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Published: 2023-10-17
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Freshwater fishes of Iraq: a revised and updated annotated checklist—2023

Department of Biology; Faculty of Art and Sciences; Nevşehir Hacı BektaşVeli University; Nevşehir; Türkiye
School of Environmental and Animal Sciences; Unitec Institute of Technology; Auckland; New Zealand
Department of Fisheries; Faculty of Natural Resources; University of Tehran; Karaj; Iran
Ichthyology and Molecular Systematics Research Laboratory; Zoology Section; Department of Biology; School of Science; Shiraz University; Shiraz; Iran
Department of Fisheries; Faculty of Natural Resources; University of Tehran; Karaj; Iran
Health Services Vocational School; Nevşehir Hacı BektaşVeli University; Nevşehir; Türkiye
Staatliches Museum für Naturkunde in Stuttgart; Rosenstein 1; 70191 Stuttgart; Germany
Pisces Freshwater ichthyofauna diversity endemic IUCn exotic Euphrates tigris

Abstract

The current status of the freshwater fishes of Iraq is revised, and an updated checklist is presented. The confirmed freshwater fishes of Iraq comprise 98 species belonging to 16 orders, 28 families, and 56 genera. Among these, 21 species (21.4%) are alien, and three species (3.0%) are considered endemic to Iraq. The orders with the largest numbers of species in the ichthyofauna of Iraq are Cypriniformes (57 species), followed by Siluriformes (12 species), Mugiliformes and Cyprinodontiformes (six species in each), Acanthuriformes and Cichliformes (three species in each), Centrarchiformes and Gobiiformes (two species in each), and the others represent only one species. At the family level, the Cyprinidae have the greatest number of species (28; 28.3% of the total species), followed by Nemacheilidae (16 species), Leuciscidae (eight species), Mugilidae (six species), Sisoridae (five species), and Xenocyprididae (four species in each). A total of 20 species that have been previously reported from Iraq have been excluded from the checklist. According to IUCN Red List criteria, among 77 naturally distributed species (alien species not included), 10 species (13.0%) are categorized as threatened extinctions, with three (3.9%) as CR, one (1.3%) as EN, and six (7.8%) as VU. A total of 38 taxa were assessed (49.4%) as LC, 27 species were not assigned NE (35.1%), and two species (2.0%) were classified as DD, i.e., with insufficient knowledge.

 

References

  1. Abdullah, Y.S. & Abdullah, S.M.A. (2018) Ichthyofauna of Darbandikhan Lake in Kurdistan Region, Iraq. ZANCO Journal of Pure and Applied Sciences, 30 (6), 130–134. https://doi.org/10.21271/ZJPAS.30.6.12
  2. Abdulrahman, N.M., Ahmed, V.M., Ameen, H.J., Salih, H.A., Latif, B.M., Ali, S.S. & Ahmed, A.A. (2017) Comparative study of on fish health indices of rainbow trout (Oncorhynchus mykiss) reared in two sites in Sulaimani governorate, north of Iraq. Biological and Applied Environmental Research, 1 (2), 228–236.
  3. Abed, J.M., Jawad, L.A. & Ibánˇez, A.N.A.L. (2020) Morphometric and meristic variations of Photopectoralis bindus (Valenciennes, 1835) in the waters Iraq: implications for stock identification and management. Bollettino del Museo Civico di Storia Naturale di Verona Botanica Zoologia, 44, 61–74.
  4. Abood, A.N. (2010) Taxonomical study on mugilid species in Iraqi marine waters and the Garmet Ali river. M. Sc. Thesis, College of Agriculture, University of Basrah, Basra, 168 pp.
  5. Agha, G.F., Bilal, S.J. & Abdullah, S.M.A. (2023) DNA barcoding of some species of the genus Capoeta Valenciennes, 1842 from Kurdistan Region, Iraq. ZANCO Journal of Pure and Applied Sciences, 35 (1), 136–142. https://doi.org/10.21271/ZJPAS.35.1.14
  6. Al-Dubaikel, A.Y. (1986) Composition of fish species in Shatt Al-Basrah canal and their trophic relations. M. Sc. Thesis, College of Agriculture, University of Basrah, 118 pp. [in Arabic]
  7. Al-Faisal, A.J. & Mutlak, F.M. (2014) First record of the Nile tilapia Oreochromis niloticus (Linnaeus, 1758) from the Shatt al-Arab River, southern Iraq. Mesopotamian Journal of Marine Sciences, 29 (1), 45–50. https://doi.org/10.58629/mjms.v29i1.139
  8. Al-Faisal, A.J. & Mutlak, F.M. (2018) Survey of the marine fishes in Iraq. Bulletin of the Iraq Natural History Museum, 15 (2), 163–177. https://doi.org/10.26842/binhm.7.2018.15.2.0163
  9. Al-Faisal, A.J.M., Mutlak, F.M. & Abdullah, S.A. (2014) Exotic freshwater fishes in the Southern Iraq. Marsh Bulletin, 9 (1), 65–78.
  10. Al-Faisal, A.J. (2020) Updating checklist of freshwater fishes of Iraq. Mesopotamia Environmental Journal, 5 (4), 1–7.
  11. Al-Hassan, L.A.J, Hussain, N.A. & Soud, K.D. (1989) A preliminary annotated checklist of the fishes of Shatt Al-Arab river, Basrah, IRAQ. Polish Archives of Hydrobiology, 36 (2), 283–288.
  12. Al-Hassan, L.A.J. & Hussain, N.A. (1985) Hydrological parameters influencing the penetration of Arabian Gulf fishes into the Shatt al Arab River, Iraq. Cybium, 9 (1), 7–16.
  13. Al-Hassan, L.A.J. & Madhi, A.A. (1987) Enzyme polymorphisms in the mullet, Liza dussumeiri (sic) from Shatt al-Arab river, Khor al-Zubair and the Arabian Gulf. Biochemical Systematics and Ecology, 15 (2), 269–271. https://doi.org/10.1016/0305-1978(87)90030-5
  14. Al-Hassan, L.A.J. (1994) The silver carp, Hypophthalmichthys molitrix (Cyprinidae) in Shatt al-Arab River. Cybium, 18 (2), 204.
  15. Al-Hassan, L.A.J. & Naama, A.K. (1986) New records of some Arab Gulf fishes in the freshwater system of Iraq. Bulletin of Basrah Natural History Museum, 6, 35–44.
  16. Ali, A.H., Adday, T.K. & Khamees, N.R. (2018) Catalogue of marine fishes of Iraq. Biological and Applied Environmental Research, 2 (2), 298–368.
  17. Ali, M.D. (1985) Observations on lernaeosis and gyrodactylosis in carp fingerling raised in ponds. Journal of Biological Sciences Research, 16 (1), 125–13.
  18. Al-Nasiri, S.K. & Shamsul Hoda, S.M. (1975) Survey of fish fauna of Shatt Al-Arab (from Abul Khasib to Karmat Ali). Bulletin of the Iraq Natural History Museum, 2, 36–46.
  19. Al-Nasiri, S.K. & Shamsul Hoda, S.M. (1976) A guide to the freshwater fishes of Iraq. Bulletin of Basrah Natural History Museum, 1, 1–126.
  20. Al-Saadi, B.A., Mhaisen, F.T. & Al-Rubae, A.R.L. (2012) The first parasitological report on the redbelly tilapia, Tilapia zillii (Gervais, 1848) in Iraq. In: Proceedings of the Scientific Session "The Tilapia fish and its effect on Iraqi environment" of the Iraq. Natural History Research Centre and Museum, Baghdad, pp. 1–7.
  21. Bănărescu, P.M. & Nalbant, T.T. (1964) Süßwasserfische der Türkei. 2. Teil Cobitidae. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 61, 159–201, pls. 1–8.
  22. Banister, K.E. & Bunni, M.K. (1980) A new blind cyprinid fish from Iraq. Bulletin of the British Museum (Natural History) Zoology, 38 (3), 151–158. https://doi.org/10.5962/p.12611
  23. Barrois, T. (1894) Contribution à l’étude de quelques lacs de Syrie. Revue biologique du nord de la France, 6 (1893–1894), 224–312.
  24. Bartley, D.M. (2006) Introduced species in fisheries and aquaculture: information for responsibleusc and control. Food and Agri culture Organization, Rome. [CD-ROM]
  25. Basilewsky, S. (1855) Ichthyographia Chinae borealis. Nouveaux mémoires de la Société impériale des naturalistes de Moscou, 10, 215–263.
  26. Berg, L. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum from the basin of the Tigris River. Izvestiya Akad. Nauk SSSR, 7, 1267–1270.
  27. Berg, L.S. (1932) Eine neue Barilius-Art (Pisces, Cyprinidae) aus Mesopotamien. Zoologischer Anzeiger, 100 (11/12), 332–334.
  28. Berg, L.S. (1933) Les poissons des eaux douces de l’U.R.S.S. et des pays limitrophes. 3–e édition, revue et augmentée. Leningrad. Les poissons des eaux douces de l’U.R.S.S., 2, 544–903, 1 map.
  29. Berg, L.S. (1948) Ryby presnych vod SSSR i sopredelnych stan. [Freshwater fishes of the U.S.S.R. and adjacent countries]. Vol. 1. 4th Edition. Opredeliteli po faune SSSR. [Guide to the fauna of the U.S.S.R.], Moskva. Freshwater fishes of the U.S.S.R. and adjacent countries, No. 27, 1–466. [in Russian. English translation available, Israel Prog. Sci. Transl., Jerusalem, 1962, p. 1–504]
  30. Berg, L.S. (1949) Ryby presnych vod SSSR i sopredelnych stan. [Freshwater fishes of the U.S.S.R. and adjacent countries]. Vol. 2. 4th Edition. Opredeliteli po faune SSSR. [Guide to the Fauna of the U.S.S.R.], Moskva. Freshwater fishes of the U.S.S.R. and adjacent countries, No. 29, 467–925. [in Russian, English translation appeared in Israel Program of Scientific Translation, Jerusalem, 1964, pp. 1–496]
  31. Bleeker, P. (1850) Over eenige nieuwe soorten van Belone en Hemiramphus van Java. Natuurkundig Tijdschrift voor Nederlandsch Indië, 1 (2), 93–95.
  32. Bleeker, P. (1854) Species piscium Bataviensium novae vel minus cognitae. Natuurkundig Tijdschrift voor Nederlandsch Indië, 6 (2), 191–202.
  33. Bloch, M.E. (1782) D. Marcus Elieser Bloch’s, ausübenden Arztes zu Berlin. Oeconomische Naturgeschichte der Fische Deutschlands. Erster Theil. Auf Kosten des Verfassers und in Commission bei dem Buchhändler Hr. Hesse, Berlin, 128 pp., pls. 1–37. https://doi.org/10.5962/bhl.title.63303
  34. Bloch, M.E. (1786) Naturgeschichte der ausländischen Fische. Vol. 2. Aus Kosten der Verfassers, und in Commission in der Buchhandlung der Realschule, Berlin, viii + 160 pp., pls. 145–180.
  35. Bloch, M.E. (1793) Naturgeschichte der ausländischen Fische. Vol. 7. J. Morino, Berlin, xiv + 144 pp., pls. 325–360.
  36. Bloch, M.E. (1794) Naturgeschichte der ausländischen Fische. Vol. 8. Verfasser, Berlin, iv + 174, pls. 361–396.
  37. Bloch, M.E. (1795) Naturgeschichte der ausländischen Fische. Vol. 9. Schlesinger, Berlin, ii + 192 pp., pls. 397–429.
  38. Bloch, M.E. & Schneider, J.G. (1801) M.E. Blochii, Systema Ichthyologiae Iconibus cx Ilustratum. Post obitum auctoris opus inchoatum absolvit, correxit, interpolavit Jo. Gottlob Schneider, Saxo. Berolini. Sumtibus Auctoris Impressum et Bibliopolio Sanderiano Commissum, Berolini, lx + 584, pls. 1–110. https://doi.org/10.5962/bhl.title.5750
  39. Borkenhagen, K. & Krupp, F. (2013) Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae). ZooKeys, 339, 1–53. https://doi.org/10.3897/zookeys.339.4903
  40. Brandt, J.F. & Ratzeburg, J.T.C. (1833) Medizinische Zoologie, oder getreue Darstellung und Beschreibung der Thiere, die in der Arzneimittellehre in Betracht kommen, in systematischer Folge herausgegeben. Vol. 2. A. Hirschwald, Berlin, iv + 364 pp., pls. 1–36 + 1.
  41. Burchell, W.J. (1822) Travels in the interior of southern Africa. 2 Vols. Printed for Longman, Hurst, Rees, Orme, and Brown, London, xi + 582 + 4 pp., 1 map & 648 pp. https://doi.org/10.5962/bhl.title.100911
  42. Cambray, J.A. (2003) Impact on indigenous species biodiversity caused by the globalisation of alien recreational freshwater fishes. Hydrobiologia, 500, 217–230. https://doi.org/10.1023/A:1024648719995
  43. Cantor, T.E. (1842) General features of Chusan, with remarks on the flora and fauna of that island. Annals and Magazine of Natural History, New Series, 9 (58, 59 & 60), 265–278, 361–370 & 481–493. https://doi.org/10.5962/bhl.title.6704
  44. Castelnau, F.L. (1855) Poissons. In: Animaux nouveaux or rares recueillis pendant l'expédition dans les parties centrales de l'Amérique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; exécutée par ordre du gouvernement Français pendant les années 1843 a 1847. Part 7. Tome 2. Zoologie. Chez P. Bertrand, Paris, pp. i–xii + 1–112, pls. 1–50. https://doi.org/10.5962/bhl.title.110120
  45. Çiçek, E., Birecikligil, S. & Fricke, R. (2015) Freshwater fishes of Turkey; a revised and updated annotated checklist. Biharean Biologist, 9, 141–157.
  46. Çiçek, E., Eagderi, S. & Sungur, S. (2022) A review of the alien fishes of Turkish inland waters. Turkish Journal of Zoology, 46, 1–13. https://doi.org/10.55730/1300-0179.3104
  47. Çiçek, E., Sungur, S. & Fricke, R. (2020) Freshwater lampreys and fishes of Turkey; a revised and updated annotated checklist 2020. Zootaxa, 4809 (2), 241–270.
  48. Çi̇çek, E., Eagderi, S., Seçer, B. & Sungur, S. (2021) Capoeta kosswigi Karaman, 1969 a junior synonym of Capoeta damascina (Valenciennes, 1842) (Teleostei: Cyprinidae). Turkish Journal of Zoology, 45 (3), 235–240. https://doi.org/10.3906/zoo-2012-36
  49. Çiçek, E., Fricke, R., Sungur, S. & Eagderi, E. (2018) Endemic freshwater fishes of Turkey. FishTaxa, 3 (4), 1–39.
  50. Çiçek, E., Sungur, S., Fricke, R. & Seçer, B. (2023) Freshwater lampreys and fishes of Türkiye; an annotated checklist-2023. Turkish Journal of Zoology. [accepted]
  51. Coad, B.W. (2009) A new species of tooth-carp, Aphanius mesopotamicus, from Iran and Iraq (Actinopterygii, Cyprinodontidae). ZooKeys, 31, 149–163. https://doi.org/10.3897/zookeys.31.131
  52. Coad, B.W. & Najafpour, N. (1997) Barbus sublimus, a new species of cyprinid fish from Khuzestan Province, Iran. Ichthyological Exploration of Freshwaters, 7 (3), 273–278.
  53. Coad, B.W. & Nalbant, T.T. (2005) A new genus and a new species of a remarkable nemacheilid fish from Iran (Pisces: Ostariophysi: Nemacheilidae). Travaux du Muséum d’Histoire Naturelle "Grigore Antipa", 48, 303–308.
  54. Coad, B.W. (1991) Fishes of the Tigris-Euphrates basin: A critical checklist. Syllogeus, Ottawa, 68, 1–49.
  55. Coad, B.W. (1996) Exotic and transplanted fishes in southwest Asia. Proceedings of the 8th Congress of Societas Europaea Ichthyologorum (SEI), Oviedo, Spain, 26 September 26–2 October1994, 1996, 81–106.
  56. Coad, B.W. (2010) Freshwater fishes of Iraq. Pensoft Publication, Sofia, 274 pp., 16 pls.
  57. Coad, B.W. & Hussain, N.A. (2007) First record of the exotic species Hemiculter leucisculus (Actinopterygii: Cyprinidae) in Iraq. Zoology in the Middle East, 40 (1), 107–109. https://doi.org/10.1080/09397140.2007.10638212
  58. Collette, B.B. & Parin, N.V. (1978) Five new species of halfbeaks (Hemiramphidae) from the Indo-west Pacific. Proceedings of the Biological Society of Washington, 91 (3), 731–747.
  59. Cucherousset, J. & Olden, J.D. (2011) Ecological impacts of non-native freshwater fishes. Fisheries, 36, 215–230. https://doi.org/10.1080/03632415.2011.574578
  60. Cuvier, G. (1816) Le Règne Animal distribué d’après son organisation pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Tome II. Les reptiles, les poissons, les mollusques et les annélides. 1st Edition. Chez Déterville, Paris, xviii + 532 pp., pls. 9–10 in Vol. 4.
  61. Cuvier, G. & Valenciennes, A. (1828) Histoire naturelle des poissons. Tome Second. Livre Troisième. Des poissons de la famille des perches, ou des percoïdes. Chez F. G. Levrault, Paris, xxi + 2 + 490 pp., pls. 9–40.
  62. Cuvier, G. & Valenciennes, A. (1829) Histoire naturelle des poissons. Tome Troisième. Suite du Livre troisième. Des percoïdes à dorsale unique à sept rayons branchiaux et à dents en velours ou en cardes. Chez F. G. Levrault, Paris, xxviii + 2 + 500 pp., pls. 41–71.
  63. Cuvier, G. & Valenciennes, A. (1830) Histoire naturelle des poissons. Tome Sixième. Livre sixième. Partie I. Des Sparoïdes; Partie II. Des Ménides. Chez F. G. Levrault, Paris, xxiv + 6 + 559 pp., pls. 141–169.
  64. Cuvier, G. & Valenciennes, A. (1831) Histoire naturelle des poissons. Tome Septième. Livre septième. Des Squamipennes. Livre huitième. Des poissons à pharyngiens labyrinthiformes. Chez F. G. Levrault, Paris, xxix + 531 pp., pls. 170–208.
  65. Cuvier, G. & Valenciennes, A. (1833) Histoire naturelle des poissons. Tome Neuvième. Suite du livre neuvième. Des Scombéroïdes. Chez F. G. Levrault, Paris, xxix + 3 + 512 pp., pls. 246–279.
  66. Cuvier, G. & Valenciennes, A. (1835) Histoire naturelle des poissons. Tome Dixième. Suite du livre neuvième. Scombéroïdes. Livre dixième. De la famille des Teuthyes. Livre onzième. De la famille des Taenioïdes. Livre douzième. Des Athérines. Chez F. G. Levrault, Paris, xxiv + 482 + 2 pp., pls. 280–306.
  67. Cuvier, G. & Valenciennes, A. (1836) Histoire naturelle des poissons. Tome Onzième. Livre treizième. De la famille des Mugiloïdes. Livre quatorzième. De la famille des Gobioïdes. Chez F. G. Levrault, Paris, xx + 506 + 2 pp., pls. 307–343.
  68. Cuvier, G. & Valenciennes, A. (1837) Histoire naturelle des poissons. Tome Douzième. Suite du livre quatorzième. Gobioïdes. Livre quinzième. Acanthoptérygiens à pectorales pédiculées. Chez F. G. Levrault, Paris, xxiv + 507 + 1 pp., pls. 344–368.
  69. Cuvier, G. & Valenciennes, A. (1842) Histoire naturelle des poissons. Tome Seizième. Livre dix-huitième. Les Cyprinoïdes. Chez P. Bertrand, Paris, xx + 472 pp., pls. 456–487.
  70. Cuvier, G. & Valenciennes, A. (1844) Histoire naturelle des poissons. Tome Dix-Septième. Suite du livre dix-huitième. Cyprinoïdes. Chez P. Bertrand, Paris, xxiii + 497 + 2 pp., pls. 487–519.
  71. Cuvier, G. & Valenciennes, A. (1847) Histoire naturelle des poissons. Tome Vingtième. Livre vingt et unième. De la famille des Clupéoïdes. Chez P. Bertrand, Paris, xviii + 1 + 472 pp., pls. 591–606.
  72. Cuvier, G. & Valenciennes, A. (1847) Histoire naturelle des poissons. Tome Dix-Neuvième. Suite du livre dix-neuvième. Brochets ou Lucioïdes. Livre vingtième. De quelques familles de Malacoptérygiens, intermédiaires entre les Brochets et les Clupes. Chez P. Bertrand, Paris, xix + 544 + 6 pp., pls. 554–590.
  73. Davidson, N.C. (2014) How much wetland has the world lost? Long-term and recent trends in global wetland area. Marine and Freshwater Research, 65, 934–941. https://doi.org/10.1071/MF14173
  74. Day, F. (1872) Notes on fish, collected by Dr. Stoliczka in Kachh. Journal of the Asiatic Society of Bengal, 41 (1–4), 258–260.
  75. Day, F. (1888) Observations on the fishes of India.–Part I. Proceedings of the Zoological Society of London, 1888 (3) (Art. 4) (for 1 May 1888), 258–265. https://doi.org/10.1111/j.1469-7998.1888.tb06706.x
  76. Eagderi, S., Mouludi-Saleh, A., Esmaeli, H.R., Sayyadzadeh, G. & Nasri, M. (2022) Freshwater lamprey and fishes of Iran; a revised and updated annotated checklist—2022. Turkish Journal of Zoology, 46 (6), 500–522. https://doi.org/10.55730/1300-0179.3104
  77. Elvira B. (2001) Identification of non-native freshwater fishes established in Europe and assessment of their potential threats to the biological diversity. Convention on the conservation of European wildlife and natural habitats. Council of Europe T-PVS 6. Standing Committee, 21st Meeting, Strasbourg, 26–30 November 2001. Document prepared by Benigno Elvira Department of Animal Biology I, Faculty of Biology, University Complutense of Madrid, EMadrid and funded by the Ministry of the Walloon Region, 35 pp.
  78. Esmaeili, H.R. (2021) Checklist of freshwater fishes of southwestern wetlands of Iran. In: Jawad, L.A. (Ed.), Southern Iraq's Marshes: Their Environment and Conservation. Springer International Publishing, Cham, pp. 295–318. https://doi.org/10.1007/978-3-030-66238-7_17
  79. Esmaeili, H.R., Gholamifard, A. & Freyhof, J. (2011) Ichthyofauna of Zarivar Lake (Iran) with the first records of Hemiculter leucisculus and Alburnus hohenackeri in the Tigris drainage. Electronic Journal of Ichthyology, 7 (1), 1–6.
  80. Esmaeili, H.R., Jufaili, S.A., Masoumi, A.H. & Zarei, F. (2022b) Ichthyodiversity in southeastern Arabian Peninsula: Annotated checklist, taxonomy, short description and distribution of Inland fishes of Oman. Zootaxa, 5134 (4), 451–503. https://doi.org/10.11646/zootaxa.5134.4.1
  81. Esmaeili, H.R., Khaefi, R. & Nejad, R.Z. (2016b) Historical review on the taxonomy of Squalius berak Heckel, 1843 (Teleostei: Cyprinidae). FishTaxa, 1 (3), 118–126.
  82. Esmaeili, H.R., Masoudi, M. & Mehraban, H.R. (2014) Assignment of Acanthopagrus populations in the Persian Gulf drainage system of Iran to Acanthopagrus arabicus Iwatsuki, 2013 (Perciformes: Sparidae). Iranian Journal of Ichthyology, 1 (1), 23–28.
  83. Esmaeili, H.R., Masoudi, M., Chermahini, M.A., Esmaeili, A.H., Zarei, F. & Ebrahimi, M. (2017) Invasion of the Neotropical and Nearctic fishes to Iran. FishTaxa, 2 (3), 126–133.
  84. Esmaeili, H.R., Sayyadzadeh, G., Zarei, F., Eagderi, S. & Mousavi-Sabet, H. (2022a) Mystus cyrusi, a new species of bagrid catfish (Teleostei: Bagridae) from Middle East. Zootaxa, 5099 (3), 325–343. https://doi.org/10.11646/zootaxa.5099.3.2
  85. Esmaeili, H.R., Teimori, A., Zarei, F. & Sayyadzadeh, G. (2020) DNA barcoding and species delimitation of the Old World tooth-carps, family Aphaniidae Hoedeman, 1949 (Teleostei: Cyprinodontiformes). PLoS ONE, 15 (4), e0231717, 1–26. https://doi.org/10.1371/journal.pone.0231717
  86. Esmaeili, H.R., Coad, B.W., Gholamifard, A., Nazari, N. & Teimory, A. (2010) Annotated checklist of the freshwater fishes of Iran. Zoosyst Rossica, 19, 361–386. https://doi.org/10.31610/zsr/2010.19.2.361
  87. Esmaeili, H.R., Mehraban, H., Abbasi, K., Keivany, Y. & Coad, B. (2017) Review and updated checklist of freshwater fishes of Iran: taxonomy, distribution and conservation status. Iranian Journal of Ichthyology, 4 (Supplement 1), 1–114.
  88. Esmaeili, H.R., Sayyadzadeh, G., Özuluğ, M., Geiger, M. & Freyhof, J. (2014) Three new species of Turcinoemacheilus from Iran and Turkey (teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 24 (3), 257–273.
  89. Freyhof, J., Esmaeili, H.R., Sayyadzadeh, G. & Geiger, M. (2014) Review of the crested loaches of the genus Paracobitis from Iran and Iraq with the description of four new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 25 (1), 11–38. https://doi.org/10.11646/zootaxa.4048.1.4
  90. Freyhof, J. (2016) Redescription of Garra elegans (Günther, 1868), a poorly known species from the Tigris River drainage (Teleostei: Cyprinidae). Zootaxa, 4173 (5), 496–500. https://doi.org/10.11646/zootaxa.4173.5.7
  91. Freyhof, J. & Abdullah, Y.S. (2017) Two new species of Oxynoemacheilus from the Tigris drainage in Iraqi Kurdistan (Teleostei: Nemacheilidae). Zootaxa, 4238 (1), 73–87. https://doi.org/10.11646/zootaxa.4238.1.5
  92. Freyhof, J., Kaya, C. & Turan, D. (2017) Oxynoemacheilus kentritensis, a new species from the upper Tigris drainage in Turkey with remarks on O. frenatus (Teleostei: Nemacheilidae). Zootaxa, 4258 (6), 551–560. https://doi.org/10.11646/zootaxa.4258.6.4
  93. Freyhof, J., Esmaeili, H.R., Sayyadzadeh, G. & Geiger, M.F. (2014) Review of the crested loaches of the genus Paracobitis from Iran & Iraq with the description of four new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 25 (1), 11–38.
  94. Freyhof, J. & Geiger, M.F. (2017) Oxynoemacheilus zarzianus, a new loach from the Lesser Zab River drainage in Iraqi Kurdistan (Teleostei: Nemacheilidae). Zootaxa, 4273 (2), 258–270. https://doi.org/10.11646/zootaxa.4273.2.6
  95. Freyhof, J., Kaya, C., Abdullah, Y.S. & Geiger, M.F. (2021b) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453–491. https://doi.org/10.11646/zootaxa.4969.3.2
  96. Freyhof, J., Kaya, C. & Ali, A. (2021a) A critical checklist of the inland fishes native to the Euphrates and Tigris drainages. In: Tigris and Euphrates Rivers: their environment from headwaters to mouth. Aquatic Ecology Series. Vol. 11. Springer, Cam, pp. 815–854. https://doi.org/10.1007/978-3-030-57570-0_35
  97. Freyhof, J., Abdullah, Y.S., Ararat, K., Ibrahim, H. & Geiger, M.F. (2016) Eidinemacheilus proudlovei, a new subterranean loach from Iraqi Kurdistan (Teleostei; Nemacheilidae). Zootaxa, 4173 (3), 225–236. https://doi.org/10.11646/zootaxa.4173.3.2
  98. Freyhof, J., Kaya, C. & Geiger, M.F. (2022) A practical approach to revise the Oxynoemacheilus bergianus species group (Teleostei: Nemacheilidae). Zootaxa, 5128 (2), 151–194. https://doi.org/10.11646/zootaxa.5128.2.1
  99. Fricke, R., Eschmeyer, W.N. & Van der Laan, R. (Eds.) (2023) Eschmeyer’s catalog of fishes: genera, species, references. Electronic Version. Available from: http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (accessed 30 April 2023) (2) (PDF) Patterns in Freshwater Fish Diversity. Available from: https://www.researchgate.net/publication /354892201_Patterns_in_Freshwater_Fish_Diversity#fullTextFileContent (accessed 27 May 2023)
  100. Froese, R. & Torres, A. (1999) Fishes under threat: an analysis of the fishes in the 1996 IUCN Red List. In: Pullin, R.S.V., Bartley, D.M. & Kooiman, J. (Eds.), Towards policies for conservation and sustainable use of quatic genetic resources. ICLARM Conference Proceedings, 59, pp. 131–144.
  101. Froese, R. & Pauly, D. (Eds.) (2022) FishBase. World Wide Web Electronic Publication. Version August 2022. Available from: http://www.fishbase.org (accessed 6 February 2023)
  102. Garcia-Berthou, E. & Moyle, P.B. (2011) Rivers. In: Simberloff, D. & Rejmanek, M. (Eds.), Encyclopaedia of Biological Invasions. University of California Press, Berkeley, Los Angeles, Califonria, pp. 609–612. https://doi.org/10.1525/9780520948433-136
  103. Geldiay, R. & Balık, S. (2007) Türkiye Tatlısu Balıkları. Ege Üniversitesi Fen Fakültesi Kitaplar Serisi, No. 46. Ege Üniversitesi Basımevi, Bornova-İzmir, 644 p.
  104. Gervais, F.L.P. (1848) Sur les animaux vertébrés de l’Algérie, envisagés sous le double rapport de la géographie zoologique et de la domestication. Annales des Sciences Naturelles, Paris, Zoologie, Série 3, 10, 202–208. https://doi.org/10.5962/bhl.title.51497
  105. Gholamhosseini, A., Razbanian, M., Esmaeili, H.R. & Eagderi, S. (2022) Molecular systematics and morphological variation in the Mesopotamian spiny eel Mastacembelus mastacembelus (Teleostei: Mastacembelidae). The European Zoological Journal, 89 (1), 546–555. https://doi.org/10.1080/24750263.2022.2057604
  106. Gholami, Z., Esmaeili, H.R., Erpenbeck, D. & Reichenbacher, B. (2014) Phylogenetic analysis of Aphanius from the endorheic Kor River Basin in the Zagros Mountains, south-western Iran (Teleostei: Cyprinodontiformes: Cyprinodontidae). Journal of Zoological Systematics and Evolutionary Research = Zeitschrift für zoologische Systematik und Evolutionsforschung, 52 (2), 130–141. https://doi.org/10.1111/jzs.12052
  107. Girard, C.F. (1859) Ichthyological notices. Proceedings of the Academy of Natural Sciences of Philadelphia, 11, 56–68.
  108. Günther, A. (1868) Catalogue of the fishes in the British Museum. Vol. 7. Catalogue of the Physostomi, containing the families Heteropygii, Cyprinidae, Gonorhynchidae, Hyodontidae, Osteoglossidae, Clupeidae, Chirocenridae, Alepocephalidae, Notopteridae, Halosauridae. in the collection of the British Museum. Vol. 7. Department of Zoology, British Museum (Natural History), London, xx + 512 pp.
  109. Günther, A. (1874) A contribution to the fauna of the river Tigris. Annals and Magazine of Natural History, Series 4, 14 (79) (Art. 8), 36–38, pls. 8–9. https://doi.org/10.1080/00222937408680918
  110. Hamilton, F. (1822) An account of the fishes found in the river Ganges and its branches. Edinburgh & London, vii + 405 pp., 39 pls. https://doi.org/10.5962/bhl.title.59540
  111. Heckel, J.J. (1843) Ichthyologie [von Syrien]. In: von Russegger, J., Reisen in Europa, Asien und Afrika, mit besonderer Rücksicht auf die naturwissenschaftlichen Verhältnisse der betreffenden Länder unternommen in den Jahren 1835 bis 1841, etc. E. Schweizerbart‘sche Verlagshandlung. Stuttgart. Ichthyologie [von Syrien], 1 (pt 2), 991–1099.
  112. Heckel, J.J. (1847) Naturhistorischer Anhang. [Various subtitles]. In: von Russegger, J., Reisen in Europa, Asien und Afrika: mit besonderer Rücksicht auf die naturwissenschaftlichen Verhältnisse der betreffenden Länder; unternommen in den Jahren 1835 bis 1841. E. Schweizerbart’sche Verlagshandlung, Stuttgart, 2 (3), pp. 207–357.
  113. Hora, S.L. & Misra K.S. (1943) On a small collection of fish from Iraq. Journal of the Asiatic Society of Bengal, Science, Series 3, 9 (1), 1–15.
  114. Hussain, N.A. & Ali, T.S. (1987) Some biological aspects of Thryssa hamiltoni and Thryssa mystax in Khor Al-Zubair, northwest Arabian Gulf. Indian Journal of Fisheries, 34 (2), 152–162.
  115. Hussain, N.A., Mohamed, A.M., Al-Noor, S.S., Mutlak, F.M., Abed, I.M. & Coad, B.W. (2009) Structure and ecological indices of fish assemblages in the recently restoed Al-hammar Marsh, southern Iraq. BioRisk, 3, 173–186. https://doi.org/10.3897/biorisk.3.11
  116. Hussain, N.A., Naiama, A.K. & Al-Hassan, L.A.J. (1988) Annotated check list of the fish fauna of Khor Al-Zubair, north west of the Arabian Gulf, Iraq. Acta Ichthyologica et Piscatoria, 18 (1), 17–24. https://doi.org/10.3750/AIP1988.18.1.02
  117. Hussain, N.A., Rasen, A.K., Al, B.Y. & Coad, B.W. (2012) Bull shark occurrence Carcharhinus leucas (Valenciennes, 1839) at the inland waters of southern Iraq. Journal of Duhok University, 15 (1), 140–143.
  118. Hussain, N.A., Younis, K.H. & Yousif, U.H. (1999) Seasonal fluctuations of the fish assemblage of intertidal mudflats of the Shatt Al-Arab estuary, Iraq, northwestern Arabian Gulf. Mesopotamian Journal of Marine Science, 14 (1), 33–53.
  119. Iwatsuki, Y. (2013) Review of the Acanthopagrus latus complex (Perciformes: Sparidae) with descriptions of three new species from the Indo-West Pacific Ocean. Journal of Fish Biology, 83 (1), 64–95. https://doi.org/10.1111/jfb.12151
  120. Jawad, L. (2003) Impact of environmental change on the freshwater fish fauna of Iraq. International Journal of Environmental Studies, 60 (6), 581–593. https://doi.org/10.1080/0020723032000087934
  121. Jawad, L., Mutlak, F. & Abbas, A.F. (2016) On the record of vertebral deformities in Mastacembelus mastacembelus collected from the lower reaches of Euphrates River, Iraq. Boletim do Instituto de Pesca, 42 (1), 216–220. https://doi.org/10.20950/1678-2305.2016v42n1p216
  122. Jawad, L.A. & Qasim, A.M. (2019) Neotropical piranha, Serrasalmus maculatus (Actinopterygii: Characiformes: Serrasalmidae), in the Tigris River, Baghdad, Iraq—a case of deliberate introduction by the aquarium trade. Acta Ichthyologica et Piscatoria, 49 (2), 199–201. https://doi.org/10.3750/AIEP/02526
  123. Jawad, L.A. (2015) The Indian catfish, Heteropneustes fossilis (Bloch, 1794) Envenomation—A report of four cases from Basrah and Maysan, South of Iraq. TOFIQ Journal of Medical Sciences, 2 (2), 52–61.
  124. Jawad, L.A. (2021) First Reports on Cases of Hallucinatory Fish Poisoning (Ichthyoallyeinotoxism) and Scombrotoxic Fish Poisoning in Iraq. Tigris and Euphrates Rivers: Their Environment from Headwaters to Mouth. Aquatic Ecology Series. Vol. 11. Springer, Cham, pp. 1499–1504. https://doi.org/10.1007/978-3-030-57570-0_75
  125. Jawad, L.A., Abed, J., Mohsen, Z. & Al-Janabi, M. (2021a) A confirmed record of the European catfish Silurus glanis L., 1758 (Actinopterygii: Siluriformes: Siluridae) from the southern marshes of Iraq, with a new anatomical set of characters to separate S. glanis and S. triostegus. Integrative Systematics: Stuttgart Contributions to Natural History, 3 (1), 85–100. https://doi.org/10.18476/insy.v03.a6
  126. Jawad, L.A., Abed, J.M., Hussain, S.A. & Ünlü, E. (2018) Vertebral Dimensions in the Genus Carasobarbus (Teleostei: Cyprinidae). Journal of Ichthyology, 58, 780–794. https://doi.org/10.1134/S003294521806005X
  127. Jawad, L.A., Al-Faisal, A. & Al-Mukhtar, M. (2012) Presence of the non–indigenous fish species, Gibel carp, Carassius gibelio (Family: Cyprinidae) in Basrah Province freshwater systems, Iraq. Water Research and Management, 2 (4), 41–44.
  128. Jawad, L.A., Al-Mukhtar, M. & Subr, S.H. (2016) Occurrence of the population of Photopectoralis bindus (Valenciennes, 1835) in Shatt al-Arab river, Basrah, Iraq. International Journal of Marine Science, 6, 1–4. https://doi.org/10.5376/ijms.2016.06.0057
  129. Jawad, L.A., Al-Sheikhly, O.F. & Al-Fayadhi, K.H. (2022) The oscar, Astronotus ocellatus (Agassiz, 1831) (Cichlidae): a deliberate aquarium trade introduction in the Euphrates River, Iraq. Thalassia Salentina, 44, 41–50.
  130. Jawad, L.A., Habbeb, F.S. & Al-Mukhtar, M.A. (2018) Morphometric and meristic characters of two cichlids, Coptodon zillii and Oreochromis aureus collected from Shatt al-Arab River, Basrah, Iraq. International Journal of Marine Science, 8 (2), 12–25. https://doi.org/10.5376/ijms.2018.08.0002
  131. Jawad, L.A., Hussein, S.A. & Fahad, K.K. (2009) Glyptothorax kurdistanicus (Berg, 1931) (Pisces, Siluriformes, Sisoridae) in the lower reaches of the Tigris River, Iraq?. Journal of Applied Ichthyology, 25 (6), 779–781. https://doi.org/10.1111/j.1439-0426.2009.01314.x
  132. Jawad, L.A., Jassim, A.K., Tahir Ankush, M.A.W. & Abed, J.M. (2019) Flushed with the flood: the rainbow trout Oncorhynchus mykiss in the Shatt Al-Arab River, Basrah, Iraq. Thalassia Salentina, 41, 119–126.
  133. Jawad, L.A., Mutlak, F., Mohamed, A.-R.M., Al-Mamry, J.M., Hameed, E.K. & Ibrahim, M. (2021b) Juveniles and larvae of eleven fish species collected from three countries of the Arabian Gulf and the Arabian Sea. Proceedings of the Zoological Institute RAS, 325 (4), 457–468. https://doi.org/10.31610/trudyzin/2021.325.4.457
  134. Jawad, L.A. & Qasim, A.M. (2019) Neotropical piranha, Serrasalmus maculatus (Actinopterygii: Characiformes: Serrasalmidae), in the Tigris River, Baghdad, Iraq—a case of deliberate introduction by the aquarium trade. Acta Ichthyologica et Piscatoria, 49 (2), 199–201. https://doi.org/10.3750/AIEP/02526
  135. Jawad, L.A., Al-Sheikhly, O.F. & Al-Dirawi, A.M. (2021) The Danube Sturgeon Acipenser gueldenstaedtii Brandt & Ratzeburg, 1833 (Actinopterygii, Acipenseridae) in the Euphrates River, Iraq. Acta Scientiarum Polonorum. Zootechnica, 20 (2), 33–38. https://doi.org/10.21005/asp.2021.20.2.04
  136. Kamangar, B.B., Prokofiev, A.M. Ghaderi, E. & Nalbant, T.T. (2014) Stone loaches of Choman River system, Kurdistan, Iran (Teleostei: Cypriniformes: Nemacheilidae). Zootaxa, 3755 (1), 33–61. https://doi.org/10.11646/zootaxa.3755.1.2
  137. Karaman, M.S. (1971) Süßwasserfische der Türkei. 8. Teil. Revision der Barben Europas, Vorderasiens und Nordafrikas. Mitteilungen aus dem hamburgischen Zoologischen Museum und Institut, 67, 175–254.
  138. Kaya, C., Turan, D. & Ünlü, E. (2016) The latest status and distribution of fishes in upper Tigris River and two new records for Turkish freshwaters. Turkish Journal of Fisheries and Aquatic Sciences, 16 (3), 545–562. https://doi.org/10.4194/1303-2712-v16_3_07
  139. Kennedy, W.P. (1937) Some additions to the fauna of Iraq. Journal of the Bombay Natural History Society, 39, 745–749.
  140. Khalaf, K.T. (1961) The marine and freshwater fishes of Iraq. Al-Rabitta Press, Baghdad: 164 pp.
  141. Khaefi, R., Esmaeili, H.R., Geiger, M.F. & Eagderi, S (2017a) Taxonomic review of the cryptic Barbus lacerta species group with description of a new species (Teleostei: Cyprinidae). FishTaxa, 2 (2), 90–115.
  142. Khaefi, R., Teimori, A. & Esmaeili, H.R. (2017) Phylogenetic relationships and taxonomy of Luciobarbus barbulus (Heckel, 1847) (Teleostei: Cyprinidae). Journal of Ichthyology, 57, 835–845. https://doi.org/10.1134/S0032945217060078
  143. Khamees, N.R., Ali, A.H., Abed, J.M. & Adday, T.K. (2013) First record of striped catfish Pangasianodon hypophthalmus (Sauvage, 1878) (Pisces: Pangasiidae) from inland waters of Iraq. Basrah Journal of Agricultural Sciences, 26 (1), 184–197. https://doi.org/10.33762/bagrs.2013.111655
  144. Kner, R. (1858) Zur Familie der Characinen. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe, 32 (22), 163–168.
  145. Kottelat M. & Freyhof, J. (2007) Handbook of European Freshwater Fishes. Kottelat, Cornol and Freyhof, Berlin, xiii + 646 pp.
  146. Koumans, F.P. (1941) Gobioid fishes of India. Memoirs of the Indian Museum, 13 (3), 205–329.
  147. Krupp, F. & Schneider, W. (2008) Die Fischfauna des Nahr al-Habur, Nordost Syrien. In: Kühne, H. (Ed.), Umwelt und Subsistenz der assyrischen Stadt Dur-Katlimmu am unteren Habur, Berichte der Ausgrabung von Tall Seh Hamad/Dur-Katlimmu. (BATSH). Band 8. Harrassowitz, Wiesbaden, pp. 41–51.
  148. Lacepède, B.G.E. (1801) Histoire naturelle des poissons. Vol. 3. Chez F. G. Levrault, Paris, lxvi + 558 pp., pls. 1–34.
  149. Lacepède, B.G.E. (1803) Histoire naturelle des poissons. Vol. 5. Chez F. G. Levrault, Paris, lxviii + 803 pp., index, pls. 1–21.
  150. Ladiges, W. (1960) Süßwasserfische der Türkei, I. Teil Cyprinidae. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 58, 105–150.
  151. Landsberger, B. (1962) The fauna of ancient Mesopotamia. MSL VIII/2. 2nd Part. Pontificium Institutum Biblicum, Roma 204, Piazza Pilotta 35. Publié avec le concourse financier de l‘UNESCO et sous less auspices du Conseil International de la Philosophie et des Sciences Humaines, Paris, 180 pp.
  152. Liao, T.Y., Ünlü, E. & Kullander, S.O. (2011) Western boundary of the subfamily Danioninae in Asia (Teleostei, Cyprinidae): derived from the systematic position of Barilius mesopotamicus based on molecular and morphological data. Zootaxa, 2880 (1), 31–40. https://doi.org/10.11646/zootaxa.2880.1.3
  153. Lesueur, C.A. (1821a) Description of a new genus, and of several new species of fresh water fish, indigenous to the United States. Journal of the Academy of Natural Sciences, Philadelphia, 2 (1), 2–8.
  154. Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Impensis Direct. Laurentii Salvii, Holmiae, ii + 824 pp. https://doi.org/10.5962/bhl.title.542
  155. Linnaeus, C. (1766) Systema naturae sive regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tom. 1. Pt. 1. 12th Edition. Impensis direct. Laurentii Salvii, Holmiae, 532 pp. https://doi.org/10.5962/bhl.title.68927
  156. Mahdi, N. (19629) Fishes of Iraq. Ministry of Education, Baghdad, 82 pp.
  157. Maitland, P.S. (1992) The status of Arctic charr, Salvelinus alpinus L., in southern Scotland: a cause for concern. Freshwater Forum, 2, 212–227.
  158. McEachran, J.D. & Fechhelm, J.D. (1998) Fishes of the Gulf of Mexico. Vol. 1. Myxiniformes to Gasterosteiformes. Univeersity of Texas Press, Austin, Texas, 1112 pp. https://doi.org/10.7560/752061
  159. McKay, R.J. (1985) A revision of the fishes of the family Sillaginidae. Memoirs of the Queensland Museum, 22 (Pt 1), 1–73.
  160. McKay, R.J. & McCarthy, L.J. (1989) A revision of the sillaginid fishes of the Arabian Gulf with a description of Sillago arabica new species. Memoirs of the Queensland Museum, 27 (2), 551–553.
  161. Menon, M.A.S. (1956) On a third collection of fish from Iraq. Records of the Indian Museum, 54, 139–157. https://doi.org/10.26515/rzsi/v54/i3-4/1956/161996
  162. Misra, K.S. (1947) On a second collection of fish from Iraq. Records of the Indian Museum, 45, 115–127.
  163. Mohamed, A.-R.M. & Abood, A.N. (2017a) Compositional change in fish assemblage structure in the Shatt Al-Arab River, Iraq. Asian Journal of Applied Sciences, 5 (5), 944–958.
  164. Mohamed, A.-R.M. & Abood, A.N. (2017b) Ecological health assessment of the Shatt Al-Arab River, Iraq. IOSR Journal of Agriculture and Veterinary Science (IOSR-JAVS), 10 (10), 1–8. https://doi.org/10.9790/2380-1002021722
  165. Mohamed, A.-R.M. & Al-Wan, S.M. (2020) Biological aspects of an invasive species of Oreochromis niloticus in the Garmat Ali River, Basrah, Iraq. Journal of Agriculture and Veterinary Science, 13 (2), 15–26.
  166. Mohamed, A.-R.M. & Hameed, E.K. (2019) Impacts of saltwater intrusion on the fish assemblage in the middle part of Shatt Al-Arab River, Iraq. Asian Journal of Applied Sciences, 7 (5), 577–586. https://doi.org/10.24203/ajas.v7i5.5917
  167. Mohamed, A.-R.M., Hussain, N.A., Al-Noor, S.S., Coad, B.W. & Mutlak, F.M. (2009) Status of diadromous fish species in the restored East Hammar Marsh in Southern Iraq. American Fisheries Society Symposium, 69, 577–588.
  168. Mohamed, A.-R.M., Hussein, S.A. & Lazem, L.F. (2015) Spatiotemporal variability of fish assemblage in the Shatt Al-Arab River, Iraq. Journal of Coastal Life Medicine, 3 (1), 27–34.
  169. Mohamed, A.-R.M., Resen, A.K. & Taher, M.M. (2012) Longitudinal patterns of fish community structure in the Shatt Al-Arab River, Iraq. Basrah Journal of Science, 30 (2), 65–86.
  170. Mohamed, A.-R.M., Hussain, N.A. & Ali, T.S. (2001) Estuarine components of the ichthyofauna of the Arabian Gulf. Marina Mesopotamica, 16 (2), 209–224.
  171. Mohamed, A.-R.M., Abood, A.N. & Jawad, L.A. (2016) Presence of Carasobarbus sublimus (Coad & Najafpour, 1997) in the upper reaches of Shatt al-Arab River, Basrah. Zoology and Ecology, 27 (1), 30–34. https://doi.org/10.1080/21658005.2016.1272833
  172. Mohammadian-Kalat, T., Esmaeili, H.R. Aliabadian, M. & Freyhof, J. (2017) Re-description of Alburnus doriae, with comments on the taxonomic status of A. amirkabiri, A. mossulensis, A. sellal and Petroleuciscus esfahani (Teleostei: Cyprinidae). Zootaxa, 4323 (4), 487–502. https://doi.org/10.11646/zootaxa.4323.4.3
  173. Mousavi-Sabet, H., Eagderi, S., Saemi-Komsari, M., Kaya C. & Freyhof, J. (2022) Garra rezai, a new species from two widely disjunct areas in the Tigris drainage (Teleostei: Cyprinidae). Zootaxa, 5195 (5), 419–436. https://doi.org/10.11646/zootaxa.5195.5.2
  174. Mousavi-Sabet, H., Eagderi, S., Vatandoust, S. & Freyhof, J. (2021) Five new species of the sisorid catfish genus Glyptothorax from Iran (Teleostei: Sisoridae). Zootaxa, 5067 (4), 451–484. https://doi.org/10.11646/zootaxa.5067.4.1
  175. Mousavi-Sabet, H., Salehi, M., Sarpanah, A. & Pirali Kheirabadi, E. (2019) First records of Acipenser baerii and Huso huso (Actinopterygii: Acipenseriformes: Acipenseridae) from the Tigris-Euphrates Basin, Iran. Acta Ichthyologica Et Piscatoria, 49 (3), 265–267. https://doi.org/10.3750/AIEP/02510
  176. Mousavi-Sabet, H., Vatandoust, S., Esmaeili, H.R., Geiger, M.F. & Freyhof, J. (2015) Cobitis avicennae, a new species of spined loach from the Tigris River drainage (Teleostei: Cobitidae). Zootaxa, 3914 (5), 558–568. https://doi.org/10.11646/zootaxa.3914.5.4
  177. Müller, J. & Henle, F.G.J. (1838–41) Systematische Beschreibung der Plagiostomen. Veit und Comp., Berlin, xxii + 200 p., 60 pls. https://doi.org/10.5962/bhl.title.6906
  178. Mutlak, F.M. & Al-Faisal, A.T. (2009) A new record of two exotic cichlids fish Oreochromis aureus (Steindachner, 1864) and Tilapia zillii (Gervais, 1848) from the south of the main outfall drain in Basrah city. Mesopotamian Journal of Marine Science, 24, 160–170.
  179. Mutlak, F., Jawad, L. & Al-Faisal, A. (2017) Atractosteus spatula (Actinopterygii: Lepisosteiformes: Lepisosteidae): A deliberate aquarium trade introduction incidence in the Shatt al-Arab River, Basrah, Iraq. Acta Ichthyologica et Piscatoria, 47 (2), 205–207. https://doi.org/10.3750/AIEP/02143
  180. Myers, N., Mittermeier, R.A., Mittermeier, C.G., da Fonseca, G.A.B. & Kent, J. (2000) Biodiversity hotspots for conservation priorities. Nature, 403, 853–858. https://doi.org/10.1038/35002501
  181. Nader, I.A. & Jawdat, S.Z. (1977) New records of fishes from Iraq. Bulltin of the Biological Research Centre, Baghdad, 8, 73–87.
  182. Nelson, J.S., Grande, T.C. & Wilson, M.V.H. (2016) Fishes of the World. 5th Edition. John Wiley & Sons, Inc., Hoboken, New Jersey, 707 pp.
  183. Niebuhr, C. (1775) Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit Petrus Forskål. Post mortem auctoris edidit Carsten Niebuhr. ex officina Mölleri, Hauniae, 20 + xxxiv + 164 pp., map.
  184. Nyman, L. (1991) Conservation of freshwater fish: protection of biodiversity and genetic variability in aquatic ecosystems. Fish Devel Series, Gothenburg, 50, 1–38.
  185. Park, M. (1797) Descriptions of eight new fishes from Sumatra. The Transactions of the Linnean Society of London, 3 (9), 33–38. https://doi.org/10.1111/j.1096-3642.1797.tb00553.x
  186. Parmesan, C. (2006) Ecological and evolutionary responses to recent climate change. Annual Review of Ecology and Evolution, 37, 637–669. https://doi.org/10.1146/annurev.ecolsys.37.091305.110100
  187. Peters, W.(C.H.) (1859) Eine neue vom Herrn Jagor im atlantischen Meere gefangene Art der Gattung Leptocephalus, und über einige andere neue Fische des Zoologischen Museums. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin, 1859, 411–413.
  188. Peterson, A.T., Soberón, J. & Sánchez-Cordero, V. (1999) Conservatism of ecological niches in evolutionary time. Science, 285 (5431), 1265–1267. https://doi.org/10.1126/science.285.5431.1265
  189. Pietschmann, V. (1913) Eine neue Glyptosternum-Art aus dem Tigris. Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftlichen Klasse, 50 (8), 93–95.
  190. Postgate, J.N. (1994) Text and figure in ancient Mesopotamia: match and mismatch. In: Renfrew, A.C. & Zubrow, E. (Eds.), The Ancient Mind: Elements of cognitive archaeology. Cambridge University Press, Cambridge, pp. 176–184. https://doi.org/10.1017/CBO9780511598388.018
  191. Qasim, A.M. & Jawad, L.A. (2022) Presence of the Amazon sailfin catfish, Pterygoplichthys pardalis (Castelnau, 1855) (Pisces: Loricariidae), in the Shatt al-Arab River, Basrah, Iraq. Integrative Systematics: Stuttgart Contributions to Natural History, 5 (1), 95–103. https://doi.org/10.18476/2022.647187
  192. Qasim, A.M., Mutlak, F.M., Al-Faisal, A.J. & Jawad, L.A.J. (2019) December. Reports on the presence of the milkfish Chanos chanos (Forsskål, 1775) (Gonorynchiformes: Chanidae) in the Shatt al-Arab River, Basrah, and in the marine waters of Iraq, with notes on its tolerance to freshwater. Anales de Biología, 41, 83–87. https://doi.org/10.6018/analesbio.41.12
  193. Radkhah, A., Eagderi, S. & Mousavi-Sabet H. (2016) First record of the exotic species Hemiculter leucisculus (Pisces: Cyprinidae) in southern Iran. Limnetica, 35 (1), 175–178. https://doi.org/10.23818/limn.35.14
  194. Rauchenberger, M. (1989) Systematics and biogeography of the genus Gambusia (Cyprinodontiformes: Poecilidae). American Museum Novitates, 2951, 1–74.
  195. Richardson, J. (1845) Ichthyology. Part 3. In: Hinds, R.B. (Ed.), The zoology of the voyage of H. M. S. Sulphur, under the command of Captain Sir Edward Belcher, R. N., C. B., F. R. G. S., etc., during the years 1836–42, No. 10. Smith, Elder & Co., London, pp. 99–150, pls. 55–64.
  196. Risso, A. (1810) Ichthyologie de Nice, ou histoire naturelle des poissons du Département des Alpes Maritimes. F. Schoell, Paris, xxxvi + 388 pp., pls. 1–11. https://doi.org/10.5962/bhl.title.7052
  197. Russell, A. (1794) Natural History of Aleppo. 2nd Edition. Revised by P. Russell. Vol. 2. Printed for G.G. and J. Robinson, London, vii + 430 + xxxiv + 26 pp., index, pls. 1–16.
  198. Rzóska, J. & Banister, K.E. (1980) The fishes of the Tigris and Euphrates rivers. In: Euphrates and Tigris, Mesopotamian ecology and destiny. Dr. W. Junk, The Hague, pp. 95–108. https://doi.org/10.1007/978-94-009-9171-2_9
  199. Saad, A., Çiçek, E., Esmaeili, H.R., Fricke, R., Sungur, S. & Eagderi, S. (2023) Freshwater fishes of Syria: a revised and updated annotated checklist-2023. Zootaxa, 5350 (1), 1–623. https://doi.org/10.11646/zootaxa.5350.1.1
  200. Saggs, H.W.F. (1962) The greatness that was Babylon: a sketch of the ancient civilization of the Tigris-Euphrates valley. Hawthorn Books, New York, New York, xix + 562 pp.
  201. Saleh, K.I. (2007) First recorded of Tilapia zillii (Gervais, 1848), in natural water of Iraq (Tigris River). The First Scientific Conference of Agricultures College, University of Basra, Basra, pp. 26–27.
  202. Sauvage, H.-E. (1878) Note sur quelques poissons d’espèces nouvelles provenant des eaux douces de l’Indo-Chine. Bulletin de la Société philomathique de Paris, Série 7, 2, 233–242.
  203. Sayyadzadeh, G. & Esmaeili, H.R. (2020) Oxynoemacheilus marunensis, a new loach species from the Persian Gulf basin with remarks on O. frenatus (Teleostei: Nemacheilidae). Zootaxa, 4885 (2), 189–206. https://doi.org/10.11646/zootaxa.4885.2.2
  204. Sayyadzadeh, G., Esmaeili, H.R. & Freyhof, J. (2015) Garra mondica, a new species from the Mond River drainage with remarks on the genus Garra from the Persian Gulf basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4048 (1), 75–89. https://doi.org/10.11646/zootaxa.4048.1.4
  205. Sayyadzadeh, G., Eagderi, E. & Esmaeili, H.R. (2016) A new loach of the genus Oxynoemacheilus from the Tigris River drainage and its phylogenetic relationships among the nemacheilid fishes (Teleostei: Nemacheilidae) in the Middle East on mtDNA COI sequences. Iranian Journal of Ichthyology, 3 (4), 236–250.
  206. Sayyadzadeh, G., Esmaeili, H.R., Eagderi, S., Jouladeh-Roudbar, A., Masoudi, M. & Vatandoust S. (2017) Re-description of Oxynoemacheilus longipinnis from the Persian Gulf basin (Teleostei: Nemacheilidae). Zoology in the Middle East, 63 (3), 228–238. https://doi.org/10.1080/09397140.2017.1349243
  207. Smith, A. (1838–47) Pisces. In: Illustrations of the zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa in 1834–36, 4, 77 (unnumbered) pp., accompanying pls. 1–31.
  208. Spix, J.B. & von, Agassiz, L. (1829–31) n.k. In: Selecta genera et species piscium quos in itinere per Brasiliam annis MDCCCXVII–MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis augustissimi peracto. Collegit et pingendos curavit Dr J. B. de Spix, (Quondam ordinis Regii Coronae Civilis Bavaricae Eques, Academiae Scientiarum. Bavaricae Socius Ordinarius, Musei Regii zoologici, zootomici et ethnographici Conservator rel.). Digessit, descripsit et observationibus anatomicis illustravit Dr. L. Agassiz, praefatus est et edidit itineris socius Dr. F. C. Ph. de Martius. Parts 1 & 2. Typis C. Wolf, Monachii, pp. i–xvi + i–ii + 1–6 + 1–82, pls. 1–48 & pp. 83–138, pls. 49–101.
  209. Steindachner, F. (1864) Ichthyologische Mittheilungen. (VII.) [With subtitles I–VIII]. Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien, 14, 223–232, pls. 7–8.
  210. Tan, M. & Armbruster, J.W. (2018) Phylogenetic classification of extant genera of fishes of the order Cypriniformes (Teleostei: Ostariophysi). Zootaxa, 4476 (1), 6–39. https://doi.org/10.11646/zootaxa.4476.1.4
  211. Trewavas, E. (1955) A blind fish from Iraq, related to Garra. Annals and Magazine of Natural History, Series 12, 8 (91) (Art. 67), 551–555. https://doi.org/10.1080/00222935508655666
  212. Turan, D., Kaya, C., Ekmekçi, F.G. & Doğan, E. (2014) Three new species of Alburnoides (Teleostei: Cyprinidae) from the Euphrates River, Eastern Anatolia, Turkey. Zootaxa, 3754 (2), 101–116. https://doi.org/10.11646/zootaxa.3754.2.1
  213. Van der Laan, R., Eschmeyer, W.N. & Fricke, R. (2014) Family-group names of recent fishes. Zootaxa, 3882 (Monograph), 1–230. https://doi.org/10.11646/zootaxa.3882.1.1
  214. van Hasselt, J.C. (1823) Uittreksel uit een' brief van den Heer J. C. van Hasselt, aan den Heer C. J. Temminck, geschreven uit Tjecande, Residentie Bantam, den 28sten December 1822. Algemeene Konst- en Letter-bode voor het Jaar II Deel, 35, 130–133.
  215. Wagner, S.C. (2010) Keystone Species. Nature Education Knowledge, 3 (10), 51
  216. Walbaum, J.J. (1792) Petri Artedi sueci genera piscium. In quibus systema totum ichthyologiae proponitur cum classibus, ordinibus, generum characteribus, specierum differentiis, observationibus plurimis. Redactis speciebus 242 ad genera 52. Ichthyologiae Pars III. Ant. Ferdin. Rose, Grypeswaldiae [Greifswald], (viii) + 723 pp., pls. 1–3.
  217. World Wide Fund for Nature [WWF] (2016) Living planet report. Risk and resilience in a new era. WWW International, Gland. [unknown pagination]
  218. Yaseen, A.T. (2016) Effect of some environmental factors on the nature of fish assemblage in stream and estuary of Shatt Al-Arab. M. Sc. Thesis, College of Agriculture, Tikrit University, Tikrit, 132 pp.
  219. Zareian, H. & Esmaeili, H.R. (2017) Mitochondrial phylogeny and taxonomic status the Capoeta damascina species group (Actinopterygii: Cyprinidae) in Iran with description of a new species. Iranian Journal of Ichthyology, 4 (3), 231–269.
  220. Zareian, H., Esmaeili, H.R., Gholamhosseini, A., Japoshvili, B., Özuluğ, M. & Mayden, R.L. (2018) Diversity, mitochondrial phylogeny, and ichthyogeography of the Capoeta capoeta complex (Teleostei: Cyprinidae). Hydrobiologia, 806, 363–409. https://doi.org/10.1007/s10750-017-3375-0
  221. Zareian, H., Esmaeili, H.R., Zamanian Nejad, R. & Vatandoust, S. (2015) Hemiculter leucisculus (Basilewsky, 1855) and Alburnus caeruleus Heckel, 1843: new data on their distributions in Iran. Caspian Journal of Environmental Science, 13, 11–20.
  222. Ziyadi, M.S., Jawad, L.A., Almukhtar, M.A. & Pohl, T. (2015) Day's goby, Acentrogobius dayi Koumans, 1941 (Pisces: Gobiidae) in the desert Sawa Lake, south-west Baghdad, Iraq. Marine Biodiversity Records, 8, e148. https://doi.org/10.1017/S1755267215001220