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Issue: Vol. 5289 No. 1: 18 May 2023
Type: Monograph
Published: 2023-05-18
DOI: 10.11646/zootaxa.5289.1.1
Page range: 1-82
Abstract views: 238
PDF downloaded: 34

Taxonomic study of the subfamily Pyraustinae (Lepidoptera: Crambidae)in Iran

Iranian Research Institute of Plant Protection (IRIPP); Agricultural Research; Education and Extension Organization (AREEO); Tehran
Neerland 20; 4614 GD Bergen op Zoom; The Netherlands
Department of Zoology; Swedish Museum of Natural History; Stockholm; Sweden
Račianska 61; SK-83102 Bratislava; Slovak Republic
Lepidoptera Pyraustinae faunistic new synonyms new records Iran

Abstract

Sixty-seven species of the subfamily Pyraustinae from 17 genera and two tribes are listed. Anania verbascalis verbascalis ([Denis & Schiffermüller], 1949) is considered as a senior synonym of A. verbascalis parvalis (Osthelder, 1935) syn. n.; Pyrausta virginalis Duponchel, 1832 as a senior synonym of P. perkeo Amsel, 1970 syn. n.; and P. ferrealis (Hampson, 1900) as a senior synonym of P. euergestalis Amsel, 1954 syn. n. Thirteen species, namely Anania coronata (Hufnagel, 1767), A. murcialis (Ragonot, 1895), A. terrealis (Treitschke, 1829), Pyrausta ferrealis (Hampson, 1900), P. armeniaca Slamka, 2013, P. zeitunalis Caradja, 1916, P. cingulata (Linnaeus, 1758), P. delicatalis Caradja, 1916, P. tatarica Kemal et al., 2020, Loxostege sedakowialis (Eversmann, 1852), L. wagneri Zerny in Wagner, 1929, L. mucosalis (Herrich-Schäffer, 1848) and L. peltaloides (Rebel in Wagner, 1932), are newly reported from Iran. Additionally, a redescription of the male of P. delicatalis and the female of L. sedakowialis, as well as description of the hitherto unknown female of P. delicatalis are presented. The intraspecific variations of the species if present are discussed and additional new diagnostic characters to separate the closely related species are presented. Data on the geographical distribution of the Iranian species of this subfamily are provided.

 

References

  1. Alipanah, H., Malm, T. & Asselbergs, J. (2020) A new species of Sitochroa Hübner, 1825 (Lep.: Crambidae: Pyraustinae), with taxonomic notes on the genus. Nota Lepidopterologica, 43, 61–76. http://doi.org/10.3897/nl.43.49128 DOI: https://doi.org/10.3897/nl.43.49128
  2. Allard, G. (1867) Notes sur les insectes de l'Algérie. Annales de la Société Entomologique de France, Series 4, 7, 311–322, pl. 6.
  3. Amsel, H.G. (1949a) Die Microlepidopteren der Brandt'schen Iran-Ausbeute. 1. Teil. Bulletin de la Société Fouad Ier d’Entomologie, 33, 227–269, 4 pls.
  4. Amsel, H.G. (1949b) On the Microlepidoptera collected by E. P. Wiltshire in Irak and Iran in the years 1935 to 1938. Bulletin de la Société Fouad Ier d’Entomologie, 33, 271–326.
  5. Amesl, H.G. (1950) Die Microlepidopteren der Brandt’schen Iran-Ausbeute. 2. Teil. Arkiv för Zoologi, New Series, 1 (17), 223–257.
  6. Amsel, H.G. (1951) Die Microlepidopteren der Brandt’schen Iran-Ausbeute. 3. Teil. Arkiv för Zoologi, New Series, 1 (36), 525–563.
  7. Amsel, H.G. (1952) Über einige von Hampson beschriebene palaearktische Pyraliden (Lepidoptera, Pyralidae). Mitteilungen der Münchener Entomologischen Gesellschaft, 42, 40–70.
  8. Amsel, H.G. (1954) Die Microlepidopteren der Brandt’schen Iran-Ausbeute. 4. Teil. Arkiv för Zoologi, New Series, 6 (16), 255–326, pls. 1–33.
  9. Amsel, H.G. (1959) Microlepidotera aus Iran. (Ergebnisse der Entomologischen Reisen Willi Richter, Stuttgart, im Iran 1954 und 1956-Nr.25). Stuttgarter Beiträge zur Naturkunde, 28, 1–38.
  10. Amsel, H.G. (1961) Die Microlepidopteren der Brandt’schen Iran-Ausbeute. 5. Teil. Arkiv för Zoologie, New Series, 13 (17), 323–445, pls. 1–9.
  11. Amsel, H.G. (1970) Afghanische Pyraustinae (Lepidoptera: Pyralidae). Beiträge zur Naturkundlichen Forschung in Südwestdeutschland, 29 (1), 25–66, pls. 1–4.
  12. Asselbergs, J. (2008) Order Lepidoptera, superfamily Pyraloidea. In: van Harten, A. (Ed.) Arthropod fauna of the UAE Vol. I. Dar Al Ummah Printing, Publishing, Distribution & Advertising, Abu Dabi, pp. 469–561.
  13. Bethune-Baker, G.T. (1894) Notes one some Lepidoptera received from the neighbourhood of Alexandria. Transactions of the Entomological Society of London, 33–51, pl. 1.
  14. Bremer, O. (1864) Lepidopteren Ost-Sibiriens, insbesondere des Amur-Landes, gesammelt von den Herren G. Radde, R. Maack und P. Wulffius. Mémoires de l'Académie impériale des Sciences de St-Pétersbourg, Series 7, 8 (1), 1–104, pls. 1–8. https://doi.org/10.5962/bhl.title.62141 DOI: https://doi.org/10.5962/bhl.title.62141
  15. Butler, A.G. (1878) Illustrations of typical specimens of Lepidoptera Heterocera in the collection of the British Museum Part 2. The Trustees of British Museum (Natural History), London, 62 pp., 20 pls.
  16. Butler, A.G. (1879) Illustrations of typical specimens of Lepidoptera Heterocera in the collection of the British Museum Part 3. Trustees of British Museum (Natural History), London, xviii + 82 pp., 20 pls.
  17. CABI (2022) Ostrinia nubilalis (European maize borer). Invasive Species Compendium, detailed coverage of invasive species threatening livelihoods and the environment worldwide. Available from: https://www.cabi.org/isc/datasheet/46129 (accessed 12 January 2022)
  18. Caradja, A.V. (1916) Beitrag zur Kenntnis der geographischen Verbreitung der Pyraliden und Tortriciden des europäischen Faunengebietes, nebst Beschreibung neuer Formen. Deutsche entomologische Zeitschrift Iris, 30 (1), 1–88 + 151–152 [errata].
  19. Caradja, A.V. (1925) Ueber Chinas Pyraliden, Tortriciden, Tineiden nebst kurze Betrachtungen, zu denen das Studium dieser Fauna Veranlassung gibt (Eine biogeographische Skizze). Memoriile Sectiunii Stiintifice. Academia Romana, Series 3, 3 (7), 257–383, pls. 1–2.
  20. Caradja, A.V. (1935) Schwedisch-chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Sü Ping-chang. Lepidoptera. 3. Familiae, Pyralidae, Pterophoridae, Tortricidae–Tineidae l.s. Arkiv för Zoologi, 27 A (8), 1–10.
  21. Caradja, A.V. (1938) Materialien zu einer Microlepidopteren-Fauna von Atuntse in Nord-Yünnan. Ein Nachtrag zur gleichbetitelten Arbeit in D.E.Z. "Iris" 1938. Stettiner Entomologische Zeitung, 99 (2), 247–250.
  22. Caradja, A.V. (1939) Materialien zu einer Mikrolepidopterenfauna des Yangtsetales bei Batang. Deutsche Entomologische Zeitschrift Iris, 53, 15–26.
  23. Caradja, A.V. & Meyrick, E. (1935) Materialien zu Einer Microlepidopteren-Fauna der Chinesischen Provinzen Kiangsu, Chekiang und Hunan. R. Friedländer & Sohn, Berlin, 96 pp., 3 pls.
  24. Chrétien, P. (1915) Contribution à la connaissance des Lépidoptères du Nord de l'Afrique. Annales de la Société Entomologique de France, 84 (3), 289–374.
  25. Christoph, H.T. (1873) Weiterer Beitrag zum Verzeichnisse der in Nord-Persien einheimischen Schmetterlinge. Horae Societatis Entomologicae Rossicae, 10 (3), 3–55.
  26. Christoph, H.T. (1876–1877) Sammelergebniss aus Nordpersien, Krasnowodsk in Turkmenien und dem Daghestan. Horae Societatis Entomologicae Rossicae, 12, 181–299, pls. 5–8.
  27. Christoph, H.T. (1881) Neue Lepidopteren des Amurgebietes. Bulletin de la Société Impériale des Naturalistes de Moscou, 56, 1–80.
  28. Christoph, H.T. (1882) Einige neue Lepidopteren aus Russisch-Armenien. Horae Societatis Entomologicae Rossicae, 17, 104–122.
  29. Christoph, H.T. (1888) Zu einigen neuen Lepidopteren des palaearctischen Faunengebietes. Trudy Russkago Entomologiceskago Obscestva, 22, 308–314.
  30. Clarke, J.F.G. (1941) The preparation of slides of the genitalia of Lepidoptera. Bulletin of the Brooklyn Entomological Society, 36, 149–161.
  31. Clarke, J.F.G. (1971) The Lepidoptera of Rapa Island. Smithsonian Contributions to Zoology 56. Smithsonian Institution Press, Washington, 182 pp. https://doi.org/10.5479/si.00810282.56 DOI: https://doi.org/10.5479/si.00810282.56
  32. Clerck, C.A. (1764) Icones Insectorum Rariorum cum Nominibus eorum trivialibus, Locisque e. C. Linnaei Systemi Naturalis Allegatis. s.n., Stockholm, viii + iii + 3 pp., pls. 1–55. [1759–1764]
  33. Costa, A. (1843) Esposizione sommaria delle osservazioni raccolte durante l'anno 1842 intorno allo sviluppo ed apparizione successiva degli insetti nei contorni di Napoli. Annali dell'Accademia degli Aspiranti Naturalisti di Napoli, 1, 37–68.
  34. Costantini, A. (1923) Lepidoptera pro fauna italica nova, additis specierum formarumque novarum descriptionibus. II. Neue Beiträge zur systematischen Insektenkunde, 2 (12), 105–107, pl. 7.
  35. Curtis, J. (1830) s.n. In: British Entomology; being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Vol. VII. London, J. Curtis, pl. 299, 2 unnumbered pp.
  36. Denis, J.N.C.M. & Schiffermüller, I. [(1775)] Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend herausgegeben von einigen Lehrern am k.k. Theresianum. Augustin Bernardi, Wien, frontispiece, 323 pp., 3 pls.
  37. De Prins, J. & De Prins, W. (2011–2021) Afromoths, online database of Afrotropical moth species (Lepidoptera). World Wide Web electronic publication. Available from: http://www.afromoths.net (accessed 14 October 2022)
  38. Dufrane, A. (1957) Microlépidoptères de la faune Belge (huitième note). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 33 (32), 1–16.
  39. Duponchel, P.-A.-J. (1831–1834) Nocturnes 5 (2). Histoire Naturelle des Lépidoptères ou Papillons de France, 8 (2), 5–402, errata, pls. 211–236.
  40. Duponchel, P.-A.-J. (1842–1844) Nocturnes. Supplement aux tomes quatrième et suivants. Histoire naturelle des Lépidoptères ou Papillons de France, Paris, Supplement 4, 1–554, Avertissement, pls. 1–90. [imprint "1842"]
  41. Esper, E.J.C. (1786–1805) s.n. In: Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Vierter Theil. Europaeische Gattungen Die Eulenphalenen. Band II. W. Walther, Erlangen, pp. 1–372 + 373–698 + 1–85, col. Taf. LXXX–CXXV.c; CXXVI–CXCVIII.
  42. Eversmann, E.F. (1842) Quaedam Lepidopterorum species novae, in Rossia orientali observatae, nunc descriptae et depictae. Bulletin de la Société Impériale des Naturalistes de Moscou, 15 (3), 543–565, pls. 5–6.
  43. Eversmann, E.F. (1852) Mittheilung über einige neue Falter Russlands. Bulletin de la Société Impériale des Naturalistes de Moscou, 25 (1), 148–169.
  44. Fabricius, J.C. (1775) Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. Kortii, Flensburgi et Lipsiae, 832 pp. https://doi.org/10.5962/bhl.title.36510 DOI: https://doi.org/10.5962/bhl.title.36510
  45. Fabricius, J.C. (1781–[1782]) Species insectorum exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus. Vol. 2. Carol. Ernest. Bohnii, Hamburgi et Kilonii, pp. 1–494 + 495–514 (appendix) + 515–517 (index). https://doi.org/10.5962/bhl.title.36509 DOI: https://doi.org/10.5962/bhl.title.36509
  46. Fabricius, J.C. (1787) Mantissa Insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus. Vol. 2. Christian Gottlieb Proft, Hafniae, 382 pp.
  47. Fabricius, J.C. (1794) Entomologica systematica emendata et aucta. Secundum classes, ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. C. G. Proft et C. F. Mohr, Hafniae et Kiliae, 349 pp.
  48. Fabricius, J.C. (1798) Supplementum Entomologiae Systematicae. Proft et Storch, Hafniae, 572 pp.
  49. Fawcett, J.M. (1916) Notes on a collection of Heterocera made by Mr. W. Feather in British East Africa, 1911–13. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London, 707–737, pl. 1. https://doi.org/10.1111/j.1096-3642.1916.tb02043.x DOI: https://doi.org/10.1111/j.1096-3642.1916.tb02043.x
  50. Fourcroy, A.F. (1785) Entomologia Parisiensis; Sive catalogus insectorum quae in agro Parisiensi reperiuntur; secundum methodum Geoffraeanum in sectiones, genera & species distributus: Cui addita sunt nomina trivialia & fere trecentae novae species, 1–2, i–viii + 1–232 + 233–544. https://doi.org/10.5962/t.174486 DOI: https://doi.org/10.5962/t.174486
  51. Gaede, M. (1917) Neue Lepidopteren des Berliner Zoologischen Museums. I. Aethiopische Pyralididen. Mitteilungen aus dem Zoologischen Museum in Berlin, 8 (3), 387–401.
  52. Gao, Q., Zhang, D.-D. & Wang, S.-X. (2013) Taxonomic study of the genus Ecpyrrorrhoe Hübner (Lepidoptera, Crambidae, Pyraustinae) from China, with descriptions of three new species. Acta Zootaxonomica Sinica, 38 (2), 311–316.
  53. Guenée, M.A. (1854) Deltoïdes et Pyralites. In: Boisduval, J.B.A.D. de & Guenée, M.A. (Eds.), Histoire Naturelle des Insectes. Species Général des Lépidoptères 8. Librairie encyclopédique Roret, Paris, pp. 1–448.
  54. Hampson, G.F. (1900) New palaearctic Pyralidae. Transactions of the entomological Society of London, 68 (3), 369–401. https://doi.org/10.1111/j.1365-2311.1900.tb02380.x DOI: https://doi.org/10.1111/j.1365-2311.1900.tb02380.x
  55. Hampson, G.F. (1913) Descriptions of new species of Pyralidae of the subfamily Pyraustinae. Annals and Magazine of Natural History, including Zoology, Botany and Geology, Series 8, 11 (63 + 66), 322–342 + 509–530. https://doi.org/10.1080/00222931308693351 DOI: https://doi.org/10.1080/00222931308693351
  56. Hartig, F. & Amsel, H.G. (1952) Lepidoptera Sardinica. Fragmenta Entomologica, 1951 (1), 1–152, pls. 1–7.
  57. Haslberger, A. & Segerer, A.H. (2016) Systematische, revidierte und kommentierte Checkliste der Schmetterlinge Bayerns (Insecta: Lepidoptera). Mitteilungen der Münchner Entomologischen Gesellschaft, 106 (Supplement), 1–336.
  58. Haworth, A.H. (1803–1828) Lepidoptera Britannica: sistens digestionem novam insectorum lepidopterorum quae in Magna Britannia reperiuntur, larvarum pabulo, remporeque pascendi, expansione alarum, mensibusque volandi, synonymis atque locis observationibusque variis: etc. R. Taylor, London, 4 + xxxvi + 609 pp. DOI: https://doi.org/10.1080/14786442808674667
  59. Haworth, A.H. (1811) Lepidoptera Britannica, sistens digestionem novam insectorum lepidopterorum quae in Magna Britannia reperiuntur, larvarum pabulo, temporeque pascendi; expansione alarum, mensibusque volandi, synonymis. R. Taylor, London, 512 pp.
  60. Herrich-Schäffer, G.A.W. (1847–1855) Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner‘s Sammlung europäischer Schmetterlinge. 4: Die Zünsler und Wickler. G. J. Manz, Regensburg, 288 pp.
  61. Hübner, J. (1793) Sammlung auserlesener Vögel und Schmetterlinge, mit ihren Namen herausgegeben auf hundert nach der Natur ausgemalten Kupfern. Gedruckt mit Brinnhaußerischen Schriften, Augsburg, 16 pp., 100 pls.
  62. Hübner, J. (1796–1833) Sammlung europäischer Schmetterlinge. 6. Horde. Die Zünsler; nach der Natur geordnet, beschrieben und vorgestellt (continued by C. Geyer). Verlag J. Hübner, Augsburg, pp. [i]–[iv] + [i–ii] + [i–ii] + 1–30 + [i–ii] + [i–ii], pls. 1–32. [imprint “1796”]
  63. Hübner, J. (1796–1836) Sammlung europäischer Schmetterlinge. 8. Horde. Die Schaben; nach der Natur geordnet, beschrieben und vorgestellt (continued by C. Geyer). Verlag J. Hübner, Augsburg, pp. [1]–[12]–13–70–[71]–[78], pls. 1–71. [imprint “1796”]
  64. Hübner, J. (1816–1826) s.n. In: Verzeichniß bekannter Schmettlinge [sic] bey dem Verfasser zu finden. bey dem Verfasser zu Finden. s.n., Augsburg, pp. [1]–[3]–4–6–[7]–8–431 (Verzeichniß) + [1]–2–72 (Anzeiger). [imprint “1816”]
  65. Hufnagel, J.S. (1767) Fortsetzung der Tabelle von den Nachtvögeln, welche die 3te Art derselben, nehmlich die Spannenmesser (Phalaenas Geometras Linnaei) enthält. Berlinisches Magazin, 4 (5 + 6), 504–527 + 599–626.
  66. Inoue, H. (1988) A new species of Epipaschiinae from Japan, with some synonymic notes on the Pyralidae from east Asia (Lepidoptera). Tinea, 12 (10), 85–95.
  67. Jong, Y. de, Verbeek, M., Michelsen, V., Bjørn, P.P., Los, W., Steeman, F., Bailly, N., Basire, C., Chylarecki, P., Stloukal, E., Hagedorn, G., Wetzel, F.T., Glöckler, F., Kroupa, A., Korb, G., Hoffmann, A., Häuser, C., Kohlbecker, A., Müller, A., Güntsch, A., Stoev, P. & Penev, L. (2014) Fauna Europaea—all European animal species on the web. Biodiversity Data Journal, 2, e4034. https://doi.org/10.3897/BDJ.2.e4034 DOI: https://doi.org/10.3897/BDJ.2.e4034
  68. Karsholt, O. & Razowski, J. (1996) The Lepidoptera of Europe. A distributional checklist. Apollo Books, Stenstrup, 380 pp.
  69. Koçak, A.Ö. (1980) Some notes on the nomenclature of Lepidoptera. Communications de la Faculté des Sciences de l'Université d'Ankara, Séries C3, Zoologie, 24, 7–25. DOI: https://doi.org/10.1501/Commuc_0000000022
  70. Koçak, A.Ö. (1981) Further notes on the homonymy of the specific names of Lepidoptera. The Centre for Entomological Studies Ankara Priamus, 1 (1), 11–17.
  71. Koçak, A.Ö. & Kemal, M. (2012) List of the hitherto recorded pterygot taxa of Turkey (Insecta) (temporary report of the Entomofauna Projekt of Turkey 10). The Centre for Entomological Studies Ankara, Memoirs, 6, i–iv + 1–1649.
  72. Koçak, A.Ö. & Kemal, M. (2014) Lepidoptera of Iran based upon the Info-system of the Cesa. The Centre for Entomological Studies Ankara, Priamus, 31 (Supplement), 1–487.
  73. Kemal, M. & Koçak, A.Ö. (2017) Description of a new species, Loxostege ayhanana sp. n. from East Turkey (Lepidoptera, Pyraloidea). The Centre for Entomological Studies Ankara, Miscellaneous Papers, 164, 1–5, figs. 1–7.
  74. Kemal, M. & Koçak, A.Ö. (2018) Vernal aspect of the Lepidoptera from Nemrut Mountain and the vicinity of Karadut in the Kahta district with a description of a new species (SE Turkey, Adýyaman Province). The Centre for Entomological Studies Ankara, Miscellaneous Papers, 174, 1–21.
  75. Kemal, A., Kýzýldað, S. & Koçak, A.Ö. (2020) Description of a new Pyrausta species from East Turkey with comments about related species (Lepidoptera, Pyraloidea). The Centre for Entomological Studies Ankara, Miscellaneous Papers, 210, 1–5.
  76. Kollar, V. & Redtenbacher, L. (1850) Ueber den Charakter der Insecten-Fauna von Südpersien. Denkschriften der Kaiserlichen Akademie der Wissenschaften (Mathematisch-Naturwissenschaftliche Klasse), 1, 42–53.
  77. Krulikovsky, L. (1928) s.n. Zbirnyk prats Zoolohichnoho Muzeyu, Kiev, 5, 212.
  78. La Harpe, J.-J.C. de (1864) Premier supplément aux Pyralidides et aux Crambides de la faune suisse. Neue Denkschriften der Schweizerischen Naturforschenden Gesellschaft, 14, 30–55.
  79. Landry, B. (2015) The Pyraustinae (Lepidoptera, Pyralidae s. l.) of the Galápagos Islands, Ecuador. Revue suisse de Zoologie, 122 (1), 55–70.
  80. Lederer, J. (1858) Noch einige syrische Schmetterlinge. Wiener Entomologische Monatschrift, 2 (5), 135–152, pls. 2–4.
  81. Lederer, J. (1863) Beitrag zur Kenntniss der Pyralidinen. Wiener Entomologische Monatschrift, 7 (8 + 10–12), 243–280 + 331–504, pls. 2–18.
  82. Lederer, J. (1869) Verzeichniss der von Herrn Jos. Haberhauer bei Astrabad in Persien gesammelten Schmetterlinge. Horae Societatis Entomologicae Rossicae, 6, 6–93, 5 pls.
  83. Lederer, J. (1871) Nachtrag zum verzeichnisse Verzeichniss der von Herrn Jos. Haberhauer bei Astrabad in Persien gesammelten Schmetterlinge. Horae Societatis Entomologicae Rossicae, 8, 3–28.
  84. Leech, J.H. & South, R. (1901) Lepidoptera Heterocera from China, Japan, and Corea. Part V. Transactions of the Entomological Society of London, 385–514, pls. 14–15. https://doi.org/10.1111/j.1365-2311.1901.tb01371.x DOI: https://doi.org/10.1111/j.1365-2311.1901.tb01371.x
  85. Leraut, P.J.A. (1996) Contribution à l'étude des Pyrales de la Faune de France (Lepidoptera, Crambidae). Alexanor, 19 (4), 215–228.
  86. Leraut, P.J.A. (2001) Contribution à l'étude des pyrales paléarctiques (Lepidoptera, Pyraloidea). Nouvelle Revue d'Entomologie, 18 (2), 173–185.
  87. Leraut, P.J.A. (2012) Zygènes, Pyrales 1 et Brachodides. Papillons de nuit d'Europe, Verrières-le-Buisson, 3, 1–599.
  88. Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Laurentii Salvii, Holmiae, 824 pp. https://doi.org/10.5962/bhl.title.542 DOI: https://doi.org/10.5962/bhl.title.542
  89. Linnaeus, C. (1761) Fauna Svecica sistens Animalia sveciae regni: Mammalia, Aves, Amphibia, Pisces, Insecta, Vermes. Distributa per Classes & Ordines, Genera & Species cum Differentiis Specierum, Synonymis Auctorum, Nominibus Incolarum, Locis Natalium, Descriptionibus Insectorum. Laurentii Salvii, Stockholm, [43] + 578 pp., 2 pls. https://doi.org/10.5962/bhl.title.46380 DOI: https://doi.org/10.5962/bhl.title.46380
  90. Linnaeus, C. (1767) Systema naturae: per regna tria natura, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio 12. Laurentii Salvii, Holmiae, 794 pp. https://doi.org/10.5962/bhl.title.156772 DOI: https://doi.org/10.5962/bhl.title.559
  91. Linnaeus, C. (1771) Mantissa Plantarum altera Generum editionis VI & Specierum editionis II. Impensis Direct. Laurentii Salvii, Holmiae, pp. i–viii + 144–587.
  92. Lucas, D. (1942) Contribution à l’étude des Lépidoptères du Maroc. Bulletin de la Société des Sciences Naturelles du Maroc, 21, 61–66.
  93. Maes, K.V.N. (2002) On the systematic positions of some Palaearctic Pyraustinae. Nota lepidopterologica, 24 (2001) (4), 51–57.
  94. Mally, R., Hayden J.E., Neinhuis, C., Jordal, B.H. & Nuss, M. (2019) The phylogenetic systematics of Spilomelinae and Pyraustinae (Lepidoptera: Pyraloidea: Crambidae) inferred from DNA and morphology. Arthropod Systematics and Phylogeny, 77, 141–204. https://doi.org/10.26049/asp77-1-2019-07
  95. Mann, J. (1862) Verzeichniss der im Jahre 1851 bei Brussa in Kleinasien gesammelten Schmetterlinge. Wiener Entomologische Monatschrift, 6 (11 + 12), 356–371 + 373–409, 3 pls.
  96. Marion, H. (1952) Ebauche d‘une classification nouvelle des Pyraustidae. Revue française de Lépidoptérologie, 13, 260–270.
  97. Meagher, R.L., Brambila, J. & Hung, E. (2008) Monitoring for exotic Spodoptera species (Lepidoptera: Noctuidae) in Florida. Florida Entomologist, 91 (4), 517–522. https://doi.org/ 10.1653/0015-4040-91.4.517 DOI: https://doi.org/10.1653/0015-4040(2008)91[512:SDOAAS]2.0.CO;2
  98. Meyrick, E. (1936–1937) Exotic Microlepidoptera. Taylor and Francis, London, 160 pp.
  99. Mirzayans, H. & Kalali, G.H. (1970) Contribution a la connaissance de la faune des Lépidoptères de l’Iran. Applied Entomology and Phytopathology, 29, 15–23.
  100. Möschler, H.B. (1888) Botys retowskyi n. sp. Stettiner Entomologische Zeitung, 49, 128.
  101. Müller, O.F. (1764) Fauna Insectorum Fridrichsdalina, sive methodica descriptio insectorum agri Fridrichsdalensis, cum characteribus genericis et specificis, nominibus trivialibus, locis natalibus, iconibus allegatis, novisque pluribus speciebus additis. Gleditsch, Hafniae et Lipsiae, xxiv + 96 pp.
  102. Müller, O.F. (1730–1784) Zoologiae Danicae prodromus: seu Animalium Daniae et Norvegiae indigenarum; characteres, nomina, et synonyma imprimis popularium. V. Insecta. 3. Lepidoptera. typis Hallageriis, Havniae, pp. 111–139. [1776] https://doi.org/10.5962/bhl.title.13268 DOI: https://doi.org/10.5962/bhl.title.13268
  103. Munroe, E.G. (1976) Pyraloidea Pyralidae comprising the subfamily Pyraustinae tribe Pyraustini (part). In: Dominick, R.B. (Ed.), The Moths of America north of Mexico. Fascicle 13.2A. E.W. Classey Ltd. and The Wedge Entomological Research Foundation, London, pp. 1–78, pls. 1–4 + A–H.
  104. Munroe, E.G. & Mutuura, A. (1968) Contributions to a study of the Pyraustinae (Lepidoptera: Pyralidae) of temperate East Asia III. The Canadian Entomologist, 100 (9), 974–985. DOI: https://doi.org/10.4039/Ent100974-9
  105. Mutuura, A. (1954) Classification of the Japanese Pyrausta group based on the structure of the male and female genitalia (Pyr.: Lep.). Bulletin of the Naniwa University, Series B, 4, 7–33.
  106. Mutuura, A. & Munroe, E.G. (1970) Taxonomy and distribution of the European corn borer and allied species: genus Ostrinia [Hübner] (Lepidoptera: Pyralidae). Memoirs of the Entomological Society of Canada, 71, 1–112. DOI: https://doi.org/10.4039/entm10271fv
  107. Nolcken, J.H.W. (1848) Ueber Botys venosalis Lienig n. sp.. Arbeiten des naturforschenden Vereins zu Riga, 1 (3–4), 283–289, pl. 1, fig. 10.
  108. Nuss, M., Landry, B., Mally, R., Vegliante, F., Tränkner, A., Bauer, F., Hayden, J., Segerer, A., Schouten, R., Li, H., Trofimova, T., Solis, M.A., De Prins, J. & Speidel, W. (2003–2022) Global Information System on Pyraloidea. Available from: http://www.pyraloidea.org (accessed 16 July 2022)
  109. Oberthür, C. (1888) Faunes entomologiques. Descriptions d'insectes nouveaux ou peu connus. Études d'Entomologie, 12, 1–44, pls. 1–7.
  110. Poltavsky, A.N. & Ilyina, E.V. (2016) New records of some rare Noctuoidea and Pyraloidea in Daghestan Republic (Russia). Entomofauna, 37 (16), 265–280.
  111. Popescu-Gorj, A. & Constantinescu, A. (1977) Revision of the genus Euclasta Lederer (Lepidoptera, Pyraustinae). A taxonomic and zoogeographic study. Travaux du Muséum d'Histoire Naturelle "Grigore Antipa", 18, 157–245.
  112. Panzer, G.W.F. (1804) Systematische Nomenclatur über weiland Dr. Jacob Christian Schäffers natürlich ausgemalte Abbildungen regensburgischer Insekten (D. Jacobi Christiani Schaefferi Iconum Insectorum circa Ratisbonam indi. Johann Jakob Palm, Erlangen, xvi + 260 pp. [ibidem Vol. 3, pls. 201–280 + i–xiv] https://doi.org/10.5962/bhl.title.101964 DOI: https://doi.org/10.5962/bhl.title.101964
  113. Ragonot, E.L. (1895) Descriptions d'un genre et de deux espèces inédites de Pyralites provenant d'Espagne. Bulletin de la Société entomologique de France, 64, xxii–xxiii.
  114. Rebel, H. (1901) Famil. Pyralidae–Micropterygidae. In: Staudinger, O. & Rebel, H. (Eds.), Catalog der Lepidopteren des palaearctischen Faunengebietes. Vol. 22. R. Friedländer & Sohn, Berlin, pp. 1–368.
  115. Rebel, H. (1902) Neue Pyraliden, Pterophoriden und Tineen des palaearctischen Faunengebietes. Deutsche entomologische Zeitschrift Iris, 15, 100–126, 3 text figs., pl. 4.
  116. Rebel, H. (1917) Lepidopteren aus Neumontenegro. Sitzungsberichte der Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I. Mineralogie, Biologie, Erdkunde und Verwandte Wissenschaft, 126, 765–813.
  117. Regier, J.C., Mitter, C., Solis, M.A., Hayden, J.E., Landry, B., Nuss, M., Simonsen, T.J., Yen, S.-H., Zwick, A. & Cummings, M.P. (2012) A molecular phylogeny for the pyraloid moths (Lepidoptera: Pyraloidea) and its implications for higher-level classification. Systematic Entomology, 37 (4), 635–656. http://doi.org/ 10.1111/j.1365-3113.2012.00641.x DOI: https://doi.org/10.1111/j.1365-3113.2012.00641.x
  118. Retzius, A.J. (1783) s.n. In: Caroli De Geer (...) Genera et species insectorum e generosissimi auctoris scriptis extraxit, digessit, latine quoad partem reddidit, et terminologiam insectorum Linneanam addidit. Siegfried Lebrecht Crusium, Lipsiae, pp. 7–220 + 1–32.
  119. Reutti, C. (1853) Übersicht der Lepidopteren-Fauna des Grossherzogthum’s Baden. Beiträge zur Rheinischen Naturgeschichte (herausgegeben von der Gesellschaft zur Beförderung der Naturwissenschaften zu Freiburg im Breisgau), 3, I–VIII + 1–216, 1 pl.
  120. Robinson, G.S. (1976) The preparation of slides of Lepidoptera genitalia with special reference to the Microlepidoptera. Entomologist's Gazette, 27, 127–132.
  121. Romanoff, N.M. (1887) Les Lépidoptères de la Transcaucasie. Troisième partie. In: Romanoff, N.M. (Ed.), Mémoires sur les Lépidoptères 33. Imprimerie de M.M. Stassuléwitch, St. Petersburg, pp. 1–49, pls. 1–2.
  122. Rothschild, L.W. (1921) Captain Angus Buchanan’s Aïr expedition. V. On the Lepidoptera collected by Captain A. Buchanan in northern Nigeria and the southern Sahara in 1919–1920. Novitates Zoologicae, 28, 142–170 + 215–229.
  123. Rungs, C.E.E. (1979) Catalogue Raisonné des Lépidoptères du Maroc. Inventaire faunistique et observations écologiques. Tome I. Travaux de l’Institute Scientifi que, Série Zoologie, Rabat, 39, [i]–[x] + 1–244, 2 maps.
  124. Seizmair, M. (2022) The presence of the genus Pyrausta Schrank, 1802 (Lepidoptera, Crambidae, Pyraustinae) on the Arabian Peninsula—faunistic and taxonomic notes with description of a new species. International Journal of Zoological and Entomological Letters, 2 (1), 36–41 https://doi.org/10.22271/letters.2022.v2.i1a.30 DOI: https://doi.org/10.22271/letters.2022.v2.i1a.30
  125. Schawerda, K. (1932) Phlyctaenodes osthelderi species nova. Mitteilungen der Münchener Entomologischen Gesellschaft, 22, 12–13.
  126. Scopoli, J.A. (1763) Entomologia Carniolica exhibens insecta Carnioliae indigena et distributa in ordines, genera, species, varietates, methodo Linneana. Joannis Thomae Trattner, [2] + [8] + [22] + 420 + [4] pp., [43] pls. https://doi.org/10.5962/bhl.title.119976 DOI: https://doi.org/10.5962/bhl.title.34434
  127. Slamka, F. (2013) Pyraloidea of Europe, Pyraustinae and Spilomelinae. Vol. 3. František Slamka, Bratislava, 357 pp.
  128. Snellen, P.C.T. (1890) A catalogue of the Pyralidina of Sikkim collected by Henry J. Elwes and the late Otto Möller, with notes by H. J. Elwes. Transaction of the Entomological Society of London, 557–647, pls. 14–20. https://doi.org/10.1111/j.1365-2311.1890.tb03031.x DOI: https://doi.org/10.1111/j.1365-2311.1890.tb03031.x
  129. Solis, M.A. & Maes, K.V.N. (2003) Preliminary phylogenetic analysis of the subfamilies of Crambidae (Pyraloidea Lepidoptera). Belgian Journal of Entomology, 4, 53–95.
  130. Speidel, W. & Hanigk, H. (1990) Revision der paläarktischen Arten der Gattung Algedonia (Lepidoptera, Crambidae). Bonner Zoologische Beiträge, 41 (3–4), 259–275.
  131. Staudinger, O. (1859) Diagnosen nebst kurzen Beschreibungen neuer andalusischer Lepidopteren. Entomologische Zeitung, 20 (7–9), 211–259.
  132. Staudinger, O. (1867) Einige neue Lepidopteren (sämmtlich aus der Sammlung des verstorbenen O. Gruner). Stettiner Entomologische Zeitung, 28 (1–3), 100–110.
  133. Staudinger, O. (1879–1880) Lepidopteren-Fauna Kleinasien's (Fortsetzung). Horae Societatis entomologicae Rossicae, 15, 159–435. [pp. 159–368: 13 November 1879; pp. 369–435: 13 February 1880]
  134. Staudinger, O. (1892) Lepidopteren des Kentei-Gebietes. Deutsche entomologische Zeitschrift Iris, 5 (2), 300–393.
  135. Ström, H. (1768) s.n. In: Beskrivelse over Norske Insecter. Det Kongelige Norske Videnskabers Selskabs Skrifter, Kjobenhavn, 4, pp. 313–371, pls. 16.
  136. Swinhoe, C. (1884) On Lepidoptera collected at Kurrachee. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London, 503–529, pls. 47–48. https://doi.org/10.1111/j.1096-3642.1884.tb02862.x DOI: https://doi.org/10.1111/j.1096-3642.1884.tb02862.x
  137. Szent-Ivány, J. & Uhrik-Meszáros, T. (1942) Die Verbreitung der Pyralididen (Lepidopt.) im Karpatenbecken. Ergänzende Angaben, Berichtigungen, kritische Bemerkungen zur "Fauna Regni Hungariae". Annales Historico-Naturales Musei Nationalis Hungarici, 35, 105–196, pls. 4.
  138. Thunberg, C.P. (1784) Dissertatio Entomologica sistens Insecta Suecica, quorum Partem Primam, Resp. Johannes Borgstroem. J. Edman, Upsaliae, [6] + 24 pp., 1 pl.
  139. Toll, S. (1936) Untersuchung der Genitalien bei Pyrausta purpuralis L. und P. ostrinalis Hb., nebst Beschreibung 11 neuer Microlepidopteren-Arten. Annales Musei Zoologici Polonici, 11 (24), 403–413, pls. 47–49.
  140. Toll, S. (1948) Beitrag zur Mikrolepidopterenfauna von Nordost-Persien. Zeitschrift der Wiener Entomologischen Gesellschaft, 32, 107–116, pls. 4–6.
  141. Tränkner, A. & Nuss, M. (2010) Anania ochrofascialis (Christoph) comb. n. and A. murcialis (Ragonot) comb. n.—two vicarious species from the western Palaearctic region (Pyraloidea: Crambidae: Pyraustinae). Nota lepidopterologica, 33 (1), 59–65.
  142. Treitschke, F. (1828) Die Schmetterlinge von Europa (Fortsetzung des Ochsenheimer‘schen Werks). Gerhard Fleischer, Leipzig, 319 pp.
  143. Treitschke, F. (1829) Die Schmetterlinge von Europa (Fortsetzung des Ochsenheimer‘schen Werks). Gerhard Fleischer, Leipzig, vi + 252 pp.
  144. Treitschke, F. (1835) Die Schmetterlinge von Europa (Fortsetzung des Ochsenheimer‘schen Werks). Ernst Fleischer, Leipzig, 302 pp.
  145. Villers, C. de (1789) Caroli Linnaei entomologia, faunæ Suecicæ descriptionibus aucta. Piestre & Delamollière, Lyon [Lugduni], xvi + 656 pp.
  146. Wagner, F. (1929) Weiterer Beitrag zur Lepidopteren-Fauna Inner-Anatoliens. Mitteilungen der Münchener Entomologischen Gesellschaft, 19 (10–12), 175–206, pl. 2.
  147. Wagner, F. (1932) Vierter (V.) [sic] Beitrag zur Lepidopteren-Fauna Inner-Anatoliens. Internationale Entomologische Zeitschrift, 26 (18), 185–191.
  148. Walker, F. (1859) List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XVI—Deltoides, XVII—Pyralides, Part XVIII—Pyralides, Part XIX—Pyralides. The Trustees of the British Museum (Natural History), London, iv + 1036 pp.
  149. Walker, F. (1866) List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XXXIV. Supplement Part 4. The Trustees of the British Museum (Natural History), London, pp. i–iv + 1121–1533.
  150. Warren, W. (1892) Descriptions of new genera and species of Pyralidae contained in the British Museum collection. Annals and Magazine of Natural History, including Zoology, Botany and Geology, Series 6, 9, 172–179 + 294–302 + 389–397 + 429–442. https://doi.org/10.1080/00222939208677357 DOI: https://doi.org/10.1080/00222939208677344
  151. Wieser, C., Humer, P. & Stangelmaier, G. (2002) Schmetterlinge (Lepidoptera). In: Gutleb, B. & Weiser, C. (Eds.), Ergebnisse einer zoologischen Exkursion in den Nordiran, 2001 Arthopoda (Lepidoptera, Trichoptera, Neuroptera, Heteroptera, Coleoptera, Opiliones, Araneae, Decapoda) und Vertebrata (Amphibia, Reptilia, Aves, Mammalia). Carinthia II, 192 (112), 52–81.
  152. Wiltshire, E.P. (1977) The scientific results of the Oman flora and fauna survey 1975. Lepidoptera: Part I Families Cossidae, Pyralidae, Geometridae, Sphingidae, Arctiidae, Lymantriidae and Noctuidae. Part II A list of further Lepidoptera-Heterocera from Oman. Part III Pyral. Journal of Oman Studies Special Report, 1, 155–178.
  153. Wocke, M. (1871) Microlepidoptera. In: Staudinger, O. & Wocke, M. (Eds.), Catalog der Lepidopteren des Europäischen Faunengebietes. Dr. O. Staudinger & Königl. Hofbuchhandlung von Hermann Burdach, Dresden, pp. i–xvi + 1–192.
  154. Yepishin, V. (2022) A new synonym of the Pyraloidea from the genus Loxostege Hübner, 1825 (Lepidoptera: Crambidae: Pyraustinae). Zootaxa, 5134 (3), 448–450. https://doi.org/10.11646/zootaxa.5134.3.9 DOI: https://doi.org/10.11646/zootaxa.5134.3.9
  155. Zeller, P.C. (1847) Bemerkungen über die auf einer Reise nach Italien und Sicilien beobachteten Schmetterlingsarten. Isis von Oken, 2–4 + 6–12, 121–159 + 213–233 + 284–308 + 401–457 + 481–522 + 561–594 + 639–667.
  156. Zeller, P.C. (1852) Lepidoptera Microptera, quae J. A. Wahlberg in Caffrorum terra collegit. Kungliga Svenska Vetenskapsakademiens Handlingar, Seires 3, 40, 1–120.
  157. Zerny, H. (1914) Über paläarktische Pyraliden des k. k. naturhistorischen Hofmuseums in Wien. Annalen des K. K. Naturhistorischen Hofmuseums, 28, 295–348.
  158. Zerny, H. (1939) Mikrolepidopteren aus dem Elburs-Gebirge in Nord-Iran. Zeitschrift des Wiener Entomologen-Vereins, 24, 129–138 + 148–149 + 159–160 + 171–175.
  159. Zerny, H. (1940) Microlepidopteren aus dem Elburs-Gebirge in Nord-Iran. Zeitschrift des Wiener Entomologen-Vereins, 25, 20–24 + 42–48.
  160. Zetterstedt, J.W. (1839) Proëmium Lepidopterorum Ordo Entomologorum scopulus. Latr. Conspectus Familiarum et Generum Lepidopterorum Lapponiae. In: Zetterstedt, J.W. (Ed.), Insecta Lapponica. Voss, Lipsiae, pp. 869–1014.
  161. Zhang, D., Cai, Y. & Li, H. (2014) Taxonomic review of the genus Paratalanta Meyrick, 1890 (Lepidoptera: Crambidae: Pyraustinae) from China, with descriptions of two new species. Zootaxa, 3753 (2), 118–132. DOI: https://doi.org/10.11646/zootaxa.3753.2.2
  162. Ziegler, H. (2005–2019) Butterflies & Moths of Palaearctic Regions. Available from: http://www.euroleps.ch/seiten/s_art.php?art=pyr_scalaralis (accessed 27 January 2020)