Abstract
Three hundred and fiftysix species of ceratopogonids in 35 genera are listed for Argentina, with most species in Forcipomyia Meigen (67 species) Culicoides Latreille (51 species), Stilobezzia Kieffer (40 species) and Dasyhelea Kieffer (37 species). For each listed species, the following information is included: sex and/or life stage described in the original description, type locality, type status and sex, depository of type material, distribution, georeferenced localities for Argentina (provinces are underlined) and main references that include the original description and those that specify locations in the country. Specific epithets of synonyms are accompanied by type locality, type status and sex, and location of type material. New localities from Argentina for 102 species are provided, and the following 16 species are newly recorded from the country: Forcipomyia (Caloforcipomyia) glauca Macfie, Forcipomyia (C.) hatoensis Utmar & Wirth, F. (Euprojoannisia) bromeliae Saunders, F. (E.) quasiingrami Macfie, F. (Forcipomyia) zonogaster Ingram & Macfie, F. (F.) catarinensis Marino & Spinelli, F. (Lasiohelea) cornuta Saunders, F. (Metaforcipomyia) osaensis Spinelli, Marino & Borkent, F. (Phytohelea) bromelicola (Lutz), F. (Trichohelea) goniognatha Wirth & Messersmith, F. (T.) leptognatha Wirth & Messersmith, Brachypogon (Brachypogon) woodruffi Spinelli & Grogan, Physohelea turgidipes Ingram & Macfie, Bezzia gibbera (Coquillett), Bezzia schmitzorum Dippolito & Spinelli and Palpomyia versicolor Macfie. Leptoconops brasiliensis (Lutz) and Clastrieromyia dycei Spinelli & Grogan are excluded from the Argentinean fauna.
References
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Ronderos, M.M. & Spinelli, G.R. (1995b) Redescripción de Culicoides lahillei y descripción de Culicoides ichesi, n. sp. de la Argentina, Paraguay y Uruguay (Diptera: Ceratopogonidae). Neotrópica, 41, 77–81.
Ronderos, M.M. & Spinelli, G.R. (1997) Culicoides patagoniensis n. sp. del sur de Argentina y Chile (Diptera: Ceratopogonidae). Gayana Zoología, 61, 33–39.
Ronderos, M.M. & Spinelli, G.R. (1998) Las especies de Culicoides en el área de influencia de la represa de Yacyretá (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 57, 79–83.
Ronderos, M.M. & Spinelli, G.R. (1999) On the subgenus Forcipomyia (Lasiohelea) in the Neotropical Region (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 125, 151–161 (June).
Ronderos, M.M. & Spinelli, G.R. (2000) The larva and pupa of Culicoides bambusicola Lutz observed with SEM, and additional notes on the adult (Diptera Ceratopogonidae). Transactions of the American Entomological Society, 126, 133–144.
Ronderos, M.M. & Spinelli, G.R. (2002) Los Culicoides de la Argentina: clave de especies y nuevos registros (Diptera: Ceratopogonidae). In: Salomón, D. (Comp.), Actualizaciones en Artropodología Sanitaria Argentina. Serie Enfermedades Transmisibles. Pub. Monogr. Nº 2. Fundación Mundo Sano, Buenos Aires, pp. 89–95.
Ronderos, M.M. & Spinelli, G.R. (2009) Description of the immatures of the predaceous midge Bezzia blantoni Spinelli & Wirth (Diptera: Ceratopogonidae). Zootaxa, 2295 (1), 46–54.https://doi.org/10.11646/zootaxa.2295.1.5
Ronderos, M.M., Spinelli, G.R. & Borkent, A. (2008) A description of the larva and pupa of Culicoides charruus Spinelli & Martínez (Diptera: Ceratopogonidae) from leaf axils of Eryngium pandanifolium (Apiaceae) in Argentina. Russian Entomological Journal, 17, 115–122.
Ronderos, M.M., Spinelli, G.R. & Borkent, A. (2012a) A peculiar new species of Stilobezzia Kieffer breeding in bamboo internodes in northeastern Argentina (Diptera: Ceratopogonidae). Aquatic Insects, 34, 1–17.https://doi.org/10.1080/01650424.2012.718079
Ronderos, M.M., Spinelli, G.R. & Díaz, F. (2007) Redescription of the last instar larva and pupa of Bezzia roldani (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 66, 21–28.
Ronderos, M.M., Spinelli, G.R. & Grogan, W.L. Jr. (2017a) The Neotropical species of the predaceous midge genus Austrohelea Wirth & Grogan (Diptera: Ceratopogonidae). Zootaxa, 4276 (2), 255–269.https://doi.org/10.11646/zootaxa.4276.2.7
Ronderos, M.M., Spinelli, G.R., Huerta, H. & Díaz, F. (2003a) Immature stages of two Neotropical species of Dasyhelea Kieffer, 1911 (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 129, 295–308.
Ronderos, M.M., Spinelli, G.R., Marino, P.I. & Ferreyra Keppler, R.L. (2017b) Description of unknown or poorly known pupae and adult males of Neotropical Heteromyia Say (Culicomorpha: Ceratopogonidae), with specific key to pupae. Zoologischer Anzeiger, 266, 177–178.https://doi.org/10.1016/j.jcz.2016.12.001
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Spinelli, G.R. (1982) Cuatro especies nuevas del género Artichopogon [sic] Kieffer (Diptera: Ceratopogonidae) en la República Argentina. Revista de la Sociedad Entomológica Argentina, 41, 201–210.
Spinelli, G.R. (1983a) Notas sobre Ceratopogonidae de la República Argentina (Diptera, Nematocera) II. Nuevos aportes al conocimiento del género Forcipomyia Meigen. Neotrópica, 29, 121–129.
Spinelli, G.R. (1983b) Notas sobre Ceratopogonidae (Diptera, Nematocera) de la República Argentina I. Una nueva especie del género Alluaudomyia Kieffer, redescripción de Dasyhelea penthesileae Macfie, y nuevas citas para el género Stilobezzia Kieffer. Limnobios, 2, 403–411.
Spinelli, G.R. (1983c) Notas sobre Ceratopogonidae (Diptera: Nematocera) de la República Argentina. III. Nuevos aportes al conocimiento de la tribu Palpomyiini. Limnobios, 2, 17–24.
Spinelli, G.R. (1984) Notas sobre Ceratopogonidae (Diptera: Nematocera) de la República Argentina. IV. Descripción de adulto y pupa de Neobezzia termophila sp. nov. Neotrópica, 30, 197–200.
Spinelli, G.R. (1987a) Notas sobre Ceratopogonidae (Diptera: Nematocera) de la República Argentina. VI. Las hembras de Paradasyhelea brevipalpis y de Macrurohelea paracaudata. Limnobios, 2, 667–670.
Spinelli, G.R. (1987b) Nuevas citas de Ceratopogonidae para la Argentina y Uruguay (Insecta: Diptera). Revista de la Sociedad Entomológica Argentina, 44, 160.
Spinelli, G.R. (1988) Two new species of Neotropical Alluaudomyia Kiefer, 1913 (Insecta, Diptera, Certopogonidae [sic]). Iheringia, série Zoologia, 68, 129–136.
Spinelli, G.R. (1989) Amerohelea similis n. sp., from Argentina and Uruguay (Diptera: Ceratopogonidae). Revista de la Asociación de Ciencias Naturales del Litoral, 20, 25–28.https://doi.org/10.14409/natura.v1i20.3572Spinelli, G.R. (1990) The genus Brachypogon in Argentina, with a key to the Neotropical species (Diptera, Ceratopogonidae). Revista Brasileira de Entomologia, 34, 743–755.
Spinelli, G.R. (1994) The male of Physohelea turgidipes (Diptera: Ceratopogonidae). Neotrópica, 40, 73–74.
Spinelli, G.R. (1996) A new species in the genus Diaphanobezzia from subantarctic Argentina (Diptera: Ceratopogonidae). Neotrópica, 42, 77–79.
Spinelli, G.R. (1997a) A new genus of the tribe Sphaeromiini from subantarctic Argentina and Chile related to Mackerrasomyia Debenham (Diptera: Ceratopogonidae). Memoirs of the Entomological Society of Washington, 18, 224–229.
Spinelli, G.R. (1997b) Further notes on Neotropical Ceratopogonini (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 56, 123–124.
Spinelli, G.R. (1998) Three new species and new records of Neotropical Stenoxenini (Diptera: Ceratopogonidae). Neotrópica, 44, 51–55.
Spinelli, G.R. (2000) New records of biting midges from southern South America (Diptera: Ceratopogonidae). Neotrópica, 46, 69–70.
Spinelli, G.R. (2019) Synonymy in Neotropical Ceratopogonidae (Diptera: Culicomorpha). Revista de la Sociedad Entomológica Argentina, 78, 38–39.https://doi.org/10.25085/rsea.780308
Spinelli, G.R. & Balseiro, E.G. (1982) Observaciones relativas a la actividad nocturna de Ceratopogonidae (Diptera: Nematocera) en la zona de Los Talas (Provincia de Buenos Aires), mediante el empleo de una trampa de luz. Revista de la Sociedad Entomológica Argentina, 41, 225–232.
Spinelli, G.R., Borkent, A. & Ronderos, M.M. (2013a) Two new peculiar species of Neotropical Brachypogon Kieffer (Diptera: Ceratopogonidae). Zootaxa, 3702 (1), 90–96.https://doi.org/10.11646/zootaxa.3702.1.6
Spinelli, G.R. & Cazorla, C. G. (2003) New findings of species of the tribe Palpomyiini in Argentina (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 62, 46–48.
Spinelli, G.R. & Cazorla, C.G. (2004) A new species of Brachypogon (Isohelea) from arid zones of Argentina (Diptera: Ceratopogonidae). Zootaxa, 570 (1), 1–6.https://doi.org/10.11646/zootaxa.570.1.1
Spinelli, G.R. & Cazorla, C.G. (2005) The subgenus Stilobezzia (Eukraiohelea) in Argentina and Paraguay (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 64, 72–74.
Spinelli, G.R. & Cazorla, C.G. (2006) A new predaceous midge of the Palpomyia tibialis group from northeastern Argentina (Diptera: Ceratopogonidae). Revista de Biología Tropical, 54, 1067–1070.https://doi.org/10.15517/rbt.v54i3.13958
Spinelli, G.R. & Duret, J.P. (1993) Las especies neotropicales de Clinohelea (Diptera: Ceratopogonidae). Graellsia, 49, 39–50.
Spinelli, G.R. & Felippe-Bauer, M.L. (1990a) Two new Neotropical species of predaceous midges of the tribe Sphaeromiini (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 85, 87–90.https://doi.org/10.1590/S0074-02761990000100014
Spinelli, G.R. & Felippe-Bauer, M.L. (1990b) Sphaerohelea, a new Neotropical predaceous midge genus of the tribe Sphaeromiini (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 85, 195–198.https://doi.org/10.1590/S0074-02761990000200009
Spinelli, G.R., Greiner, E.C. & Wirth, W.W. (1993) The Neotropical bloodsucking midges of the Culicoides guttatus group of the subgenus Hoffmania (Diptera: Ceratopogonidae). Contributions of the American Entomological Institute, 27, 1–91.
Spinelli, G.R. & Grogan, W.L. (1984) Three new species of Macrurohelea from Argentina with a key to the Neotropical species (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 86, 961–967.
Spinelli, G.R. & Grogan, W.L. (1985) Clastrieromyia, a new Neotropical genus of predaceous midges related to Palpomyia and Bezzia (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 87, 329–334.
Spinelli, G.R. & Grogan, W.L. (1986) Two new species of the predaceous midge genus Clastrieromyia from Uruguay with a new record of C. schnacki for Argentina (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 88, 455–460.
Spinelli, G.R. & Grogan, W.L. (1987) A revision of the Neotropical species of Parabezzia (Diptera: Ceratopogonidae). Biología Acuática, 11, 1–45.
Spinelli, G.R. & Grogan, W.L. (1990) New species of predaceous midges of the tribe Ceratopogonini from subantarctic Argentina (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 92, 127–134.
Spinelli, G.R. & Grogan, W.L. (1993) Borkenthelea, a new predaceous midge genus from subantarctic Argentina (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 95, 321–326.
Spinelli, G.R. & Grogan, W.L. (1994) Two new Neotropical species of the subgenus Isohelea of Brachypogon, with the description of the female of B. (I.) misionensis and a key to the Neotropical species (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 53, 1–8.
Spinelli, G.R. & Grogan, W.L. (1998) A revision of the Neotropical predaceous midges of Brachypogon (Brachypogon) Kieffer (Diptera: Ceratopogonidae). Insecta Mundi, 12, 59–79.
Spinelli, G.R. & Grogan, W.L. (1999) A new species of Macrurohelea Ingram and Macfie, and new records of biting midges of the tribes Culicoidini and Ceratopogonini (Diptera: Ceratopogonidae) from Tierra del Fuego and the Magallanes. Proceedings of the Entomological Society of Washington, 101, 708–713.
Spinelli, G.R. & Grogan, W.L. (2001) A revision of the Patagonian predaceous midge genus Borkenthelea Spinelli and Grogan (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 103, 147–156.
Spinelli, G.R. & Grogan, W.L. (2003) A revision of the Neotropical biting midges of the genus Paradasyhelea Ingram and Macfie (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 105, 568–577.
Spinelli, G.R., Grogan, W.L. & Ronderos, M.M. (2009b) A revision of the Patagonian predaceous midges of the genus Palpomyia Meigen (Diptera: Ceratopogonidae). Insect Systematics and Evolution, 40, 43–70.https://doi.org/10.1163/187631209X416705
Spinelli, G.R. & Marino, P.I. (1997) Two new species of the subgenus Thyridomyia of Forcipomyia from Argentina and new records of F. (Synthyridomyia) sanctaeclarae (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 123, 187–190.
Spinelli, G.R. & Marino, P.I. (1998) First records for Argentina of three species of Forcipomyia (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 57, 39–40.
Spinelli, G.R. & Marino, P.I. (2007) A new Neotropical species of Atrichopogon Kieffer, and a redescription of A. casali Cavalieri & Chiossone (Diptera: Ceratopogonidae). Studies on Neotropical Fauna and Environment, 42, 203–209.https://doi.org/10.1080/01650520601136912
Spinelli, G.R. & Marino, P.I. (2018) Forcipomyia (Synthyridomyia) murina (Winnertz, 1852) in South America. Synonymy and redescription (Diptera: Ceratopogonidae). Biología Acuática, 33, 1–7.https://doi.org/10.24215/16684869e001
Spinelli, G.R., Marino, P.I. & Borkent, A. (2009a) First description of the male of the predaceous midge genus Notiohelea Grogan & Wirth (Diptera: Ceratopogonidae). Entomological News, 120, 305–310.https://doi.org/10.3157/021.120.0309
Spinelli, G.R., Marino, P.I. & Borkent, A. (2012a) A revision of biting midges of the subgenera Forcipomyia (Metaforcipomyia) and F. (Saliohelea) from Costa Rica (Diptera: Ceratopogonidae). Zootaxa, 3419 (1), 1–52.https://doi.org/10.11646/zootaxa.3419.1.1
Spinelli, G.R., Marino, P.I. & Gaddi, A.L. (2010) New records of biting midges from Argentina and Chile (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 69, 127–132.
Spinelli, G.R., Marino, P.I. & Huerta, H. (2015a) Revision of the Notropical species of the subgenus Atrichopogon (Psilokempia) (Diptera: Ceratopogonidae). Zootaxa, 4003 (1), 1–64.https://doi.org/10.11646/zootaxa.4003.1.1
Spinelli, G.R., Marino, P.I. & Mauad, M. (2012b) Diptera, Ceratopogonidae Newman, 1834: New records of Biting and Predaceous midges from Iberá wetlands, Argentina. Check List, 8, 886–888.
Spinelli, G.R., Marino, P.I. & Mauad, M. (2012c) Wing pattern variation in the Patagonian biting midge, Forcipomyia (Forcipomyia) multipicta Ingram & Macfie (Diptera, Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 71, 309–313.
Spinelli, G.R., Marino, P.I. & Posadas, P. (2006) The Patagonian species of the genus Atrichopogon Kieffer, with a biogeographic analysis based on Forcipomyiinae (Diptera: Ceratopogonidae). Insect Systematics and Evolution, 37, 301–324.https://doi.org/10.1163/187631206788838581
Spinelli, G.R., Marino, P.I. & Ronderos, M.M. (2005b) The fourth instar larva and pupa of the Neotropical species Forcipomyia (Forcipomyia) rioplatensis Marino & Spinelli (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 107, 108–114.
Spinelli, G.R. & Martinez, M.E. (1992) The genus Culicoides in Uruguay (Diptera: Ceratopogonidae). Insecta Mundi, 5, 175–179 (1991).
Spinelli, G.R. & Rodríguez, E.A. (1999) A new species of Dasyhelea from Argentina described as adult, pupae and fourth instar larvae (Diptera: Ceratopogonidae). Neotrópica, 45, 59–62.
Spinelli, G.R. & Ronderos, M.M. (1987) Notas sobre Ceratopogonidae (Diptera: Nematocera) de la República Argentina. V. Nuevos aportes al conocimiento del género Dasyhelea Kieffer. Neotrópica, 33, 11–17.
Spinelli, G.R. & Ronderos, M.M. (1991) Los polvorines del género Culicoides en área de influencia de la represa de Salto Grande (Diptera: Ceratopogonidae). Neotrópica, 37, 83–94.
Spinelli, G.R. & Ronderos, M.M. (1994) Nuevos registros de Culicoides para los bosques subantárticos de la Argentina y Chile (Diptera: Ceratopogonidae). Neotrópica, 40, 60.
Spinelli, G.R. & Ronderos, M.M. (1997) Description of the male of Culicoides lahillei and revalidation of Culicoides debilipalpis (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 123, 303–309.
Spinelli, G.R. & Ronderos, M.M. (2001a) The male of Culicoides irwini (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 127, 513–517.
Spinelli, G.R. & Ronderos, M.M. (2001b) First record of the genus Bezzia in Chile, with a description of a new species of the venustula group (Diptera: Ceratopogonidae). Revista Chilena de Historia Natural, 74, 751–754.https://doi.org/10.4067/S0716-078X2001000400002
Spinelli, G.R. & Ronderos, M.M. (2005) Description of the male of Culicoides pampaensis Spinelli & Wirth (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 64, 43–45.
Spinelli, G.R. & Ronderos, M.M. (2007) Description of the male of Culicoides dureti (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 66, 77–80.
Spinelli, G.R. & Ronderos, M.M. (2015) First description of the male of Diaphanobezzia patagonica (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 74, 57–60.
Spinelli, G.R., Ronderos, M.M. & Balseiro, E.G. (1989) Análisis de la actividad diaria de dípteros nematocéros en Punta Lara (Pdo. de Ensenada, Prov. de Buenos Aires). II. Ceratopogonidae, con la descripción de una especie nueva en el género Atrichopogon. Limnobios, 2, 733–737.
Spinelli, G.R., Ronderos, M.M. & Cazorla, C.G. (2012d) A new species of Bezzia Kieffer from Argentina (Diptera: Ceratopogonidae). Zootaxa, 3232, 62–68.https://doi.org/10.11646/zootaxa.3232.1.3
Spinelli, G.R., Ronderos, M.M. & Díaz, F. (2021) Two new species and new records of Neotropical Culicoides (Diptera: Ceratopogonidae). Zootaxa, 4915 (3), 401–410.https://doi.org/10.11646/zootaxa.4915.3.8
Spinelli, G.R., Ronderos, M.M. & Díaz, F. (2022a) A new species of Leptoconops Skuse from Argentina, description of the male of L. casali Cavalieri & Chiossone and a key to the Neotropical species (Diptera: Ceratopogonidae). Zootaxa, 5091 (3), 487–494.https://doi.org/10.11646/zootaxa.5091.3.8
Spinelli, G.R., Ronderos, M.M., Díaz, F. & Marino, P.I. (2005a) The bloodsucking biting midges of Argentina (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 100, 137–150.https://doi.org/10.1590/S0074-02762005000200006
Spinelli, G.R., Ronderos, M.M., Donato, M. & Siri, A. (2018) Yungahelea, a new genus of predaceous midge from northwestern Argentina (Culicomorpha: Ceratopogonidae). Anais da Academia Brasileira de Ciências, 90, 137–146.https://doi.org/10.1590/0001-3765201720170079
Spinelli, G.R., Ronderos, M.M. & Funes, A. (2016) Redescription of pupae of the New World predaceous midges, Pachyhelea pachymera (Williston, 1900) and Clastrieromyia dycei Spinelli and Grogan, 1986 (Diptera: Ceratopogonidae). Aquatic Insects, 37, 9–20.https://doi.org/10.1080/01650424.2015.1105377
Spinelli, G.R., Ronderos, M.M. & Grogan, W.L. (2015b) A new species of predaceous midge in the Patagonian genus Austrosphaeromias with a redescription of A. chilensis (Diptera: Ceratopogonidae). Iheringia, série Zoologia, 105, 94–100.https://doi.org/10.1590/1678-47662015105194100
Spinelli, G.R., Ronderos, M.M. & Grogan, W.L. (2022b) Five new species in the predaceous midge genus Macrurohelea Ingram & Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli & Grogan (Diptera: Ceratopogonidae). Zootaxa, 5093 (4), 445–464.https://doi.org/10.11646/zootaxa.5093.4.4
Spinelli, G.R., Ronderos, M.M. & Marino, P.I. (2012e) Redescription of the pupa of Parabezzia balseiroi (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 71, 163–168.https://doi.org/10.7550/rmb.28876
Spinelli, G.R., Ronderos, M.M., Omad, G, Pessacq, P. & Marino, P.I. (2013) A new species of Bezzia Kieffer from Argentinean Patagonia (Diptera: Ceratopogonidae). Annales Zoologici, 63, 263–274.https://doi.org/10.3161/000345413X669568
Spinelli, G.R., Veggiani Aybar, C., Dantur Juri, M.J., Lizarralde de Grosso, L. & Marino, P.I. (2013c) Two new species and new records of biting midges of the genus Culicoides from northwestern Argentina (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 108, 586–589.https://doi.org/10.1590/S0074-02762013000500008
Spinelli, G.R. & Wirth, W.W. (1981): A new species of predaceous midge of the genus Bezzia Kieffer from Argentina (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 40, 187–192.
Spinelli, G.R. & Wirth, W.W. (1984a) Las especies Neotropicales del género Dasyhelea, grupo cincta (Diptera: Ceratopogonidae). Limnobios, 2, 586–608.
Spinelli, G.R. & Wirth, W.W. (1984b) Ocho especies nuevas del género Culicoides Latreille de la región Neotropical. Primera descripción del macho de C. flinti Wirth, y de la hembra de C. lenti Tavares y Luna Dias (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 43, 171–185.
Spinelli, G.R. & Wirth, W.W. (1984c) The Neotropical predaceous midges of the genus Alluaudomyia (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 86, 673–702.
Spinelli, G.R. & Wirth, W.W. (1984d) A review of the Neotropical predaceous midge genus Paryphoconus (Diptera: Ceratopogonidae). Proceedings of the Biological Society of Washington, 97, 882–908.
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