Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5353 No. 6: 13 Oct. 2023
Type: Article
Published: 2023-10-13
DOI: 10.11646/zootaxa.5353.6.1
Page range: 501-532
Abstract views: 215
PDF downloaded: 22

Checklist of the subfamily Scolytinae (Coleoptera: Curculionidae) in Tucumán province, Argentina

Instituto de Entomología; Área de Zoología; Fundación Miguel Lillo; Miguel Lillo 251 (4000); San Miguel de Tucumán; Argentina
University of Texas Insect Collection; Lake Austin Center 3001 Lake Austin Boulevard; Suite 1.314; Austin; Texas 78703
Instituto de Ecología; Comportamiento y Conservación; Biología Integrativa; Fundación Miguel Lillo; Miguel Lillo 251 (4000); San Miguel de Tucumán; Argentina
Coleoptera Biodiversity inventory Argentine Northwest species

Abstract

A checklist of Scolytinae species is presented to contribute to the knowledge of the biodiversity of the subfamily in Tucumán province, Argentina. The inventory was based on collections in the field from 2016 to 2023, and a review of the material deposited in the entomological collections of the Museo de La Plata, the Museo Argentino de Ciencias Naturales Bernardino Rivadavia, and the Fundación Miguel Lillo. Previous bibliographic records were also included. In total, 54 species belonging to 11 tribes were recorded, of which six were new reports for Argentina and nine for Tucumán province; eight species were introduced. The genus Amasa was also recorded in Argentina for the first time.

 

References

  1. Adámolli, J., Sennhauser, E., Acero, J.M. & Rescia, A. (1990) Stress and disturbance: vegetation dynamics in the dry Chaco region of Argentina. Journal of Biogeography, 17, 491–500. https://doi.org/10.2307/2845381
  2. Atkinson, T.H. (2017) Curculionidae: Scolytinae: Escarabajos descortezadores y escarabajos ambrosiales. In: Cibrián-Tovar, D. (Ed.), Fundamentos de Entomología Forestal. CONACYT, México, D.F., pp. 306–313.
  3. Atkinson, T.H. (2019) Escarabajos descortezadores y ambrosiales (Coleoptera: Curculionidae: Scolytinae, Platypodinae) de Sonora, México. Dugesiana, 26 (1), 41–49. https://doi.org/10.32870/dugesiana.v26i1.7068
  4. Atkinson, T.H. (2023) Bark and ambrosia beetles of the Americas. Available from: http://www.barkbeetles.info (accessed 29 August 2023)
  5. Atkinson, T.H. & Equihua, A. (1986) Biology of Scolytidae and Platypodidae (Coleoptera) of a tropical rain forest in southeastern Mexico with an annotated checklist of species. Annals of the Entomological Society of America, 79, 414–423. https://doi.org/10.1093/aesa/79.3.414
  6. Atkinson, T.H. & Flechtmann, C.A.H. (2021) New species, new records and synonymy of Brazilian species of Hypothenemus Westwood, 1834 (Coleoptera: Curculionidae: Scolytinae). Insecta Mundi, 0846, 1–33. [https://journals.flvc.org/mundi/article/view/127953/128638]
  7. Atkinson, T.H., Martínez F.E., Saucedo, C.E. & Burgos, S.A. (1986a) Scolytidae y Platypodidae (Coleoptera) asociados a selva baja caducifolia y comunidades derivadas en el estado de Morelos. Folia Entomológica Mexicana, 69, 41–82. https://doi.org/10.21829/azm.1986.13172048
  8. Atkinson, T.H., Saucedo C.E., Martínez F.E. & Burgos S.A. (1986b) Coleópteros Scolytidae y Platypodidae asociados con las comunidades vegetales de clima templado y frío en el estado de Morelos. Acta Zoológica Mexicana, 17, 1–58. https://doi.org/10.21829/azm.1986.13172048
  9. Ayarde, H.R., Boero, C., Moris, M., Slanis, A. & Gonzáles, J.A. (1999) Relevamiento de la Biodiversidad de la Reserva Nacional de Flora y Fauna Tariquía (Tarija, Bolivia). In: González, J.A., Scrocchi, G.J. & Lavilla, E.O. (Eds.), Serie Conservación de la Naturaleza N° 14. Fundación Miguel Lillo-PROBONA, Tucumán, pp. 29–62. https://doi.org/10.30550/j.azl/2017.61.2/1
  10. Bachmann, A.O. & Lanteri, A.A. (2013) Catalog of the types of Curculionoidea (Insecta, Coleoptera) deposited at the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires. Revista del Museo Argentino de Ciencias Naturales, Nueva Serie, 15 (2), 209–280. [http://revista.macn.gob.ar/ojs/index.php/RevMus/article/view/183/174]
  11. Barnouin, T., Soldati, F., Roques, L., Faccoli, M., Kirkendall, L.R., Mouttet, R., Daubree, J.B. & Noblecourt, T. (2020) Bark beetles and pinhole borers recently or newly introduced to France (Coleoptera: Curculionidae, Scolytinae and Platypodinae). Zootaxa, 4877 (1), 51–74. https://doi.org/10.11646/zootaxa.4877.1.2
  12. Blackman, M.W. (1939) A new genus and three new species of Scolytidae from Argentina and Bolivia (Coleoptera). Revista de Entomologia, São Paulo, 10 (1), 86–96.
  13. Blackman, M.W. (1943) New species of American scolytid beetles, mostly neotropical. Proceedings of the United States National Museum, 94 (3174), 371–399. https://doi.org/10.5479/si.00963801.94-3174.371
  14. Blandford, W.F.H. (1896) Scolytidae. Biologia Centrali-Americana, Coleoptera, 4 (6), 97–144.
  15. Blandford, W.F.H. (1898) Scolytidae. Biologia Centrali-Americana, Coleoptera, 4 (6), 185–224.
  16. Bobba, M.E. & Hernández, C.M. (2005) Aspectos Geográficos del Noroeste Argentino. In: Minetti, J.L. (Ed.), El Clima del Noroeste Argentino. s.n., Tucumán, pp. 9–25. https://doi.org/10.35537/10915/55243
  17. Brèthes, J. (1921) Notas Coleopterologicas. Revista de la Facultad de Agronomia, Universidad de la Plata, 14, 163–169.
  18. Bright, D.E. (2004) Curculionidae: Scolytinae. In: Cordo, H.A., Logarzo, G., Braun, K., Di Iorio, O.R. (Eds.), Catálogo de insectos fitófagos de la Argentina y sus plantas asociadas. Sociedad Entomológica Argentina Ediciones, Buenos Aires, pp. 155–162. https://doi.org/10.22179/revmacn.11.272
  19. Bright, D.E. (2019) A taxonomic monograph of the bark and ambrosia beetles of the West Indies (Coleoptera: Curculionoidea: Scolytidae). Occasional Papers of the Florida State Collection of Arthropods, 12, 1–491.
  20. Brown, A.D. (1995) Fitogeografía y conservación de las selvas de montaña del Noroeste de Argentina. In: Churchill, S.P., Balslev, H., Forero, E. & Luteyn, J.L. (Eds.), Biodiversity and Conservation of Neotropical Montane Forest. The New York Botanical Garden, New York, New York, pp. 663–672. https://doi.org/10.1017/s0266467400010798
  21. Brown, A.D. & Pacheco, S. (2005) Propuesta de actualización del mapa ecorregional de la Argentina. In: Brown, A., Martinez Ortiz, U., Acerbi, M. & Corcuera, J. (Eds.), La Situación Ambiental Argentina 2005. s.n., Buenos Aires, pp. 28–31.
  22. Bruch, C. (1914) Catálogo sistemático de los coleópteros de la República Argentina, Pars VII. Revista del Museo de La Plata, 19 (6), 401–441. https://doi.org/10.5962/bhl.title.8799
  23. Bucher, E.H. (1987) Herbivory and semi-arid regions of Argentina. Revista Chilena de Historia Natural, 60, 265–273.
  24. Burgos, A. & Equihua-Martínez, A. (2007) Platypodidae y Scolytidae (Coleoptera) de Jalisco, México. Dugesiana, 14 (2), 59–82.
  25. Cabrera, A.L. (1976) Regiones fitogeográficas argentinas. In: Kugler, W.F. (Ed.), Enciclopedia Argentina de Agricultura y Jardinería. 2º Edición. s.n., Buenos Aires, pp. 1–85.
  26. Córdoba, S.P. & Atkinson, T.H. (2018) New records of Scolytinae (Coleoptera: Curculionidae) from Argentina. The Coleopterists Bulletin, 72 (2), 376–379. https://doi.org/10.1649/0010-065x-72.2.376
  27. Córdoba, S.P., Mendoza, E.A., Atkinson, T.H. & Manzo, M.V. (2021) Diversidad y dinámica poblacional de Scolytinae (Coleoptera: Curculionidae) en plantaciones de palto y bosque natural circundante en el NW de Argentina. Dugesiana, 28 (2), 115–130. https://doi.org/10.32870/dugesiana.v28i2.7151
  28. Del Río, M.G., Lanteri, A.A. & Suárez, S.M. (2005) Types of Scolytidae and Platypodidae (Coleoptera: Curculionoidea) housed at the Museo de La Plata entomological collection. Revista del Museo de La Plata, 46, 1–11. [https://publicaciones.fcnym.unlp.edu.ar/rmlp/article/view/2174]
  29. Dos-Santos, C.A.A., Trevisan, H., De-Carvalho, A.G., Plata-Rueda, A., Zanuncio, J.C. & De-Souza, T.S. (2022) Increased capture efficiency of Scolytinae with modified semi-funnel trap model. Brazilian Journal of Biology, 84 (e259131), 1–9. https://doi.org/10.1590/1519-6984.259131
  30. Eggers, H. (1908) Funf neue Borkenkäfer. Entomologische Blätter, 4, 214–217.
  31. Eggers, H. (1928) Ipidae (Coleoptera) da America do Sul. Archivos do Instituto de Biologia Vegetal, Rio de Janeiro, 1, 83–99.
  32. Eggers, H. (1933) Borkenkafer (Ipidae, Col.) aus Sudamerika, VI. Material des Museum Pais aus Franz, Guayana und Venezuela. Travaux du Laboratoire d’Entomologie du Museum National d’Histoire Naturelle, 1, 1–37.
  33. Eggers, H. (1934) Borkenkäfer (Ipidae, Col.) aus Südamerika, VII. Entomologische Blätter, 30 (2), 78–84.
  34. Eggers, H. (1943) Borkenkäfer (Col., Ipidae) aus Südamerika, X. Bolivia. Mitteilungen der Münchener Entomologischen Gesellschaft, 33, 344–389
  35. Eichhoff, W.J. (1869) Neue Borkenkäfer. Berliner Entomologische Zeitschrift, 12 (3–4), 273–282.
  36. Eichhoff, W.J. (1868) Neue amerikanische Borkenkäfer-Gattungen und Arten. Berliner Entomologische Zeitschrift, 12, 145–152.
  37. Eichhoff, W.J. (1872) Neue exotische Tomiciden-Arten. Berliner Entomologische Zeitschrift, 15 (2–3), 131–136. https://doi.org/10.1002/mmnd.18710150203
  38. Equihua-Martínez, A. & Atkinson, T.H. (1986) Annotated checklist of the bark and ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) associated with tropical deciduous forest in Chamela, Jalisco, Mexico. Florida Entomologist, 69, 619–635. https://doi.org/10.2307/3495205
  39. Equihua, A. & Burgos, A. (2002) Scolytidae. In: Llorente Bousquets, J. & Morrone, J.J. (Eds.), Biodiversidad, Taxonomía y Biogeografía de Artrópodos de México: Hacia una síntesis de su conocimiento. Vol. III . s.n. México, D.F., pp. 539–557. https://doi.org/10.21829/azm.2014.303100
  40. Erichson, W.F. (1842) Beitrag zur Insecten-Fauna von Vandiemensland, mit besonderer Berücksichtigung der geografischen Verbreitung der Insekten. Archiv für Naturgeschichte, 8 (1), 83–287. https://doi.org/10.5962/bhl.part.21657
  41. Estrada, A. & Atkinson, T.H. (1989) Scolytidae y Platypodidae de Escarcega, Campeche, México. Biogeografía, biología, importancia económica y una lista comentada de especies. Anales del Instituto de Bioliología, Serie Zoología, 58, 199–220.
  42. Fabricius, J.C. (1792) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tomus I. Pars II. Impensis Christ. Gottl. Proft, Hafniae, 538 pp. https://doi.org/10.5962/bhl.title.122153
  43. Fabricius, J.C. (1801) Systema eleutheratorum secundum ordines, genera, species: adiectis synonimis, locis, observationibus, descriptionibus. Tomus II. Impensis Bibliopolii Academici Novi, Kiliae, 687 pp. https://doi.org/10.5962/bhl.title.137098
  44. Ferrari, J.A. (1867) Die Forst- und Baumzuchtschädlichen Borkenkäfer (Tomicides Lac.) aus der Familie der Holzverderber (Scolytides Lac.), mit besonderer Berücksichtigung vorzüglich der Europäischen Formen, und der Sammlung des K. K. Zoologischen Kabinets in Wien. Carl Gerold‘s Sohn, Wien, pp. 96. https://doi.org/10.1002/mmnd.18680120209
  45. Fisher, R.C., Thompson, G.H. & Webb, W.E. (1953) Ambrosia beetles in forest and sawmill, their biology, economic importance and control. Forestry Abstracts, 14, 381– 389.
  46. Flechtmann, C.A.H. & Cognato, A.I. (2011) First report of Amasa truncata (Erichson (Coleoptera: Curculionidae: Scolytinae) in Brazil. The Coleopterists Bulletin, 65 (4), 417–421. https://doi.org/10.1649/072.065.0419
  47. Flechtmann, C.A.H., Ottati, A.L.T. & Berisford, C. (2001) Ambrosia and bark beetles (Scolytidae: Coleoptera) in pine and eucalypt stands in Southern Brazil. Forest Ecology and Management, 142, 183–191. https://doi.org/10.1016/s0378-1127(00)00349-2
  48. Gómez, D., Suárez, M. & Martínez, G. (2017) Amasa truncata (Erichson) (Coleoptera: Curculionidae: Scolytidae): A new exotic ambrosia beetle in Uruguay. The Coleopterists Bulletin, 71, 825–826. https://doi.org/10.1649/0010-065X-71.4.825
  49. Hagedorn, J.M. (1909) Diagnosen bisher unbeschriebener Borkenkäfer. Zweite Serie, erste Hälfte. Deutsche Entomologische Zeitschrift, 1909 (6), 733–746.
  50. Hagedorn, J.M. (1912) Borkenkäfer (Ipidae) welche in Kautschukbäumen leben. Revue Zoologique Africaine, 1, 336–346.
  51. Hayward, K.J. (1960) Insectos tucumanos perjudiciales. Revista Industrial y Agrícola de Tucumán, 42 (1), 3–144.
  52. Hopkins, A.D. (1915) Classification of the Cryphalinae, with descriptions of new genera and species. United States Department of Agriculture Report No. 99. Government Printing Office, Washington, 75 pp. https://doi.org/10.5962/bhl.title.65905
  53. Hornung, E.G. (1842) Ueber einige in den Betelnüssen vorkommende Käfer. Entomologische Zeitung, Stettin, 3, 115–117.
  54. Hulcr, J., Beaver, R.A., Puranasakal, W., Dole, S.A. & Sonthichai, S. (2008) A comparison of bark and ambrosia beetle communities in two forest types in northern Thailand (Coleoptera: Curculionidae: Scolytinae and Platypodinae). Environmental Entomology, 37 (6), 1461–1470. https://doi.org/10.1603/0046-225X-37.6.1461
  55. Hueck, K. (1978) Los bosques de Sudamérica. GTZ, Eschborn, 476 pp.
  56. Iturre, M. & Darchuck, E. (1996) Registro de escolítidos asociados al género Eucaliptus en Santiago del Estero. Quebracho, 4, 11–16.
  57. Jordal, B.H. & Smith, S.M. (2020) Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences. Zootaxa, 4813 (1), 1–67. https://doi.org/10.11646/zootaxa.4813.1.1
  58. Kirkendall, L.R. (2018) Invasive bark beetles (Coleoptera, Curculionidae, Scolytinae) in Chile and Argentina, including two species new for South America, and the correct identity of the Orthotomicus species in Chile and Argentina. Diversity, 10 (2), 1–20. https://doi.org/10.3390/d10020040
  59. Kirkendall, L.R. & Jordal, B.H. (2006) The bark and ambrosia beetles (Curculionidae, Scolytinae) of Cocos Island, Costa Rica and the role of mating systems in island zoogeography. Biological Journal of the Linnean Society, 89, 729–743. https://doi.org/10.1111/j.1095-8312.2006.00698.x
  60. Kirsch, T.F.W. (1866) Beiträge zur Käferfauna von Bogotà. Zweites Stück. Berliner Entomologische Zeitschrift, 10 (1–3), 173–216. https://doi.org/10.1002/mmnd.18660100113
  61. Knížek, M. (2011) Platypodinae, Scolytinae. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera. Vol. 7. Apollo Books, Stenstrup, pp. 201–250.
  62. Kurtz, F. (1904) Flora. In: Río, M. & Achával, L. (Eds.), Geografía de la Provincia de Córdoba. Vol. 1. Compañía Sudamericana de Billetes de Banco, Buenos Aires, pp. 270–343. https://doi.org/10.5962/bhl.title.9303
  63. Landi, L., Gómez, D., Braccini, C.L., Pereyra, V.A., Smith, S.M. & Marvaldi, A.E. (2017) Morphological and molecular identification of the invasive Xylosandrus crassiusculus (Coleoptera: Curculionidae: Scolytinae) and its South American range extending into Argentina and Uruguay. Annals of the Entomological Society of America, 110 (3), 344–349. https://doi.org/10.1093/aesa/sax032
  64. Mannerheim, C.G. von (1853) Beitrag zur Käfer-fauna der Aleutischen Inseln, der Insel Sitkha und Neu-Californiens. Bulletin de la Société Impériale des Naturalistes de Moscou, 16 (2), 175–314.
  65. Martínez, M., Cognato, A.I., Guachambala, M. & Boivin, T. (2019) Bark and Ambrosia Beetle (Coleoptera: Curculionidae: Scolytinae) diversity in natural and plantation forests in Ecuador. Environmental Entomology, 48 (3), 603–613. https://doi.org/10.1093/ee/nvz037
  66. Mendoza, E.A. (2005) El clima y la vegetación natural. In: Minetti, J.L. (Ed.), El clima del Noroeste Argentino. Magna Press, Tucumán, pp. 267–320. https://doi.org/10.3989/pirineos.2008.v163.21
  67. Mendoza, E.A. & González, J. (2011) Las ecoregiones del Norte argentino basadas en la clasificación climática de Kóppen. Serie de Conservación de la Naturaleza, 17, 1–44. https://doi.org/10.11144/javeriana.10554.59463
  68. Morello, J. & Saravia-Toledo, C. (1959) El bosque Chaqueño. I. Paisaje primitivo, paisaje cultural en el Oriente de Salta. Revista Agronómica del Noroeste Argentino, 3, 5–12.
  69. Motschulsky, V. (1866) Essai d’un catalogue des insectes de l’île de Ceylan. Supplément. Bulletin de la Société Impériale des Naturalistes de Moscou, 39 (2), 393–446.
  70. Müller, P.J.W. (1818) Vermischte Bemerkungen über einige Käferarten. Magazin de Entomologie Germar, Halle, 3, 244–249.
  71. Nel, W.J., De Beer, Z.W., Wingfield, M.J. & Duong, T.A. (2020) The granulate ambrosia beetle, Xylosandrus crassiusculus (Coleoptera: Curculionidae, Scolytinae), and its fungal symbiont found in South Africa. Zootaxa, 4838 (3), 427–435. https://doi.org/10.11646/zootaxa.4838.3.7
  72. Noguera, F.A. & Atkinson, T.H. (1990) Biogeography and biology of bark and ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) in a mesic montane forest in Mexico with an annotated checklist of species. Annals of the Entomological Society of America, 83 (3), 453–466. https://doi.org/10.1093/aesa/83.3.453
  73. Olson, D.M., Dinerstein, E., Wikramanayake, E.D., Burgess, N.D., Powell, G.V.N., Underwood, E.C., D’Amico, J.A., Itoua, I., Strand, H.E., Morrison, J.C., Loucks, C.J., Allnutt, T.H., Ricketts, T.H., Kura, Y., Lamoreux, J.F., Wettengel, W.W., Hedao, P. & Kassem, K.R. (2001) Terrestrial ecoregions of the world: A new map of life on earth: A new global map of terrestrial ecoregions provides an innovative tool for conserving biodiversity. BioScience, 51 (11), 933–938. https://doi.org/10.1641/0006-3568(2001)051%5B0933:teotwa%5D2.0.co;2
  74. Panzer, G.W.F. (1791) Beschreibung eines noch unbekannten sehr kleinen Kapuzkäfers. Naturforscher, 25, 35–38.
  75. Parodi, L.R. (1942) Viaje a la Provincia de Catamarca. Revista del Museo de La Plata, Nueva Serie, Sección Ofisial, 64–77.
  76. Ragonese, A.E. (1967) Vegetación y Ganadería en la República Argentina. Colección Científica del INTA, Buenos Aires, 213 pp.
  77. Ratzeburg, J.T.C. (1837) Die Forst-insekten: oder Abbildung und Beschreibung der in den Wäldern Preussens und der Nachbarstaaten als schädlich oder nützlich bekannt gewordenen Insekten. Erster Theil. Die Käfer. Nicolai, Berlin, 202 pp.
  78. Rudinsky, J.A. (1962) Ecology of Scolytidae. Annual Review of Entomology, 7, 327–348. https://doi.org/10.1146/annurev.en.07.010162.001551
  79. Santoro, F.H. (1966) Panorama entomológico relacionado con la Silvicultura y la tecnología forestal en la República Argentina. Revista Forestal Argentina, 10 (2), 53–59.
  80. Schedl, K.E. (1937a) Scolytidae und Platypodidae-Zentral und sudamerikanische Arten. Archivos do Instituto de Biologia Vegetal, Rio de Janeiro, 3 (2), 155–170. https://doi.org/10.5962/bhl.title.115964
  81. Schedl, K.E. (1937b) Scolytidae und Platypodidae (Coleoptera). 44 Beitrag. Arbeiten über Morphologische und Taxonomische Entomologie aus Berlin-Dahlem, 4 (1), 66–70.
  82. Schedl, K.E. (1938) Scolytidae und Platypodidae. 53 Beitrag. Diagnosen neuer und Fundort bereits bekannter argentinischer Arten. Revista de la Sociedad Entomológica Argentina, 10 (1), 21–28.
  83. Schedl, K.E. (1949) Neotropical Scolytoidea, I. 97th Contribution to the morphology and taxonomy of the Scolytoidea (Col.). Revista Brasileira de Biologia, 9 (3), 261–284.
  84. Schedl, K.E. (1950) Neotropical Scolytoidea, II. 107 Contribution. Dusenia, 1 (3), 145–180.
  85. Schedl, K.E. (1958) Fauna Argentinensis, VII. 136. Beitrag zur Morphologie und Systematik der Scolytoidea (Coleoptera). Acta Zoologica Lilloana, 16, 33–46. https://doi.org/10.1080/01650527509360480
  86. Schedl, K.E. (1966) Neotropische Scolytoidea, VIII. 238 Beitrag. Entomologische Arbeiten aus dem Museum Georg Frey, 17, 74–128.
  87. Smith, S.M. & Cognato, A.I. (2014) A taxonomic monograph of Nearctic Scolytus Geoffroy (Coleoptera, Curculionidae, Scolytinae). ZooKeys, 450, 1–182. https://doi.org/10.3897/zookeys.450.7452
  88. Smith, S.M. & Cognato, A.I. (2021) A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini). ZooKeys, 1044, 609–720. https://doi.org/10.3897/zookeys.144.62246
  89. Storer, C.G., Breinholt, J.W. & Hulcr, J. (2015) Wallacellus is Euwallacea: molecular phylogenetics settles generic relationships (Coleoptera: Curculionidae: Scolytinae: Xyleborini). Zootaxa, 3974 (3), 391–400. https://doi.org/10.11646/zootaxa.3974.3.6
  90. Vervoorst, F. (1982) Noroeste. In: Vervoorst (Ed.), Conservación de la vegetación natural en la República Argentina. Simposio XVIII Jornadas Argentinas de Botánica, Fundación Miguel Lillo, Serie de Conservación de la Naturaleza. s.n., Tucumán, pp. 9–24.
  91. Viana, M.J. (1965) Datos ecológicos de Scolytidae argentinos (Coleoptera). Revista de la Sociedad Entomológica Argentina, 27 (1–4), 119–130. [https://www.biotaxa.org/RSEA/article/view/43077]
  92. Westwood, J.O. (1834) Description of a minute coleopterous insect, forming the type of a new subgenus allied to Tomicus, with some observations upon the affinities of the xylophaga. Transactions of the Entomological Society of London, 1, 34–36. https://doi.org/10.1111/j.1365-2311.1837.tb03174.x
  93. Wichmann, H.E. (1914) Zur Kenntnis der Ipiden, II. Entomologische Blätter, 10, 136–139.
  94. Wood, S.L. (2007) Bark and ambrosia beetles of South America (Coleoptera, Scolytidae). Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, 900 pp. https://doi.org/10.1134/s001387380902016x
  95. Wood, S.L. & Bright, D.E. (1992) A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: taxonomic index. Great Basin Naturalist Memoirs, 13, 1–1553. [https://scholarsarchive.byu.edu/gbnm/vol13/iss1/21/]