entomologische Abhandlungen, 58 (12), 243–246.

  5. Ahrens, D. (2000a) Weiteres Material der Gattung Lasioserica Brenske, 1896, mit der Beschreibung acht neuer Arten (Coleoptera: Melolonthidae: Sericini). Acta Musei Moraviae, Scientiae Biologicae, 85, 1–28.

  6. Ahrens, D. (2000b) Synopsis der Gattung Gastroserica Brenske, 1897 des ostasiatischen Festlandes (Coleoptera: Melolonthidae: Sericini). Entomologische Abhandlungen des Staatlichen Museums für Tierkunde Dresden, 59, 73–121.

  7. Ahrens, D. (2000c) Serica korgei Petrovitz, 1967 aus der Türkei- ein neues Synonym von Autoserica castanea Arrow, 1913 (Coleoptera: Melolonthidae: Sericini). Reichenbachia, 33 (51), 395–298.

  8. Ahrens, D. (2001a) Neue Arten der Microserica splendidula (Fabricius, 1801)—Gruppe von West-Malayasia, Nias und Sumatra (Coleoptera: Melolonthidae). Koleopterologische Rundschau, 71, 111–120.

  9. Ahrens, D. (2001b) Further records of Microserica Brenske, 1894 from the Himalayas with description of a new species of the subgenus Parvulomaladera Ahrens, 1995. I. Part (Coleoptera, Melolonthidae, Sericini). Lambillionea, CI (2), 216–224.

  10. Ahrens, D. (2001c) Zwei neue Chrysoserica-Arten aus dem Himalaya. (Coleoptera, Melolonthidae). Bolletino della Società Entomologica Italiana, 133 (2), 133–147.

  11. Ahrens, D. (2002a) Notes on distribution and synonymy of sericid beetles from Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea, Sericini). Annales Historico-Naturales Musei Nationalis Hungarici, 94, 53–91.

  12. Ahrens, D. (2002b) Revision der Arten der Microserica viridicollis Arrow, 1913—Gruppe (Coleoptera, Melolonthinae, Sericini). Linzer biologische Beiträge, 34/1, 383–412.

  13. Ahrens, D. (2003a) Neue Arten aus der Gruppe der Amiserica insperata (Brenske, 1898) (Coleoptera, Scarabaeoidea, Sericini). Spixiana, 26, 65–81.

  14. Ahrens, D. (2003b) Zur Identität der Gattung Neoserica Brenske, 1894, nebst Beschreibung neuer Arten (Coleoptera, Melolonthidae, Sericini). Koleopterologische Rundschau, 73, 169–226.

  15. Ahrens, D. (2003c) Zur Identität der aus Madagaskar beschriebenen Autoserica stupida Brenske, 1900. (Coleoptera, Scarabaeidae, Sericini). Beiträge zur Entomologie, 53, 231–238.

  16. Ahrens, D. (2003d) On the Identity of Maladera infuscata (Moser, 1915), with description of a new species from Taiwan (Coleoptera, Scarabaeidae, Sericini). Zootaxa, 392 (1), 1–8. https://doi.org/10.11646/zootaxa.392.1.1

  17. Ahrens, D. (2004a) New species of the genus Microserica Brenske, 1894 from Sumatra and the Malay Peninsula, with notes on synonymy. (Coleoptera, Scarabaeoidea, Melolonthidae). Annali del Museo Civico di Storia Naturale “G. Doria”, 95, 35–63.

  18. Ahrens, D. (2004b) Monographie der Sericini des Himalaya (Coleoptera, Scarabaeidae). Dissertation.de—Verlag im Internet GmbH, Berlin, 534 pp.

  19. Ahrens, D. (2004c) Revision of the Microserica lineata group (Coleoptera: Scarabaeidae: Sericini). Zootaxa, 664, 1–14.

  20. Ahrens, D. (2005a) Description of the two new Microserica species from Laos and Thailand (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa, 1064 (1), 31–37. https://doi.org/10.11646/zootaxa.1064.1.3

  21. Ahrens, D. (2005b) A taxonomic review on the Serica (s. str.) MacLeay, 1819 species of Asiatic mainland (Coleoptera, Scarabaeidae, Sericini). Nova Supplementa Entomologica, 18, 1–163.

  22. Ahrens, D. (2005c) Taxonomic revision of the genus Anomalophylla Reitter, 1887 (Coleoptera, Scarabaeidae: Sericini). Zootaxa, 1076 (1), 1–62. https://doi.org/10.11646/zootaxa.1076.1.1

  23. Ahrens, D. (2006a) Sericinae. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea—Scirtoidea—Dascilloidea—Buprestoidea—Byrrhoidea. Apollo Books, Stenstrup, pp. 229–248.

  24. Ahrens, D. (2006b) The phylogeny of the genus Lasioserica inferred from adult morphology – implications on the evolution of montane fauna of the South Asian orogenic belt (Coleoptera: Scarabaeidae: Sericini). Journal of Zoological Systematics and Evolutionary Research, 44, 34–53. https://doi.org/10.1111/j.1439-0469.2005.00340.x

  25. Ahrens, D. (2006c) Revision und phylogenetische Analyse der Gattung Pachyserica Brenske, 1897 (Coleoptera, Melolonthidae, Sericini). Revue Suisse de Zoologie, 113, 487–557. https://doi.org/10.5962/bhl.part.80360

  26. Ahrens, D. (2007a) Revision der Serica nigroguttata Brenske, 1897 - Gruppe (Coleoptera, Melolonthidae, Sericini). Bulletin de l’Institut Royal des Sciences naturelles de Belgique, 77, 5–37.

  27. Ahrens, D. (2007b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1–51. https://doi.org/10.11646/zootaxa.1504.1.1

  28. Ahrens, D. (2009) A cladistic analysis reveals the occurrence of Taiwanoserica in Sichuan (China) with description of four new species (Coleoptera, Scarabaeidae). Annales de la Société Entomologique de France, New Series, 45 (3), 285–296. https://doi.org/10.1080/00379271.2009.10697612

  29. Ahrens, D. (2011) A revision of the genus Archeohomaloplia Nikolaev, 1982. (Coleoptera: Scarabaeidae: Sericini). Bonn Zoological Bulletin, 60, 117–138.

  30. Ahrens, D. (2021) Two new species of the Neoserica (sensu stricto) group from China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Journal of Natural History, 2021, 54 (45–46), 2927–2936. https://doi.org/10.1080/00222933.2021.1884761

  31. Ahrens, D. & Bezděk, A. (2016) Sericini. In: Löbl, I. & Löbl, D (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea—Scirtoidea—Dascilloidea—Buprestoidea—Byrrhoidea. Revised and Updated Edition. Brill, Leiden, pp. 281–317.

  32. Ahrens, D. & Fabrizi, S. (2009a) A review on the genus Gynaecoserica Brenske, 1897 (Coleoptera, Scarabaeidae, Sericini). Journal of Natural History, 43, 1505–1584. https://doi.org/10.1080/00222930902968809

  33. Ahrens, D. & Fabrizi, S. (2009b) New species of Sericini from the Eastern Himalaya and Tibet (Coleoptera, Scarabaeidae). In: Hartmann, M. & Weipert, M. (Eds.), Biodiversität und Naturausstattung im Himalaya III. Verein der Freunde und Förderer des Naturkundemuseums Erfurt e.V., Erfurt, pp. 249–284.

  34. Ahrens, D. & Fabrizi, S. (2011) New species of Sericini from the Himalaya and adjacent mountains (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 60, 139–164.

  35. Ahrens, D. & Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1–355.

  36. Ahrens, D. & Pacholátko, P. (2003) New data on distribution of the species of Gastroserica Brenske, 1897, with description of new taxa from China and Laos (Coleoptera: Scarabaeidae: Sericini). Zootaxa, 342 (1), 1–18. https://doi.org/10.11646/zootaxa.342.1.1

  37. Ahrens, D. & Pacholátko, P. (2007) Two new species of Gastroserica Brenske, 1897 from China and Myanmar (Coleoptera: Scarabaeidae: Sericini). Entomologische Zeitschrift, 117, 137–139.

  38. Ahrens, D. & Vogler, A. P. (2008) Towards the phylogeny of chafers (Sericini): analysis of alignment-variable sequences and the evolution of segment numbers in the antennal club. Molecular Phylogenetics and Evolution, 47, 783–798. https://doi.org/10.1016/j.ympev.2008.02.010

  39. Ahrens, D., Liu, W.G., Fabrizi, S., Bai, M. & Yang, X.K. (2014a) A taxonomic review of the Neoserica (sensu lato) septemlamellata group (Coleoptera: Scarabaeidae: Sericini). ZooKeys, 402, 76–102. https://doi.org/10.3897/zookeys.402.7360

  40. Ahrens, D., Liu, W.G., Fabrizi, S., Bai, M. & Yang, X.K. (2014b) A taxonomic review of the Neoserica (sensu lato) abnormis group (Coleoptera: Scarabaeidae: Sericini). ZooKeys, 439, 28–82. https://doi.org/10.3897/zookeys.439.8055

  41. Ahrens, D., Liu, W.G., Fabrizi, S., Bai, M. & Yang, X.K. (2014c) A revision of the species of the Neoserica (sensu lato) vulpes group (Coleoptera: Scarabaeidae: Sericini). Journal of Natural History, 49 (17–18), 1073–1130. [online version; printed 2015] https://doi.org/10.1080/00222933.2014.974707

  42. Ahrens, D., Liu, W.G., Fabrizi, S., & Bai, M. (2021) Taxonomic review on the Trioserica Moser, 1922 species of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa, 4999 (4), 343–355. https://doi.org/10.11646/zootaxa.4999.4.4

  43. Arrow, G.J. (1913) Zoological results of the Abor Expedition 1911–1912. XII. Coleoptera, IV. Lamellicornia. Records of Indian Museum, 8, 191–196. https://doi.org/10.26515/rzsi/v8/i3/1913/163144

  44. Arrow, G.J. (1946a) Notes on Aserica and some related genera of melolonthine beetles, with descriptions of a new species and two new genera. Annals and Magazine of Natural History Series, 11, 13, 264–283. https://doi.org/10.1080/00222934608654552

  45. Arrow, G.J. (1946b) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Melolonthidae. Arkiv för Zoologie, 38A (9), 1–33.

  46. Barlow, E. (1899) List of the Melolonthini contained in the colection of the Indian Museum. Indian Museum Notes, 4, 233–272.

  47. Blanchard, C. (1850) Ier Famille—Scarabaeidae. In: Milne-Edwards, H., Blanchard, C. & Lucas, H., Catalogue de la collection entomologique du Muséum d’Histoire Naturelle de Paris. Classe des insectes. Ordre des Coléoptères. Tome I. Premiè re livraison. Gide et Baudry, Paris, pp. 1–128.

  48. Bohacz, C. & Ahrens, D. (2019) A new species of the Neoserica (sensu lato) vulpes group from China, with a corrigendum on Neoserica ailaoshanica Liu, Fabrizi, Bai, Yang Ahrens, 2014 (Coleoptera: Scarabaeidae: Sericini). Journal of Natural History, 53, 47–48, 2991–2997. https://doi.org/10.1080/00222933.2020.1758819

  49. Bohacz, C. & Ahrens, D. (2020) New species of the Microserica lineata group from Laos and Thailand (Coleoptera: Scarabaeidae: Sericini). Zootaxa, 4859 (2), 263–274. https://doi.org/10.11646/zootaxa.4859.2.5

  50. Brenske, E. (1894) Die Melolonthiden der palaearktischen und orientalischen Region im Königlichen naturhistorischen Museum zu Brüssel. Beschreibung neuer Arten und Bemerkung zu bekannten. Mémoire de la Société Entomologique de Belgique, 2, 3–87.

  51. Brenske, E. (1896) Insectes du Bengale. Melolonthidae. Annales de la Société Entomologique de Belgique, 40, 150–164.

  52. Brenske, E. (1897) Die Serica-Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345–468. https://doi.org/10.1002/mmnd.18970420310

  53. Brenske, E. (1898) Die Serica-Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205–403. https://doi.org/10.1002/mmnd.47918980303

  54. Brenske, E (1900a) Die Serica-Arten der Erde. IV. Berliner Entomologische Zeitschrift, 45, 39–96.

  55. Brenske, E. (1902) Die Serica-Arten der Erde. VII. Berliner Entomologische Zeitschrift, 47, 1–70.

  56. Eberle, J., Fabrizi, S., Lago P. & Ahrens, D. (2017) A historical biogeography of megadiverse Sericini—another story "out of Africa"? Cladistics, 33, 183–197. https://doi.org/10.1111/cla.12162

  57. Fabrizi, S., Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. (2021) A monograph of the genus Maladera of China (Coleoptera, Scarabaeidae, Sericini). Zootaxa, 4922 (1), 1–400. https://doi.org/10.11646/zootaxa.4922.1.1

  58. Fairmaire, M.L. (1891) Coléoptères de l’Intérieur de la Chine (7). Annales de la Société Entomologique de Belgique, 35, 187–219.

  59. Frey, G. (1965b) Neue Sericinen aus dem nordöstlichen Himalaya. Khumbu Himal. Ergebnisse des Forschungsunternehmens Nepal Himalaya, 2, 88–93. https://doi.org/10.1007/978-3-642-92901-4_6

  60. Frey, G. (1969a) Neue Melolonthiden aus Nepal. Entomologische Arbeiten aus dem Museum G. Frey, 20, 518–525.

  61. Frey, G. (1972) Neue Sericinen aus Indien und Indochina, sowie Abbildungen von Parameren bekannter Arten (Col., Scar., Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey, 23, 186–216.

  62. Frey, G. (1975b) Ergebnisse der Bhutan Expedition 1972 des Naturhistorischen Museums in Basel. Col.: Fam. Scarabaeidae subf. Melolonthidae. Entomologica Basiliensia, 1, 223–241.

  63. Hope, F.W. (1831) Synopsis of the species of Nepaul Insects in the collection of Major General Hardwicke. Zoological Miscellany, 1, 21–33.

  64. Kobayashi, H. (2015) Two new species of sericid-beetles (Coleoptera, Scarabaeidae, Melolonthinae, Sericini) from Taiwan. Kogane, 17, 25–32.

  65. Liu, W.G., Ahrens, D., Bai, M. & Yang, X.K. (2011) A key to species of the genus Gastroserica Brenske of the China (Coleoptera, Scarabaeidae, Sericini), with the description of two new species and two new records for China. ZooKeys, 139, 23–44. https://doi.org/10.3897/zookeys.139.1702

  66. Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. (2014a) An update to the taxonomy of the genus Gastroserica Brenske (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 426, 87–110. https://doi.org/10.3897/zookeys.426.7578

  67. Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. (2014b) Towards an improved knowledge of Sericini of the Tibetan highland: new species and records (Coleoptera, Scarabaeidae). Bonn Zoological Bulletin, 63, 157–172.

  68. Liu, W.G., Fabrizi, S., Bai, M., Yang, X.K. & Ahrens, D. (2014c) A taxonomic revision of the Neoserica (sensu lato) pilosula group (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 440, 89–113. https://doi.org/10.3897/zookeys.440.8126

  69. Liu, W.G., Fabrizi, S., Bai, M., Yang, X.K. & Ahrens, D. (2014d) A taxonomic revision of the Neoserica (sensu lato) calva group (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 448, 47–81. https://doi.org/10.3897/zookeys.448.8368

  70. Liu, W.G., Fabrizi, S., Bai, M., Yang, X.K. & Ahrens, D. (2014e) A review of the Tetraserica species of China (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 448, 83–121. https://doi.org/10.3897/zookeys.443.8429

  71. Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. (2015a) New species and records of the Neoserica (sensu stricto) group (Coleoptera, Scarabaeidae, Sericini). Journal of Natural History, 49 (39–40), 2379–2395. https://doi.org/10.1080/00222933.2015.1034208

  72. Liu, W.G., Bai, M., Eberle, J., Yang, X.K. & Ahrens, D. (2015b) A phylogeny of Sericini with particular reference to Chinese species using mitochondrial and ribosomal DNA (Coleoptera: Scarabaeidae). Organisms Diversity and Evolution, 15, 343–350. https://doi.org/10.1007/s13127-015-0204-z

  73. Liu, W.G., Fabrizi, S., Bai, M., Yang, X.K. & Ahrens, D. (2016) A taxonomic revision of Neoserica (sensu lato): the species groups N. lubrica, N. obscura, and N. silvestris (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 635, 123–160. https://doi.org/10.3897/zookeys.635.9915

  74. Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. (2017a) New species and records of Lasioserica and Gynaecoserica from China (Coleoptera, Scarabaeidae, Sericini). Bonn Zoological Bulletin, 66, 29–36.

  75. Liu, W.G., Fabrizi, S., Bai, M., Yang, X.K. & Ahrens, D. (2017b) New species of Nipponoserica and Paraserica from China (Coleoptera: Scarabaeidae: Sericini). ZooKeys, 721, 65–91. https://doi.org/10.3897/zookeys.721.13918

  76. Liu, W.G., Fabrizi, S., Bai, M., Yang, X.K. & Ahrens, D. (2019) A taxonomic revision of Chinese Neoserica (sensu lato): final part (Coleoptera: Scarabaeidae: Sericini). Bonn Zoological Bulletin Supplement, 64, 1–71. https://doi.org/10.20363/BZB-S-2019.64

  77. Medvedev, S.I. (1952) Plastinchatousye (Scarabaeidae), podsem. Melolonthinae. Ch. 2 (chrushchi). Fauna SSSR, zhestkokrylye. Tom 10. Vyp. 2. Izdatelstvo Akademii Nauk SSSR, Moskva, Leningrad, 274 pp.

  78. Moser, J. (1915) Neue Serica-Arten. Deutsche Entomologische Zeitschrift, 1915, 337–393. https://doi.org/10.1002/mmnd.48019150402

  79. Moser, J (1918) Neue Melolonthiden aus der Sammlung des Deutschen Entomologischen Museums zu Berlin-Dahlem. Stettiner Entomologische Zeitung, 79, 209–247.

  80. Niijima, Y. & Kinoshita, E. (1923) Die Untersuchungen über japanische Melolonthiden. II. Research Bulletin of the College Experimental Forestry Hokkaido Imperial University, 2, 1–253.

  81. Nomura, S. (1973a) Notes on Japanese Scarabaeidae, III. Entomological Review of Japan, 25, 13–20.

  82. Nomura, S. (1973b) On the Sericini of Japan. I. Tôhô-Gakuhô, 23, 119–152.

  83. Nomura, S. (1974) On the Sericini of Taiwan. Tôhô-Gakuhô, 24, 81–115.

  84. Pacheco, T.L., Monné, M.L. & Ahrens, D. (2021) The morphospace of Neotropical Sericini (Coleoptera: Scarabaeidae) - a comparative analysis towards the relationship of shape disparity vs. species diversity. Organisms, Diversity and Evolution, 22, 177–188. https://doi.org/10.1007/s13127-021-00530-y

  85. Pacheco, T.L., Wipfler, B., Monné, M.L. & Ahrens, D. (2022a) The genus Raysymmela Saylor, 1947 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis. Insect Systematics & Evolution, 53, 400–441. https://doi.org/10.1163/1876312X-bja10031

  86. Pacheco, T.L., Monné, M.L., Vaz-de-Mello, F.Z. & Ahrens, D. (2022b) First non-feeding species of Sericini chafers (Coleoptera, Scarabaeidae): new genus and phylogenetic position. Organisms, Diversity and Evolution, 22, 733–748. https://doi.org/10.1007/s13127-022-00555-x

  87. Pacheco, T.L., Wipfler, B., Monné, M.L. & Ahrens, D. (2022c) The genus Symmela Erichson, 1847 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis. Journal of Natural History, 56 (9–12), 607–705. https://doi.org/10.1080/00222933.2022.2084649

  88. Sabatinelli, G. (1993) Catalogue of the Scarabaeoidea Melolonthidae from Himalaya and Norther India: Macrodactylini, Hoplini, Diplotaxini, Melolonthini and Sericini. Memoirie della Società Entomologica Italiana, 71 (1992), 593–636.

  89. Shrestha, J., Eberle, J. & Ahrens, D. (2012) Further data on the distribution of Himalayan Sericini from the collection of Naturel History Museum Erfurt (Insecta: Coleoptera: Scarabaeidae) with description of a new Xenoserica from West-Nepal. Vernate, 31, 379–386.

  90. Sreedevi, K., Ranasinghe, S., Fabrizi, S. & Ahrens, D. (2019) New species and records of Sericini scarab beetles from the Indian subcontinent (Coleoptera, Scarabaeidae) II. European Journal of Taxonomy, 567, 1–26. https://doi.org/10.5852/ejt.2019.567

  91. Yu, C.K., Kobayashi, H. & Chu, Y.I. (1998) The Scarabaeidae of Taiwan. Mu-Sheng Entomological Corporation, Taipei, 263 pp.