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Type: Article
Published: 2023-05-04
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Cimbicidae (Hymenoptera, ‘Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands

Department of Earth; Ocean and Atmospheric Sciences; University of British Columbia; 2020 – 2207 Main Mall; Vancouver; British Columbia; Canada; V6T 1Z4; Canada; Department of Biological Sciences; Simon Fraser University; 8888 University Drive; Burnaby; British Columbia; V5A 1S6; Canada; Museum of Comparative Zoology; 26 Oxford Street; Cambridge; Massachusetts; 02138; United States of America; Royal British Columbia Museum; 675 Belleville Street; Victoria; British Columbia; V8W 9W2; Canada
A.A. Borissiak Paleontological Institute; Russian Academy of Sciences; Moscow 117647; Russia; Invertebrate Paleontology Department; Natural History Museum; Cromwell Road; London SW7 5BD; United Kingdom
Hymenoptera Cimbicidae Okanagan Highlands Florissant Green River Formation Paleogene

Abstract

We erect the Cenocimbicinae, a new subfamily of Cimbicidae (Hymenoptera, Symphyta), restricted to the Selandian Menat Formation of France, the oldest occurrence of the family, and the Ypresian Okanagan Highlands of far-western North America. We describe new taxa from the Okanagan Highlands: Allenbycimbex morrisae gen. et sp. nov. and Leptostigma n. gen. with seven new species: L. alaemacula n. sp., L. brevilatum n. sp., L. fasciatum n. sp., L. longiclava n. sp., L. longipallidum n. sp., L. longitenebricum n. sp., and L. proxivena n. sp. We revise the Cimbicidae from the Ypresian Green River Formation and the Priabonian Florissant Formation, both in Colorado, USA. The oldest fossil of a modern cimbicid subfamily appears with a single pachylostictine specimen in the Green River Formation, and all Cimbicidae are in modern subfamilies after the Ypresian (we did not examine one larva known from Priabonian Baltic amber). Pseudocimbex clavatus Rohwer 1908 from the Florissant Formation is not a cimbicid; we treat it as Tenthredinoidea incertae sedis. We transfer Cimbex vetusculus Cockerell to Floricimbex n. gen.

 

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