Abstract
The structure and musculature of the male terminalia are described in three type species of the eupitheciine genera Chloroclystis Hübner, Sigilliclystis Galsworthy, and Ziridava Walker (Lepidoptera: Geometridae: Larentiinae). The following traits are characteristic for all three genera: (1) the tergal flexors of the valvae, muscles m4 running to the medial part of the transtilla or membrane anterior to the transtilla; (2) the sternal extensors of the valvae, muscles m8(3), broadly running to the inner wall of the saccus, with long lateral bundles; (3) the twisted muscles m7(6) running from the lower margin of the saccus. The peculiar structure of the transtilla is described and illustrated.
References
- Beljaev, E.A. (2008) Phylogenetic relationships of the family Geometridae and its subfamilies (Lepidoptera). Meetings in memory of N.A. Cholodkovsky, St. Petersburg, 60, 1–238. [in Russian]
- Ebert, G. (2001) Die Schmetterlinge Baden-Württembergs, Band 8—Nachtfalter VI. Stuttgart (Hohenheim), Ulmer, 541 pp.
- Galsworthy, A.C. (1999) New and revised eupitheciine species (Geometridae, Larentiinae) from Hong Kong and South East Asia. Transactions of the Lepidopterological Society of Japan, 50 (3), 223–234.
- Galsworthy, A.C. (2003) Bosara Walker: revisional notes on the Bosara refusaria group of species (Geometridae, Larentiinae). Transactions of the Lepidopterological Society of Japan, 54 (3), 147–155.
- Holloway, J.D. (1977) The Lepidoptera of Norfolk Island, their Biogeography and Ecology. Series Entomologica. Vol. 13. W. Junk, The Hague, 29 b/w photos, 140 figs.
- Holloway, J.D. (1982) Taxonomic Appendix. In: Holloway, J.D. (Ed.), An Introduction to the Moths of South East Asia. The Malaysian Nature Society and E. W. Classey, Kuala Lumpur, pp. 174–271.
- Holloway, J.D. (1997) The Moths of Borneo: family Geometridae, subfamilies Sterrhinae and Larentiinae. Malayan Nature Journal, 51, 1–242.
- Inoue, H. (2002) Notes on the tribe Eupitheciini (Geometridae, Larentiinae) from Japan, Taiwan and Southeast Asia, with description of ten new species. Tinea, 17 (2), 92–112.
- Klots, A.B. (1970) Lepidoptera. In: Tuxen, S.L. (Ed.), Taxonomist’s Glossary of Genitalia in Insects. Munksgaard, Copenhagen, pp. 115–130.
- Kuznetzov, V.I. & Stekolnikov, A.A. (2001) New approaches to the system of Lepidoptera of the world fauna (on the base of functional morphology of abdomen). Proceedings of the Zoological Institute of St. Petersburg, 282, 1–462. [in Russian]
- Marriott, P. (2011) Moths of Victoria, Waves and Carpets—Geometroidea, Part 3. Entomological Society of Victoria, Melbourne, 36 pp.
- McQuillan, P.B. & Edwards, E.D. (1996) Geometridae. In: Nielsen, E.S, Edwards, E.D. & Rangsi, T.V. (Eds.), Checklist of the Lepidoptera of Australia. Monographs on Australian Lepidoptera. Vol. 4. CSIRO, Melbourne, pp. 200–228.
- Mironov, V. (2003) Larentiinae II (Perizomini and Eupitheciini). In: Hausmann, A. (Ed.), The geometrid moths of Europe 4. Appollo Books, Stenstrup, pp. 1–463. https://doi.org/10.1163/9789004308633 DOI: https://doi.org/10.1163/9789004308633
- Nichols, S.W. (Ed.) (1989) The Torre-Bueno Glossary of Entomology. New York Entomological Society in corporation with American Museum of Natural History, New York, 840 pp.
- Pierce, F.N. (1914) The Genitalia of the Group Geometridae of the Lepidoptera of the British Islands. F.N. Pierce, The Elms, Dingle, Liverpool, 230 pp.
- Schmidt, O. (2001) The Australian species of Anachloris Meyrick (Lepidoptera: Geometridae: Larentiinae): taxonomy, male genitalia musculature and systematic position. Australian Journal of Entomology, 40, 219–230. https://doi.org/10.1046/j.1440-6055.2001.00232.x DOI: https://doi.org/10.1046/j.1440-6055.2001.00232.x
- Schmidt, O. (2013) The structure and musculature of male terminalia in the tribe Xanthorhoini Pierce and related tribes (Lepidoptera: Geometridae: Larentiinae), with particular reference to the Palaearctic and Australian regions. Zootaxa, 3721 (6), 552–572. https://doi.org/10.11646/zootaxa.3721.6.3 DOI: https://doi.org/10.11646/zootaxa.3721.6.3
- Schmidt, O. (2014a) Comparison of the structure and musculature of male terminalia in the tribe Cidariini Duponchel (Lepidoptera: Geometridae: Larentiinae) once again throws into doubt a sister relationship with the Xanthorhoini. Zootaxa, 3856 (3), 399–418. https://doi.org/10.11646/zootaxa.3856.3.6 DOI: https://doi.org/10.11646/zootaxa.3856.3.6
- Schmidt, O. (2014b) The structure and musculature of male terminalia in the tribes Eupitheciini, Melanthiini, Perizomini and Rheumapterini: Gnathoi or Agnathoi, that is the question (Lepidoptera, Geometridae, Larentiinae). Spixiana, 37 (2), 267–281.
- Schmidt, O. (2015) One century after: a reappraisal of the gnathos (sensu Pierce, 1914) in Larentiinae (Lepidoptera: Geometridae). Zootaxa, 3974 (3), 328–340. https://doi.org/10.11646/zootaxa.3974.3.2 DOI: https://doi.org/10.11646/zootaxa.3974.3.2
- Schmidt, O. (2017a) Comparison of the structure and musculature of male terminalia in the tribes Operophterini, Phileremini and Triphosini (Lepidoptera: Geometridae: Larentiinae). Zootaxa, 4237 (2), 347–358. https://doi.org/10.11646/zootaxa.4237.2.8 DOI: https://doi.org/10.11646/zootaxa.4237.2.8
- Schmidt, O. (2017b) Myths and facts about the absence of socii in the male genitalia of Larentiinae (Lepidoptera: Geometridae). Zootaxa, 4286 (2), 254–266. https://doi.org/10.11646/zootaxa.4286.2.8 DOI: https://doi.org/10.11646/zootaxa.4286.2.8
- Schmidt, O. (2020) The structure and musculature of male terminalia in the Palaearctic and Indo-Australian species of the eupitheciine genera Eupithecia Curtis, Gymnoscelis Mabille, and Pasiphila Meyrick (Lepidoptera: Geometridae: Larentiinae). Zootaxa, 4896 (2), 224–238. https://doi.org/10.11646/zootaxa.4896.2.4 DOI: https://doi.org/10.11646/zootaxa.4896.2.4
- Schmidt, O. & Tautel, C. (2022) Notes on the Indo-Australian genus Ziridava Walker (Lepidoptera: Geometridae: Larentiinae), with description of two new species. Zootaxa, 5100 (1), 105–118. https://doi.org/10.11646/zootaxa.5100.1.5 DOI: https://doi.org/10.11646/zootaxa.5100.1.5
- Scoble, M.J. (1995) The Lepidoptera: Form, Function and Diversity. Oxford University Press, Oxford, 424 pp.
- Scoble, M.J. (1999) Geometrid Moths of the World: A Catalogue (Lepidoptera, Geometridae). CSIRO Publishing, Collingwood, Victoria, 1200 pp. https://doi.org/10.1071/9780643101050 DOI: https://doi.org/10.1071/9780643101050
- Scoble, M.J. & Hausmann (2007) Online list of valid and available names of the Geometridae of the World. Available from: http://www.lepbarcoding.org/geometridae/species_checklists.php (accessed 20 July 2017)
- Xue, D.Y. & Zhu, H.F. (1999) Fauna Sinica (Insecta: Lepidoptera: Geometridae: Larentiinae). Science Press, Beijing, 1099 pp., 25 pls.