Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5290 No. 1: 18 May 2023
Type: Monograph
Published: 2023-05-18
DOI: 10.11646/zootaxa.5290.1.1
Page range: 1-73
Abstract views: 422
PDF downloaded: 29

A key to European species of Hyposoter Förster, 1869(Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species

Scientific Associate; Department of Life Sciences; Natural History Museum; Cromwell Road; London SW7 5BD; UK
National Museums of Scotland; Chambers Street; Edinburgh EH1 1JF; U.K.
Ålavikvegen 4; 4250 Kopervik; Norway
Hymenoptera Taxonomy parasitoid host Lepidoptera distribution new species

Abstract

An identification key is presented to European species of Hyposoter falling morphologically within the definition of Townes (1970). Eighteen new species are described: Hyposoter aglyphus Galsworthy & Shaw sp. nov., H. albosignatus Galsworthy & Shaw sp. nov., H. castaneus Galsworthy & Shaw sp. nov., H. flavicoxa Galsworthy & Shaw sp. nov., H. flaviventer Galsworthy & Shaw sp. nov., H. heidiella Galsworthy & Haraldseide sp. nov., H. horstmanni Galsworthy & Shaw sp. nov., H. jubator Galsworthy & Shaw sp. nov., H. karmensis Galsworthy & Haraldseide sp. nov., H. monensis Galsworthy & Shaw sp. nov., H. morairae Galsworthy & Shaw sp. nov., H. naso Galsworthy & Shaw sp. nov., H. pechipogator Galsworthy & Shaw sp. nov., H. pseudodidymator Galsworthy & Shaw sp. nov., H. pseudovalidus Galsworthy & Shaw sp. nov., H. rivulator Galsworthy & Shaw sp. nov., H. ruficoxator Galsworthy & Shaw sp. nov., H. rufonigrator Galsworthy & Shaw sp. nov. Angitia alpicola Smits van Burgst, 1914 is transferred to Hyposoter comb. nov. H. carbonarius insulator Aubert, 1960 is raised to species level stat. nov. H. dubitatus (Holmgren, 1860) is returned to full species status from synonymy with H. sicarius (Gravenhorst, 1829) stat. rev. The following new synonymies are proposed: Limneria obscurella Holmgren, 1860 of Limneria neglecta Holmgren, 1860 syn. nov.; Anilasta berberatae Habermehl, 1922 of Campoplex seniculus Gravenhorst, 1829 syn. nov.; Hyposoter nigrior Aubert, 1993 of Anilasta tibialis Hedwig, 1938 syn. nov.; Rhythmonotus singularis Schmiedeknecht, 1909 of Campoplex tricolor Ratzeburg, 1844 syn. nov.; Hyposoter postcaedator Aubert, 1964 of Hyposoter alpicola Smits van Burgst, 1914 syn. nov. Lectotypes are designated for the following names: Hyposoter postcaedator Aubert, 1964; Hyposoter carbonarius insulator Aubert, 1960; Hyposoter rufovariatus meridionellator Aubert, 1965; Hyposoter praecaedator Aubert, 1963; Hyposoter rapacitor Aubert, 1971. Notes are provided on all included species.

 

References

  1. Aubert, J.-F. (1960) Description préliminaire de quelques espѐces et sous-espѐces méditeranéennes de la famille des Ichneumonides. Bulletin de la Société Entomologique de Mulhouse, 1960 (Septembre–Octobre), 62–65. https://doi.org/10.3406/bsef.1960.20555 DOI: https://doi.org/10.3406/bsef.1960.20555
  2. Aubert, J.-F. (1961) Les Ichneumonides de Corse (Hym). Annales de la Société Entomologique de France, 130, 159–188.
  3. Aubert, J.-F. (1964) Les Ichneumonides du rivage méditerranéen français (Hym.). 7e série: Ichneumoninae, Cryptinae, Ophioninae et Mesochorinae de l’Hérault et des Bouches-du-Rhone. Bulletin de la Société Entomologique de France, 69, 144–164. https://doi.org/10.3406/bsef.1964.20764 DOI: https://doi.org/10.3406/bsef.1964.20764
  4. Aubert, J.-F. (1965) Les Ichneumonides du rivage méditerranéen français (8e série). Vie et Milieu, 16, 549–573.
  5. Aubert, J.-F. (1966a) Les Ichneumonides du rivage méditerranéen français (9e série: Ouest de l’Hérault et Aude) (Hym.). (suite et fin). Bulletin de la Société Entomologique de France, 71, 166–176. https://doi.org/10.3406/bsef.1966.20879 DOI: https://doi.org/10.3406/bsef.1966.20879
  6. Aubert, J.-F. (1966b) Fixations d’Ichneumonides lectotypes dans la collection C.G.Thomson conservée Lund. Opuscula Entomologica, 31 (1/2), 125–132.
  7. Aubert, J.-F. (1968a) Révision des travaux concernant les Ichneumonides de France et 6e supplément au catalogue de Gaulle. Bulletin de la Société Linnéenne de Lyon, 37, 133–4. https://doi.org/10.3406/linly.1968.6002 DOI: https://doi.org/10.3406/linly.1968.6002
  8. Aubert, J.-F. (1968b) Fixation des types, lectotypes et paratypes dans les collections d’ichneumonides et premiѐre liste de types perdus ou conservés. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 41, 175–201.
  9. Aubert, J.-F. (1971) Supplément aux Ichneumonides pétiolées avec neuf espèces nouvelles. Bulletin de la Société Entomologique de Mulhouse, 1971, 35–43.
  10. Aubert, J.-F. (1972) Étude commentée de nouveaux lectotypes choisis dans les collections HOLMGREN et THOMSON (Hym. Ichneumonidae). Entomologica Scandinavica, 3, 145–152. https://doi.org/10.1163/187631272X00229 DOI: https://doi.org/10.1163/187631272X00229
  11. Aubert, J.-F. (1973) Révision des travaux concernant les Ichneumonides de France et 8e supplément au Catalogue de Gaulle (100 espѐces nouvelles pour la faune française). Bulletin Mensuel de la Société Linnéenne de Lyon, 42 (2), 17–28. https://doi.org/10.3406/linly.1973.10045 DOI: https://doi.org/10.3406/linly.1973.10045
  12. Aubert, J.-F. (1974) Douze Ichneumonides pétiolées inédites. Bulletin de la Société Entomologique de Mulhouse, 1974, 1–6
  13. Aubert, J.-F. (1975) Les Ichneumonides pétiolées ouest-paléarctiques de MORLEY. Bulletin de la Société Entomologique de Mulhouse, 1975, 13–17.
  14. Aubert, J.-F. (1980) Notes sur diverse Ichneumonides mal connues ou inédites. Bulletin de la Société Entomologique de Mulhouse, 1980 (Janvier–Mars), 1–6.
  15. Aubert, J.-F. (1997) 10ème supplément au catalogue de GAULLE: 129 espèces d’Ichneumonides nouvelles pour la faune française (Hymenoptera, Ichneumonidae). Nouvelle Revue d’Entomologie, 14, 99–114.
  16. Bignell, G.C. (1883) Note on Limneria rufa, Brdg., L.brischkei, Brdg. and Rhogas reticulator Nees. Entomologist, 16, 69.
  17. Bridgman, J.B. (1881) Some additions to Mr. Marshall’s catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1881, 143–168. https://doi.org/10.1111/j.1365-2311.1873.tb01517.x DOI: https://doi.org/10.1111/j.1365-2311.1873.tb01517.x
  18. Bridgman, J.B. (1882) Further additions to the Rev. T.A.Marshall’s catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1882, 141–164. https://doi.org/10.1111/j.1365-2311.1882.tb01574.x DOI: https://doi.org/10.1111/j.1365-2311.1882.tb01574.x
  19. Bridgman, J.B. & Fitch, E.A. (1885) Introductory papers on Ichneumonidae. No. V. Ophionidae (continued). Entomologist, 18, 100–108.
  20. Bridgman, J.B. (1889) Further additions to the Rev. T.A.Marshall’s catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1889, 409–439. https://doi.org/10.1111/j.1365-2311.1889.tb02334.x DOI: https://doi.org/10.1111/j.1365-2311.1889.tb02334.x
  21. Brischke, C.G.A. (1880) Die Ichneumoniden der Provinzen West- und Ost-Preussen. Schriften der Naturforschenden Gesellschaft in Danzig, 4 (4), 108–210.
  22. Broad, G.R. (2016) Checklist of British and Irish Hymenoptera—Ichneumonidae. Biodiversity Data Journal, 4, e9042. https://doi.org/10.3897/BDJ.4.e9042 DOI: https://doi.org/10.3897/BDJ.4.e9042
  23. Broad, G.R., Shaw, M.R. & Fitton, M.G. (2018) Ichneumonid wasps (Hymenoptera, Ichneumonidae): their classification and biology. Royal Entomological Society, London, 417 pp.
  24. Carr, L.A. (1924) The Ichneumonidae of the Lichfield District, Staffordshire. Transactions of the North Staffordshire Field Club, 58 (Appendix), 1–70.
  25. Desvignes, T. (1856) Catalogue of British Ichneumonidae in the collection of the British Museum. British Museum, London, 120pp.
  26. Fitton, M.G. (1976) The western Palaearctic Ichneumonidae (Hymenoptera) of British Authors. Bulletin of the British Museum (Natural History), Entomology Series, 32, 301–373.
  27. Fitton, M.G. (1982) A catalogue and reclassification of the Ichneumonidae (Hymenoptera) described by C.G.Thomson. Bulletin of the British Museum (Natural History), Entomology Series, 45 (1), 1–119.
  28. Förster, A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 25 (1868), 135–221.
  29. di Giovanni, F. & Riedel, M. (2017) New records of Campopleginae for Italy (Hymenoptera: Ichneumonidae). Fragmenta entomologica, 49 (1), 109–114. https://doi.org/10.4081/fe.2017.239 DOI: https://doi.org/10.4081/fe.2017.239
  30. Gravenhorst, J.L.C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https://doi.org/10.5962/bhl.title.65750 DOI: https://doi.org/10.5962/bhl.title.65750
  31. Habermehl, H. (1922) Neue und wenig bekannte paläarktische Ichneumoniden (Hym.) Deutsche Entomologische Zeitschrift, 1922, 348–359. DOI: https://doi.org/10.1002/mmnd.48019220315
  32. Habermehl, H. (1923) Neue englische Ichneumoniden nebst Bemerkungen über Ichneumon macrocerus C.G. Thoms. ♀, Cratichneumon foersteri Wesm. ♀, und Grypocntrus cinctellus Ruthe ♂ (Hym.). Konowia, 2, 34–43. DOI: https://doi.org/10.1002/mmnd.48019230403
  33. Hedwig, K. (1932) Alte und neue Ichneumoniden. (Hym.). Deutsche Entomologische Zeitschrift, 1932, 24–35 DOI: https://doi.org/10.1002/mmnd.48019320103
  34. Hedwig, K. (1938) Neue schlesische Ichneumoniden (Hymenoptera). Arbeiten über Morphologische und Taxonomische Entomologie, 5, 221–227.
  35. Hedwig, K. (1940) Alte und neue Ichneumoniden. Deutsche Entomologische Zeitschrift, 1940, 190–203. DOI: https://doi.org/10.1002/mmnd.48019390107
  36. Hinz, R. (1957) Zur systematik und ökologie der Ichneumoniden 1 (Hym). Deutsche Entomologische Zeitschrift, 4, 86–90. https://doi.org/10.1002/mmnd.19570040112 DOI: https://doi.org/10.1002/mmnd.4800040112
  37. Hinz, R. (1964) Über einige Typen der HOLMGRENschen Gattung Limneria (Hym. Ichn. Ophioninae). Entomophaga, 9, 67–73. https://doi.org/10.1007/BF02375741 DOI: https://doi.org/10.1007/BF02375741
  38. Holmgren, A.E. (1860) Försök till uppställning och beskrifning af de i Sverige funna Ophionider (Monographia Ophionidum Sueciae). Kongliga Svenska Vetenskapsakademiens Handlingar, 2 (8), 1–158.
  39. Horstmann, K. (1969) Typenrevision der europäischen Arten der Gattung Diadegma Förster (syn. Angitia Holmgren). Beiträge zur Entomologie, 19, 413–472.
  40. Horstmann, K. (1972) Type revision of the species of Cryptinae and Campopleginae described by J.B.Bridgman (Hymenoptera: Ichneumonidae). Entomologist, 105, 217–228.
  41. Horstmann, K. (1978) Typenrevision der von G. Szépigleti beschriebenen paläarktischen Arten der Unterfamilie Campopleginae (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 31, 37–45.
  42. Horstmann, K. (1986) Typenrevision der von Karl Pfankuch beschriebenen Arten und Formen der Familie Ichneumonidae (Hymenoptera). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 8, 25–264.
  43. Horstmann, K. (1991) Revision der von Spinola und Rondani beschriebenen westpaläarktischen Ichneumoniden-Arten (Hymenoptera). Zeitschrift Arbeitsgemeinschaft Ősterreichischer Entomologen, 43, 43–39.
  44. Horstmann, K. (1995) Typenrevision einiger Ichneumonidae aus der Sammlung Strobl (Hymenoptera). Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen, 47 (1/2), 31–35.
  45. Horstmann, K. (1999) Revisionen von Schlupfwespen-Arten III (Hymenoptera: Ichneumonidae). Mitteilungen Münchener Entomologischen Gesellschaft, 89, 47–57.
  46. Horstmann, K. (2000) Typenrevisionen der von Gravenhorst beschriebenen oder gedeuteten Campoplex-Arten (Hymenoptera, Ichneumonidae). Linzer biologische Beiträge, 32 (2), 1203–14.
  47. Horstmann, K. (2008) Neue westpalaarktische arten der campopleginae (Hymenoptera: Ichneumonidae). Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen, 60 (1–2), 3–27.
  48. Horstmann, K. (2009) Typenrevisionen der von Kiss beschriebenen Taxa der Ichneumonidae III. Verschiedene Unterfamilien (Hymenoptera, Ichneumonidae). Linzer Biologische Beiträge, 41 (1), 673–689.
  49. Horstmann, K. (2013) Revisionen von Schlupfwespen-Arten XVII (Hymenoptera: Ichneumonidae). Mitteilungen Münchener Entomologischen Gesellschaft, 103, 1–14.
  50. Horstmann, K., Fiedler, K. & Baumgarten, H.-Th. (1997) Zur Taxonomie und Bionomie einiger Ichneumonidae (Hymenoptera) als Parasitoide westpaläarktischer Lycaenidae (Lepidoptera). Nachrichtenblatt der Bayerischen Entomologen, 46 (1/2), 2–7.
  51. Johnson, W. F. (1929) Irish Ichneumonidae. Entomologist’s Monthly Magazine, 65, 134–137.
  52. Jussila, R. (1992) Ichneumonidae (Hymenoptera) new to Finland II. Entomologica Fennica, 3 (3), 159–162. https://doi.org/10.33338/ef.83607 DOI: https://doi.org/10.33338/ef.83607
  53. Kaitaniemi, P. & Ruohomaki, K. (1999) Effects of autumn temperature and oviposition date on timing of larval development and risk of parasitism in a spring folivore. Oikos, 84 (3), 435–442. https://doi.org/10.2307/3546422 DOI: https://doi.org/10.2307/3546422
  54. Kerslake, J.E., Kruuk, L.E.B., Hartley, S.E. & Woodin, S.J. (1996) Winter moth (Operophtera brumata (Lepidoptera: Geometridae)) outbreaks on Scottish heather moorlands: effects of host plant and parasitoids on larval survival and development. Bulletin of Entomological Research, 86 (2), 155–164. https://doi.org/10.1017/S0007485300052391 DOI: https://doi.org/10.1017/S0007485300052391
  55. Kiss, von Zilah A. (1924) Beitrag zur Kenntnis der ungarischen und siebenbürgischen Ichneumoniden-(Schlupfwespen-) Fauna. Verhandlungen und Mitteilungen des Siebenbürgischen Vereins für Naturwissenschaften in Hermannstadt, 72/74, 32–146.
  56. Kiss, von Zilah A. (1926) Zweiter Beitrag zur Kenntnis der ungarischen und siebenbürgischen Ichneumoniden-
  57. (Schlupfwespen-) Fauna. Verhandlungen und Mitteilungen des Siebenbürgischen Vereins für Naturwissenschaften in Hermannstadt, 75/76, 74–120.
  58. Klopfstein, S. & Baur, H. (2011) Catalogue of the type specimens of Ichneumonidae (Hymenoptera) in the Jacques F. Aubert collection at the Musée de Zoologie, Lausanne, Switzerland. Zootaxa, 3081 (1), 1–90. https://doi.org/10.11646/zootaxa.3081.1.1 DOI: https://doi.org/10.11646/zootaxa.3081.1.1
  59. Klopfstein, S., Broad, G.R., Urfer, K., Vårdal, H. & Haraldseide, H. (2022) An interactive key to the European genera of Campopleginae (Hymenoptera, Ichneumonidae) and 20 new species for Sweden. Entomologisk tidskrift, 143 (2022), 121–156.
  60. Kolomiets, N.G. (1962) [Parasites and predators of Dendrolimus sibiricus Tschet]. Izdatel’stvo Sibirskogo Otdeleniya An SSSR, Novosibirsk, 173 pp. [in Russian]
  61. Koponen, M., Jussila, R. & Vikberg, V. (2003) A checklist of Finnish Hymenoptera Parasitica, part 5. Family Ichneumonidae, subfamilies Tersilochinae, Ophioninae, Anomaloninae, Paxylommatinae, Cremastinae and Campopleginae (Hymenoptera, Parasitica). Sahlbergia, 8 (1), 27–48.
  62. Kukal, O. & Kevan, P.G. (1987) The influence of parasitism on the life history of a high arctic insect, Gynaephora groenlandica (Wocke) (Lepidoptera: Lymantriidae). Canadian Journal of Zoology, 65 (1), 156–163. https://doi.org/10.1139/z87-022 DOI: https://doi.org/10.1139/z87-022
  63. Meyer, N.F. (1926) Einige neue Ichneumoniden und Cynipiden. Revue Russe d’Entomologie, 20, 260–264.
  64. Morley, C. (1913) The fauna of British India including Ceylon and Burma, Hymenoptera, Vol. 3. Ichneumonidae. British Museum, London, 531 pp.
  65. Morley, C. (1915) Ichneumonologia Britannica. V. The Ichneumons of Great Britain. Ophioninae. H. & W. Brown, London, 400pp.
  66. Nouhuys, S. van & Ehrnsten, J. (2004) Wasp behaviour leads to uniform parasitism of a host available only a few hours per year. Behavioural Ecology, 15, 661–665. https://doi.org/10.1093/beheco/arh059 DOI: https://doi.org/10.1093/beheco/arh059
  67. Nouhuys, S. van & Kaartinen, R. (2008) A parasitoid wasp uses landmarks while monitoring potential resources. Proceedings of the Royal Society (B), 275, 377–385. https://doi.org/10.1098/rspb.2007.1446 DOI: https://doi.org/10.1098/rspb.2007.1446
  68. Oehlke, J. (1963) Revision der im Deutschen Entomologischen Institut aufbewahrten Typen paläarktischer Ichneumoniden (Hymenoptera: Ichneumonidae). Beiträge zur Entomologie, 13, 403–410.
  69. Oehlke, J. & Townes, H.K. (1969) Schmideknechts Ichneumonidentypen aus der Kollektion des Museums Rudolstadt. Beiträge sur Entomologie, 19, 395–412.
  70. Perkins, J.F. (1953) Notes on the British Ichneumoninae with descriptions of new species (Hym., Ichneumonidae). Bulletin of the British Museum (Natural History), Entomology, 3 (4), 9–176. https://doi.org/10.5962/bhl.part.1050 DOI: https://doi.org/10.5962/bhl.part.1050
  71. Perkins, J.F. (1962) On the type species of Förster’s genera (Hymenoptera: Ichneumonidae). Bulletin of the British Museum (Natural History), 11, 385–483.
  72. Pfankuch, K. (1921) Neue Ichneumoniden. Entomologisches Jahrbuch, 30, 149–159.
  73. Plaas, G. (1932) Der Stachelbeerspanner Abraxas grossulariata L. in Schleswig-Holstein. Zeitschrift für Wissenschaftliche Insektenbiologie, 26 & 27, 183–191 & 12–17.
  74. Quicke, D.J.L., Laurenne, N.M., Fitton, M.G. & Broad, G.R. (2009) A thousand and one wasps: a 28S rDNA and morphological phylogeny of the Ichneumonidae (Insecta, Hymenoptera) with an investigation into alignment parameter space and elision. Journal of Natural History, 43, 1305–1421. https://doi.org/10.1080/00222930902807783 DOI: https://doi.org/10.1080/00222930902807783
  75. Ratnasingham, S. & Hebert, P.D.N. (2007) BOLD: The Barcode of Life Data System (www.barcodinglife. org). Molecular Ecology Notes, 7, 355–364. https://doi.org/10.1111/j.1471-8286.2007.01678.x DOI: https://doi.org/10.1111/j.1471-8286.2007.01678.x
  76. Ratnasingham, S. & Hebert, P.D.N. (2013) A DNA Based Registry for All Animal Species: The Barcode Index Number (BIN) System. PLoS ONE, 8 (7), e66213. https://doi.org/10.1371/journal.pone.0066213 DOI: https://doi.org/10.1371/journal.pone.0066213
  77. Ratzeburg, J.T.C. (1844) Die Ichneumonen der Forstinsecten in forstlicher und entomologischer Beziehung. Nicolaischen Buchhandlung, Berlin, 224pp. https://doi.org/10.5962/bhl.title.11094 DOI: https://doi.org/10.5962/bhl.title.11094
  78. Roman, A. (1912) Die Ichneumonidentypen C.P.Thunbergs. Zoologiska Bidrag fran Uppsala, 1, 229–293.
  79. Rondani, C. (1877) Vesparia Parasita non vel minus cognita. Bollettino della Societa Entomologica Italiana, 9, 166–206.
  80. Rudow, F. (1917) Ichneumoniden und ihre Wirte. Entomologische Zeitschrift, 31, 71–72.
  81. Schmiedeknecht, O. (1909) Opuscula Ichneumonologica. IV. Band. Fasc. XXI–XXIII. Ophioninae. s.n., Blankenburg in Thüringen. [unknown pagination]
  82. Shaw, M.R. (2003) Adverse comment on the supposed British status of the web-spinning sawfly Cephalcia arvensis Panzer (Hymenoptera: Pamphiliidae). British Journal of Entomology and Natural History, 16 (1), 2.
  83. Shaw, M.R. (2012) Larval parasitoids of Rivula sericealis (Scopoli) (Lepidoptera: Noctuidae) in Britain, including notes on the biology of Cotesia subordinaria (Tobias) (Hymenoptera: Braconidae, Microgastrinae), a solitary-cum-gregarious parasitoid. Entomologist’s Gazette, 63, 251–257.
  84. Shaw, M.R., Horstmann, K. & Whiffin, A.L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist’s Gazette, 67, 177–222.
  85. Shaw, M.R., Stefanescu, C. & van Nouhuys, S. (2009) Parasitoids of European butterflies. In: Settele, J., Shreeve, T., Konvicka, M. & van Dyck, H. (Eds.), Ecology of Butterflies in Europe. Cambridge University Press, Cambridge, pp. 130–156.
  86. Smits van Burgst, C.A.L. (1913) Tunesien Hymenoptera. Privatdruk, den Haag, 37 pp.
  87. Smits van Burgst, C.A.L. (1914) Ichneumonidae captured in the environs of Bozen (Tyrol) in June 1913. Deutsche Entomologische Zeitschrift, 1914, 325–333. DOI: https://doi.org/10.1002/mmnd.48019140312
  88. Strobl, G. (1904) Ichneumoniden Steiermarks (und der Nachbarländer), V. Fam. Ophionidae. Mitteilungen Naturwissenschaftlichen Vereines für Steiermark, Graz, 40, 43–160. [1903]
  89. Szepligeti, G. (1916) Ichneumoniden aus der Sammlung des ungarischen National Museums II. Annales Musei Nationalis Hungarici, 14, 225–380.
  90. Thomson, C.G. (1887) XXXV. Försök till uppställning och beskrifning af aterna inom slägtet Campoplex (Grav.) Opuscula Entomologica, Lund, XI, 1043–1182.
  91. Thomson, C.G. (1890) XLIII Försök till uppställning och beskrifning af aterna inom slägtet Campoplex (Grav.) Opuscula Entomologica, Lund, XI, 1043–1182.
  92. Thunberg, C.P. (1824) Ichneumonidea, Insecta Hymenoptera illustrata. Mémoires de l’Académie Imperiale des Sciences de Saint Petersbourg, 9, 285–368.
  93. Townes, H.K. (1970) The genera of Ichneumonidae, Part 3. Memoirs of the American Entomological Institute, 13, 1–307.
  94. Townes, H.K., Momoi, S. & Townes, M. (1965) A catalogue and reclassification of the eastern Palearctic Ichneumonidae. Memoirs of the American Entomological Institute, 5, 1–661.
  95. Tschek, C. (1871) Ichneumonologische Fragmente. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 21, 37–68.
  96. Viereck, H.L. (1910) Descriptions of new species of Ichneumon flies. Proceedings of the United States National Museum, 38 (1754), 379–384. https://doi.org/10.5479/si.00963801.38-1754.379 DOI: https://doi.org/10.5479/si.00963801.38-1754.379
  97. Viereck, H.L. (1911) Descriptions of one new genus and eight new species of Ichneumon flies. Proceedings of the United States National Museum, 40 (1832), 475–480. https://doi.org/10.5479/si.00963801.1832.475 DOI: https://doi.org/10.5479/si.00963801.1832.475
  98. Viereck, H.L. (1914) Type species of the genera of Ichneumon flies. United States National Museum Bulletin, 83, 1–186. https://doi.org/10.5479/si.03629236.83.1 DOI: https://doi.org/10.5479/si.03629236.83.1
  99. Wahl, D.B. (1991) The status of Rhimphoctona, with special reference to the higher categories within the Campopleginae and the relationships of the subfamily (Hymenoptera: Ichneumonidae). Transactions of the American Entomological Society, 117,193–213.
  100. Yu, D.S.K., van Achterberg, C. & Horstmann, K. (2016) Taxapad 2016. Ichneumonoidea 2015. Database on flash-drive. Nepean, Ontario. http://www.taxapad.com (accessed 9 April 2023)