Abstract
The current status of the freshwater fishes of Israel is revised, and an updated checklist is presented. The confirmed freshwater fishes of Israel comprise 55 species belonging to nine orders, 14 families, and 39 genera. Among these, 19 species (34.6%) are alien, and four species (7.3%) are considered endemic to Israel. The orders with the largest numbers of species in the ichthyofauna of Israel are Cypriniformes (19 species), followed by Cichliformes (14 species), Cyprinodontiformes (seven species), Mugiliformes (six species), Siluriformes (four species), Salmoniformes (two species), and the others represent one species in each. At the family level, the Cichlidae have the greatest number of species (14 species; 25.5% of the total species), followed by Cyprinidae (10 species), Leuciscidae and Mugilidae (six species in each), Aphaniidae (four species), Nemacheilidae and Poecilidae (three species in each). According to IUCN Red List criteria, among 36 naturally distributed species (alien species not included), two species are extinct (EX) and nine species (25.0%) are categorized as threatened extinctions, with two (5.6%) CR, four (11.1%) EN, and three (8.3%) VU.
References
- Abell, R., Thieme, M.L., Revenga, C., Bryer, M., Kottelat, M., Bogutskaya, N., Coad, B. Mandrak, N., Balderas, S.C., Bussing, W., Stiassny, M. L. J., Skelton, P., Allen, G.R., Unmack, P., Naseka, A., Ng, R., Sindorf, N., Robertson, J., Armijo, E., Higgins, J.V., Heibel, T.J., Wikramanayake, E., Olson, D., López, H.L., Reis, R.E., Lundberg, J.G., Pérez, M.H.S. & Petry, P. (2008) Freshwater Ecoregions of the World: A New Map of Biogeographic Units for Freshwater Biodiversity Conservation. BioScience, 58 (5), 403–414. https://doi.org/10.1641/B580507
- Ahl, E. (1926) Einige neue Fische der Familie Cichlidae aus dem Nyassa-See. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1926, 51–62.
- Baird, S.F. & Girard, C.F. (1853) Descriptions of new species of fishes collected by Mr. John H. Clark, on the U.S. and Mexican Boundary Survey, under Lt. Col. Jas. D. Graham. Proceedings of the Academy of Natural Sciences of Philadelphia, 6, 387–390.
- Bănărescu, P.M. & Nalbant, T.T. (1966) Zwei neue Schmerlen der Gattung Noemacheilus (Pisces, Cobitidae) aus Jordanien. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 63, 329–336, pl. 5.
- Bănărescu, P.M., Nalbant, T.T. & Goren, M. (1982) The nomacheiline loaches from Israel (Pisces: Cobitidae: Noemachilinae). Israel Journal of Zoology, 31, 1–25.
- Bariche, M. & Freyhof, J. (2016) Status of Pseudophoxinus libani and P. kervillei, two minnows from the Levant (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 27 (3), 203–210.
- Beckman, W.C. (1962) The Freshwater Fishes of Syria and their General Biology and Management. FAO, Rome, 297 pp.
- Ben-Tuvia, A. (1981) Men-induced changes in the freshwater fish fauna of Israel. Fishery Management, 12, 139–148. https://doi.org/10.1111/j.1365-2109.1981.tb00023.x
- Berg, L.S. (1948) Ryby presnych vod SSSR i sopredelnych stan. [Freshwater fishes of the U.S. S. R. and adjacent countries]. Vol. 1. 4th Edition. Opredeliteli po faune SSSR. [Guide to the fauna of the U.S.S.R.]. Freshwater fishes of the U.S.S.R. and adjacent countries No. 27. Academy of Sciences of the U.S.S.R., Zoological Institute, Moskva, 466 pp. [in Russian, English translation appeared in Israel Program of Scientific Translation, Jerusalem, 1962, 504 pp.]
- Berg, L.S. (1949) Ryby presnych vod SSSR i sopredelnych stan. [Freshwater fishes of the U.S.S.R. and adjacent countries]. Vol. 2. 4th Edition, Opredeliteli po faune SSSR. [Guide to the Fauna of the U.S.S.R.]. Freshwater fishes of the U.S.S.R. and adjacent countries No. 29. Academy of Sciences of the U.S.S.R., Zoological Institute, Moskva, 467–925. [in Russian. English translation appeared in Israel Program of Scientific Translation, Jerusalem, 1964, 496 pp.]
- Blackwell, T., Ford, A.G.P., Ciezarek, A.G., Bradbeer, S.J., Juarez, C.A.G., Smith, A.M., Ngatunga, B.P., Shechonge, A., Tamatamah, R., Etherington, G., Haerty, W., Di Palma, F., Turner, G.F. & Genner, M.J. (2021) Newly discovered cichlid fish biodiversity threatened by hybridization with non-native species. Molecular Ecology, 30 (4), 895–911. https://doi.org/10.1111/mec.15638
- Borkenhagen, K. & Freyhof, J. (2009) New records of the Levantine endemic cichlid Tristramella simonis from Syria. Cybium, 33 (4), 335–336.
- Borkenhagen, K. & Krupp, F. (2013) Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae). ZooKeys, 339, 1–53. https://doi.org/10.3897/zookeys.339.4903
- Boulenger, G.A. (1907) Zoology of Egypt: The fishes of the Nile. Hugh Rees, Ltd., London, li + 578 pp., 97 pls. https://doi.org/10.5962/bhl.title.51710
- Boulenger, G.A. (1908) Diagnoses of new fishes discovered by Capt. E. L. Rhoades in Lake Nyassa. Annals and Magazine of Natural History, Series 8, 2 (9), 238–243. https://doi.org/10.1080/00222930808692478
- Boulenger, G.A. (1912) Descriptions of three new African cichlid fishes of the genus Tilapia, preserved in the British Museum. Annals and Magazine of Natural History, Series 8, 10 (55), 138–140. https://doi.org/10.1080/00222931208693206
- Burchell, W.J. (1822) Travels in the interior of southern Africa. 2 Vols. London, xi + 582 + 4 pp., 1 map & 648 pp. https://doi.org/10.5962/bhl.title.100911
- Burgess, W.E. (1977) Studies on the family Cichlidae: 8. Pseudotropheus lombardoi, a new species of Lake Malawi Mbuna with reversed sexual coloration (Pisces: Cichlidae). Tropical Fish Hobbyist, 26 (2), 63–67.
- Cambray, J.A. (2003) Impact on indigenous species biodiversity caused by the globalisation of alien recreational freshwater fishes. Hydrobiologia, 500, 217–230. https://doi.org/10.1023/A:1024648719995
- Carpenter, K.E., Krupp, F., Jones, D.A. & Zajonz, U. (1997) FAO species identification guide for fishery purposes. The living marine resources of Kuwait, eastern Saudi Arabia, Bahrain, Qatar, and the United Arab Emirates. FAO, Rome, 293 pp., 17 pls.
- Castelnau, F.L. (1855) Poissons. In: Animaux nouveaux or rares recueillis pendant l’expédition dans les parties centrales de l’Amérique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; exécutée par ordre du gouvernement Français pendant les années 1843 a 1847. Vol. 2. Part 7. Zoologie. P. Bertrand, Paris, xii + 112 pp., 50 pls. https://doi.org/10.5962/bhl.title.110120
- Chervinski, J. (1977) Adaption of Chalon labrosus (Risso) and Liza saliens (Risso) (Pisces, Mugilidae) to fresh water. Aquaculture, 11, 75–79. https://doi.org/10.1016/0044-8486(77)90155-7
- Çiçek, E., Eagderi, S. & Sungur, S. (2022) A review of the alien fishes of Turkish inland waters. Turkish Journal of Zoology, 46, 1–13. https://doi.org/10.55730/1300-0179.3104
- Çiçek, E., Sungur, S. & Fricke, R. (2020) Freshwater lampreys and fishes of Turkey; a revised and updated annotated checklist 2020. Zootaxa, 4809 (2), 241–270.
- Cucherousset, J. & Olden, J.D. (2011) Ecological impacts of non-native freshwater fishes. Fisheries, 36, 215–230. https://doi.org/10.1080/03632415.2011.574578
- Cuvier, G. (1829) Le Règne Animal, distribué d’après son organisation, pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Vol. 2. 2nd Edition. Fortin, Masson et Cie, Paris, xv + 406 pp. https://doi.org/10.5962/bhl.title.1964
- Cuvier, G. & Valenciennes, A. (1842) Histoire naturelle des poissons. Tome Seizième. Livre dix-huitième. Les Cyprinoïdes. Vol. 16. P. Bertrand, Paris, xx + 472 pp., pls. 456–487.
- Diogo, R., Chardon M. & Vandewalle, P. (2003) On the osteology and myology of the cephalic region and pectoral girdle of Heteropneustes fossilis (Siluriformes: Heteropneustidae), with commens on the phylogenetic relationships between Heteropneustes and the clariid catfishes. Animal Biology, 53 (4), 379–396. https://doi.org/10.1163/157075603322556283
- Doadrio, I. & Carmona, J.A. (2006) Phylogenetic overview of the genus Squalius (Actinopterygii, Cyprinidae) in the Iberian Peninsula, with description of two new species. Cybium, 30 (3), 199–214.
- Dor, M. (1987) Zoological Lexicon, Vertebrata. 2 Vols. Dvir Publications, Tel Aviv, 807 pp. [in Hebrew]
- Dunz, A.R. & Schliewen, U.K. (2012) Description of a rheophilic Tilapia species Smith, 1840 (Teleostei: Cichlidae) from Guinea with comments on Tilapia rheophila Daget, 1962. Zootaxa, 3314 (1), 17–30. https://doi.org/10.11646/zootaxa.3314.1.2
- Duquenne-Delobel, E., Doadrio, I. & Denys, G.P.J. (2022) Revalidation of the genus Ichthyocoris Bonaparte, 1840 (Actinopterygii: Blenniiformes: Blenniidae). Acta Ichthyologica et Piscatoria, 52 (1), 35–41. https://doi.org/10.3897/aiep.52.79458
- Durand, J.-D. & Borsa, P. (2015) Mitochondrial phylogeny of grey mullets (Acanthopterygii: Mugilidae) suggests high proportion of cryptic species. Comptes Rendus Biologies, 338 (4), 266–277. https://doi.org/10.3897/aiep.52.79458
- Durand, J.-D., Chen, W.-J. Shen, K.-N., Fu, C.-Z. & Borsa, P. (2012) Genus-level taxonomic changes implied by the mitochondrial phylogeny of grey mullets (Teleostei: Mugilidae). Comptes Rendus Biologies, 335 (10–11), 687–697. https://doi.org/10.1016/j.crvi.2012.09.005
- Ege, V. (1939) A revision of the genus Anguilla Shaw. A systematic, phylogenetic and geographical study. Dana Report, No. 16, 1–257, pls. 1–6. https://doi.org/10.1163/9789004629585
- Elvira, B. (2001) Identification of non-native freshwater fishes established in Europe and assessment of their potential threats to the biological diversity. Convention on the conservation of European wildlife and natural habitats. Council of Europe. Standing Committee 21st Meeting, Strasbourg, 26–30 November 2001, T-PVS, 6, 1–35.
- Esmaeili, H.R., Teimori, A., Zarei, F. & Sayyadzadeh, G. (2020) DNA barcoding and species delimitation of the old world tooth-carps, family Aphaniidae Hoedeman, 1949 (Teleostei: Cyprinodontiformes). PLoS ONE, 15 (4), e0231717, 1–26. https://doi.org/10.1371/journal.pone.0231717
- Fishelson, L. (1962) Hybrid of two species of fishes of the genus Tilapia (Cichlidae, Teleostei). Fishermen’s Bulletin, Haifa, 4 (2), (32), 14–9. [in Hebrew]
- Freyhof, J. & Özuluğ, M. (2014) Acanthobrama thisbeae, a new species of bream from southern Anatolia, Turkey (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 25 (1), 1–10.
- Freyhof, J. & Yoğurtçuoğlu, B. (2020) A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa, 4810 (3), 421–451. https://doi.org/10.11646/zootaxa.4810.3.2
- Freyhof, J. (2016) Redescription of Garra elegans (Günther, 1868), a poorly known species from the Tigris River drainage (Teleostei: Cyprinidae). Zootaxa, 4173 (5), 496–500. https://doi.org/10.11646/zootaxa.4173.5.7
- Freyhof, J., Els, J., Feulner, G.R., Hamidan, N.A. & Krupp, F. (2020) Freshwater fishes of the Arabian Peninsula. Motivate Media Group, Dubai, 272 pp.
- Freyhof, J., Erk’akan, F., Özeren, S.C. & Perdices, A. (2012) An overview of the western Palaearctic loach genus Oxynoemacheilus (Teleosei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 22 (4), 301–312.
- Fricke, R., Eschmeyer, W.N. & Van der Laan, R. (Eds.) (2023) Eschmeyer‘s catalog of fishes: genera, species, references. Electronic Version. Available from: http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (accessed 30 April 2023)
- Froese, R. & Pauly, D. (Eds.) (2022) FishBase. World Wide Web Electronic Publication. Version August 2022. Available from: http://www.fishbase.org (accessed 6 February 2023)
- Fryer, G. (1956) New species of cichlid fishes from Lake Nyasa. Revue de Zoologie et de Botanique Africaines, 53 (1–2), 81–91.
- Garcia-Berthou, E. & Moyle, P.B. (2011) Rivers. In: Simberloff, D. & Rejmanek, M. (Eds.), Encyclopaedia of Biological Invasions. University of California Press, Berkeley and Los Angeles, California, pp. 609–612. https://doi.org/10.1525/9780520948433-136
- Geiger, M.F., Herder, F., Monaghan, M.T., Almada, V., Barbieri, R., Bariche, M., Berrebi, P., Bohlen, J., Casal-Lopez, M., Delmastro, G.B., Denys, G.P.J., Dettai, A., Doadrio, I., Kalogianni, E., Kärst, H., Kottelat, M., Kovacic, M., Laporte, M., Lorenzoni, M., Marcic, Z., Özulug, M., Perdices, A., Perea, S., Persat, H., Porcelotti, S., Puzzi, C., Robalo, J., Šanda, R., Schneider, M., Šlechtová, V., Stoumboudi, M., Walter, S. & Freyhof, J. (2014) Spatial heterogeneity in the Mediterranean biodiversity hotspot affects barcoding accuracy of its freshwater fishes. Molecular Ecology Resources, 14, 1210–1221. https://doi.org/10.1111/1755-0998.12257
- Gervais, F.L.P. (1848) Sur les animaux vertébrés de l’Algérie, envisagés sous le double rapport de la géographie zoologique et de la domestication. Annales des Sciences Naturelles, Paris, Série 3, Zoologie, 10, 202–208. https://doi.org/10.5962/bhl.title.51497
- Givati, A., Fredj, E. & Silver, M. (2016) Chapter 6—Operational Flood Forecasting in Israel. In Flood Forecasting. A Global Perspective. Thomas E.A., III & Thomas, C. Academic Press, pp. 153–167. https://doi.org/10.1016/b978-0-12-801884-2.00006-2
- Godwin, J.C., Stehen, D.A., Werneke, D. & Armbruster, J.W. (2016) Two significant records of exotic tropical freshwater fishes in Southern Alabama. Notes of the Southeastern Naturalist, 15 (4), 57–60. https://doi.org/10.1656/058.015.0401
- Golani, D., Sonin, O., Snovsky, G., David, L. & Tadmor-Levi, R. (2019) The occurrence of the peacock bass (Cichla kelberi Kullander and Ferreira, 2006) in Lake Kinneret (Sea of Galilee), Israel. BioInvasions Records, 8 (3), 706–711. https://doi.org/10.3391/bir.2019.8.3.28
- Golani, D. & Lerner, A. (2007) A long term study of the sandy shore ichthyofauna in the northern Red Sea (Gulf of Aqaba), with reference to adjacent mariculture activity. Raffles Bulletin of Zoology Supplement, 14, 255–264.
- Golani, D. & Mires, D. (2000) Introduction of fishes to the freshwater system of Israel. Israel Journal of Aquaculture—Bamidgeh, 52, 47–60.
- Golani, D. & Shefler, D. (1985) The occurrence of Rainbow Trout (Salmo gairdneri) and Coho Salmon (Oncorhynchus kisutch) in Lake Kinneret (Lake Tiberias). Bamidgeh, 37, 27–31.
- Golani, D. & Snovsky, G. (2013) Occurrence of suckermouth armored catfish (Siluriformes, Loricariidae, Pterygoplichthys) in inland waters of Israel. BioInvasions Records, 2, 253–256. https://doi.org/10.3391/bir.2013.2.3.13
- Golani, D., Shefler, D. & Gelman, A. (1988) Aspects of growth and feeding habits of adult European eel (Anguilla anguilla) in Lake Kinneret (Lake Tiberias), Israel. Aquaculture, 74, 349–354. https://doi.org/10.1016/0044-8486(88)90378-X
- Golani, D., Sonin, O. & Rubinstein, G. (2015) Records of Paralichthys lethostigma and Sciaenops ocellatus in the Mediterranean and Channa micropeltes in Lake Kinneret (Sea of Galilee), Israel. Marine Biodiversity Records, 8, e39. https://doi.org/10.1017/S1755267215000081
- Golani, D., Shohat, H. & Appelbaum-Golani, B. (2022) Colonization of exotic fish species of the genera Pseudotropheus and Aulonocara (Perciformes: Cichlidae) and decline of native ichthyofauna in Nahal Amal, Israel. Annales Series Historia Naturalis, 32 (2), 294–300.
- Golani, D., Sonin, O., Snovsky, G., David, L. & Tadmor-Levi, R. (2019) The Occurrence of the peacock bass (Cichla kelberi Kullander and Ferreira, 2006) in Lake Kinneret (Sea of Galilee), Israel. BioInvasions Records, 8 (3), 706–711. https://doi.org/10.3391/bir.2019.8.3.28
- Goren, M. (1972) The populations of Pseudophoxinus zeregi (Heckel) in Israel and Syria and the status of Pseudophoxinus (Pararhodeus) drusensis (Pellegrin) (Pisces: Cyprinidae). Zoological Journal of the Linnean Society, 51, 141–145. https://doi.org/10.1111/j.1096-3642.1972.tb00778.x
- Goren, M. (1974) The freshwater fishes of Israel. Israel Journal of Zoology, 23 (2), 67–118, pls. 1–4.
- Goren, M., Fishelson, L. & Trewavas, E. (1973) The Cyprinid fishes of Acanthobrama Heckel and related genera. Bulletin of the British Museum (Natural History), Zoology, 24, 291–315.
- Goren, M. & Ortal, R. (1999) Biogeography, diversity and conservation of the inland water communities in Israel. Biological Conservation, 89, 1–9. https://doi.org/10.1016/S0006-3207(98)00127-X
- Günther, A. (1864) Report on a collection of fishes from Palestine. Proceedings of the Zoological Society, London, 1864, 488–493.
- Günther, A. (1865) Report on a collection of reptiles and fishes from Palestine. Proceedings of the Zoological Society of London, 1864 (3), 488–493.
- Günther, A. (1867) On the fishes of the states of Central America, founded upon specimens collected in fresh and marine waters of various parts of that country by Messrs. Salvin and Godman and Capt. J. M. Dow. Proceedings of the Zoological Society of London, 1866 (3), 600–604.
- Guo, Y.-S. (2021) Colored atlas of fishes of Sichuan. Vol. I. Colored atlas of fishes of Sichuan. Science press, Beijing, 475 pp.
- Hasselquist, F. (1757) Fredric Hasselquists M.D., Societ. Reg. Scient. Upsal. & Stockholm Soc., Iter Palæstinum eller resa til heliga landet, förråttad ifrån år 1749 til 1752, med besfrikningar, rön, anmärkningar öfverde märkvårdigaste naturalier, på Hennes Kongl. Maj:ts befallning, utgiven af Carolus Linnæus. Lars Salvius, Stockholm. Iter Palæstinum eller resa til heliga landet, förråttad ifrån år 1749 til 1752. Lars Salvius, Stockholm, xv + 619 + i pp. https://doi.org/10.5962/bhl.title.112563
- Heckel, J.J. (1843a) Ichthyologie [von Syrien]. In: von Russegger, J., Reisen in Europa, Asien und Afrika, mit besonderer Rücksicht auf die naturwissenschaftlichen Verhältnisse der betreffenden Länder unternommen in den Jahren 1835 bis 1841, etc. Ichthyologie [von Syrien]. Vol. 1. Pt 2. E. Schweizerbart‘sche Verlagshandlung, Stuttgart, pp. 991–1099.
- Heckel, J.J. (1843b) Abbildungen und Beschreibungen der Fische Syriens, nebst einer neuen Classification und Characteristik sämmtlicher Gattungen der Cyprinen. In: Fenzl, E., Heckel, J.J. & Redtenbenher, L. (Eds.), Abbildungen und Beschreibungen neuer und seletner Thiere und Pflanzen in Syrien und im westlichen Taurus gesammelt von Th. Kotschy. Vol. 1. Pt 2. E. Schweizerbart‘sche Verlagshandlung, Stuttgart, pp. 991–1044, pls. 1–13. https://doi.org/10.5962/bhl.title.14055
- Heckel, J.J. (1847) Naturhistorischer Anhang. [Various subtitles]. In: Russegger, J. Von, Reisen in Europa, Asien und Afrika,....unternommen in den Jahren 1835 bis 1841. 2 (3). E. Schweizerbart‘sche Verlagshandlung, Stuttgart, pp. 207–357.
- Heckel, J.J. (1848) Eine neue Gattung von Poecilien mit rochenartigem Anklammerungs-Organe. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe, 1 (1–5), 289–303. [1848]
- Hornell, J. (1935) Reporton the Fisheries of Palestine. Crown Agent for the Colonies. His Majesty’s Stationary Office, London. 106 pp.
- Huber, J.H. (2019) A nomenclatural and systematic analysis of livebearing Cyprinodontiformes (Acanthopterygii: Anablepsinae, Goodeinae, Poeciliidae). Killi-Data Series, 2019, 4–155.
- Humboldt, F.H.A. von. & Valenciennes, A. (1821) Recherches sur les poissons fluviatiles de l’Amérique Équinoxiale. In: Voyage de Humboldt et Bonpland, Deuxième partie. Observations de Zoologie et d’Anatomie comparée, Paris, 2, 145–216, pls. 45–52. [title page 1833]
- Isbrücker, I.J.H. (1980) Classification and catalogue of the mailed Loricariidae (Pisces, Siluriformes). Verslagen en Technische Gegevens, Instituut voor Taxonomische Zoöogie, Universiteit van Amsterdam, No. 22, 1–181.
- IUCN. (2023) The IUCN Red List of Threatened Species Version 2022-2 Online. Available from: https://www.iucnredlist.org/ (accessed 12 July 2023)
- Jawad, L.A., Jassim, A.K., Tahir Ankush, M.A.W. & Abed, J.M. (2019) Flushed with the flood: the rainbow trout Oncorhynchus mykiss in the Shatt Al-Arab River, Basrah, Iraq. Thalassia Salentina, 41, 119–126.
- Kosswig, C. (1950) Die Gattung Tylognathus in Vorderasien. In: Ergänzungband „Neue Ergebnisse und Probleme der Zoologie“, Festschrift Berthold Klatt. Zoologischer Anzeiger, 145, 406–415.
- Kottelat, M. & Freyhof, J. (2007) Handbook of European Freshwater Fishes. Publications Kottelat, Cornol, xiii + 646 pp.
- Krupp, F. (1985) Systematik und Zoogeographie der Süsswasserfische des levantinischen Grabenbruch-Systems und der Ostküste des Mittelmeers. Dissertation, Johannes Gutenberg Universität, Mainz, 215 pp.
- Krupp, F. & Schneider, W. (1989) The fishes of the Jordan River drainage basin and Azraq Oasis. Fauna of Saudi Arabia, 10, 347–416.
- Krupp, F. (1982) Garra tibanica ghorensis subsp. nov. (Pisces: Cyprinidae), an African element in the cyprinid fauna of the Levant. Hydrobiologia, 88 (3), 319–324. https://doi.org/10.1007/BF00008513
- Kullander, S.O. & Ferreira, E.J.G. (2006) A review of the South American cichlid genus Cichla, with descriptions of nine new species (Teleostei: Cichlidae). Ichthyological Exploration of Freshwaters, 17 (4), 289–398.
- Lee, D.S., Gilbert, C.R., Hocutt, C.H., Jenkins, R.E., McAllister, D.E. & Stauffer, Jr. J.R. (1980) Atlas of North American freshwater fishes. Publication of the North Carolina Biological Survey, No. 1980-12, i–x + 1–86. https://doi.org/10.5962/bhl.title.141711
- Levanter, H. (1984) Biological control of reservoirs by fish. 2nd Edition. Mekorot Water Co., Tel Aviv, 51 pp.
- Lévêque, C., Paugy, D. & Teugels, G.G. (Eds.) (1992) Faune des poissons d’eaux douces et saumâtres de l’Afrique de l’Ouest. Collection Fauna tropicale, No. XXVIII, 2, 389–902.
- Levin, B.A., Freyhof, J., Lajbner, Z., Perea, S., Abdoli, A., Gaffaroğlu, M., Özuluğ, M., Rubenyan, H.R., Salnikov, V.B. & Doadrio, I. (2012) Phylogenetic relationships of the algae scraping cyprinid genus Capoeta (Teleostei: Cyprinidae). Molecular Phylogenetics and Evolution, 62, 542–549. https://doi.org/10.1016/j.ympev.2011.09.004
- Linnaeus, C. (1758) Systema Naturae. Ed. X. (Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata). Impensis Direct. Laurentii Salvii, Holmiae, ii + 824 pp. https://doi.org/10.5962/bhl.title.542
- Lortet, L. (1883) Études zoologiques sur la faune du lac de Tibériade, suivies d’un aperçu sur la faune des lacs d’Antioche et de Homs. I. Poissons et reptiles du lac de Tibériade et de quelques autres parties de la Syrie. Archives du Muséum d’Histoire Naturelle de Lyon, 3, 99–189, pls. 6–18. https://doi.org/10.3406/mhnly.1883.941
- Maldonado-Ocampo, J.A., Vari, R.P. & Usma, J.S. (2008) Checklist of the freshwater fishes of Colombia. Biota Colombiana, 9, 143–237.
- McEachran, J.D. & Fechhelm J.D. (1998) Fishes of the Gulf of Mexico. Vol. 1. Myxiniformes to Gasterosteiformes. University of Texas Press, Austin, Texas, 1112 pp. https://doi.org/10.7560/752061
- Miller, R.R. (2006) Freshwater fishes of México. University of Chicago Press, Chicago, Illinois, xxvi + 490 pp.
- Nakamura, I., Inada, T., Takeda, M. & Hatanaka, H. (1986) Important fishes trawled off Patagonia. Japan Marine Fishery Resource Research Center, Tokyo, 369 pp.
- Nelson, J.S., Grande, T.C. & Wilson, M.V.H. (2016) Fishes of the World. 5th Edition. John Wiley & Sons, Inc, Hoboken, New Jersey, 707 pp.
- Pan, J.-H., Zhong, L., Zheng, C.-Y., Wu, H.-L. & Liu, J.-H. (Eds.) (1991) The freshwater fishes of Guangdong Province. Guangdong Science and Technology Press, Guangzhou, frontmatter, 17 + 589 pp.
- Pellegrin, J. (1911) Poissons de Syrie recueillis par H. Gadeau de Kerville. Bulletin de la Société Zoologique de France, 36, 107–111.
- Pellegrin, J. (1933) Description d‘un poisson nouveau de la Syrie méridionale appartenant au genre Phoxinellus. Bulletin du Muséum National d‘Histoire Naturelle, Série 2, 5 (5), 368–369.
- Peters, W.(C.H.) (1852) Diagnosen von neuen Flussfischen aus Mossambique. Bericht über die zur Bekanntmachung geeigneten Verhandlungen der Königlichen Preussischen Akademie der Wissenschaften zu Berlin, 1852, 275–276 + 681–685.
- Pietschmann, V. (1913) Eine neue Glyptosternum-Art aus dem Tigris. Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftlichen Klasse, 50 (8), 93–95.
- Prokofiev, A.M. (2010) Morphological classification of loaches (Nemacheilinae). Journal of Ichthyology, 50 (10), 827–913. https://doi.org/10.1134/S0032945210100012
- Prokofiev, A.M. (2009) Problems of the classification and phylogeny of Nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49 (10), 874–898. https://doi.org/10.1134/S0032945209100051
- Prokofiev, A.M. (2017) Loaches of the subfamily Nemacheilinae of the World. Filigran, Yaroslavl, 315 pp.
- Regan, C.T. (1914) Description of a new cyprinodont fish of the genus Mollienisia from Yucatan. Annals and Magazine of Natural History, Series 8, 13 (75), 338. https://doi.org/10.1080/00222931408693488
- Reich, K. (1978) Lake Kinneret in its development. Bamidgeh, 37, 37–64.
- Richardson, J. (1856) On some fishes from Asia Minor and Palestine. Proceedings of the Zoological Society, London, 24, 371–377.
- Risso, A. (1810) Ichthyologie de Nice, ou histoire naturelle des poissons du Département des Alpes Maritimes. F. Schoell, Paris, xxxvi + 388 pp., 11 pls. https://doi.org/10.5962/bhl.title.7052
- Risso, A. (1827) Histoire naturelle des principales productions de l’Europe méridionale, et particulièrement de celles des environs de Nice et des Alpes maritimes. Vol. 3. F. G. Levrault, Paris and Strasbourg, xvi + 480 pp., 16 pls. https://doi.org/10.5962/bhl.title.58984
- Roberts, T.R. & Vidthayanon, C. (1991) Systematic revision of the Asian catfish family Pangasiidae, with biological observations and descriptions of three new species. Proceedings of the Academy of Natural Sciences of Philadelphia, 143, 97–143.
- Roll, U., Dayan, T. & Simberloff, D. (2007) Non-indigenous insect species in Israel and adjacent areas. Biological Invasions, 9, 629–643. https://doi.org/10.1007/s10530-006-9064-y
- Rosen, D.E. & Bailey, R.M. (1963) The poeciliid fishes (Cyprinodontiformes), their structure, zoogeography, and systematics. Bulletin of the American Museum of Natural History, 126 (1), 1–176.
- Saad, A., Çiçek, E., Esmaeili, H.R., Fricke, R., Sungur, S. & Eagderi, S. (2023) Freshwater fishes of Syria: a revised and updated annotated checklist-2023. Zootaxa, 5350 (1), 1–62. https://doi.org/10.11646/zootaxa.5350.1.1
- Sauvage, H.-E. (1878) Note sur quelques poissons d’espèces nouvelles provenant des eaux douces de l’Indo-Chine. Bulletin de la Société philomathique de Paris, Série 7, 2, 233–242.
- Schmitter-Soto, J.J. (2007) A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species. Zootaxa, 1603 (1), 1–76. https://doi.org/10.11646/zootaxa.1603.1.1
- Selz, O.M. & Seehausen, O. (2019) Interspecific hybridization can generate functional novelty in cichlid fish. Proceedings of the Royal Society B: Biological Sceince, 286, 20191621. https://doi.org/10.1098/rspb.2019.1621
- Shechonge, A., Ngatunga, B.P., Tamatamah, R., Bradbeer, S.J., Harrington, J., Ford, A.G.P., Turner, G.F. & Genner, M.J. (2018) Losing cichlid fish biodiversity: genetic and morphological homogenization of tilapia following colonization by introduced species. Conservation Genetics, 19, 1199–1209. https://doi.org/10.1007/s10592-018-1088-1
- Shefler, D. & Ben-Tuvia, A. (1982) The occurrence of Gibel-carp goldfish, Carassius auratus gibelio in Lake Kinneret (Lake Tiberias). Bamidgeh, 34 (2), 63–65.
- Skelton, P.H. & Teugels, G.G. (1992) Neotype description for the African catfish Clarias gariepinus (Burchell, 1822) (Pisces: Siluroidei, Clariidae). Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology, 56, 1–7.
- Snovsky, G. & Golani, D. (2012) The occurrence of an aquarium escapee, Pangasius hypophthalmus (Sauvage, 1878), (Osteichthys, Siluriformes, Pangasiidae) in Lake Kinneret (Sea of Galilee), Israel. BioInvasions Records, 7 (1), 101–103. https://doi.org/10.3391/bir.2012.1.2.03
- Spicer, I.J. (1931) Fisheries. In: Report of the Department of Agriculture and Forests for the Years 1927–30. Printing Office, Russian Building, Jerusalem, pp. 159–160.
- Stearley, R.F. & Smith, G.R. (1993) Phylogeny of the Pacific trouts and salmons (Oncorhynchus) and genera of the family Salmonidae. Transactions of the American Fisheries Society, 122 (1), 1–33. https://doi.org/10.1577/1548-8659(1993)122%3C0001:POTPTA%3E2.3.CO;2
- Steindachner, F. (1864) Ichthyologische Mittheilungen. (VII.) [With subtitles I–VIII]. Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien, 14, 223–232, pls. 7–8.
- Steinitz, H. (1953) The freshwater fishes of Palestine. An annotated list. Bulletine of the Research Council of Israel, 3B, 207–227.
- Steinitz, H. (1952) Acanthobrama terrae-sanctae, sp.n., from Lake Tiberias, Israel. Annals and Magazine of Natural History, Series 12, 5 (51), Art. 33, 293–298. https://doi.org/10.1080/00222935208654294
- Tadmor-Levi, R., Borovski, T., MarcosHadad, E., Shapiro, J., Hulata, G., Golani, D. & David, L. (2022) Establishing and using a genetic database for resolving identification of fish species in the Sea of Galilee, Israel. PLoS ONE, 17 (5), e0267021. https://doi.org/10.1371/journal.pone.0267021
- Tal, S. & Shelubsky, M. (1951) Review of the fish fatming industry in Israel. Transaction of the American Fisheries Society, 81, 218–223. https://doi.org/10.1577/1548-8659(1951)81[218:ROTFFI]2.0.CO;2
- Tan, M. & Armbruster, J.W. (2018) Phylogenetic classification of extant genera of fishes of the order Cypriniformes (Teleostei: Ostariophysi). Zootaxa, 4476 (1), 6–39. https://doi.org/10.11646/zootaxa.4476.1.4
- Teimori, A., Esmaeili, H.R., Hamidan, N. & Reichenbacher, B. (2018) Systematics and historical biogeography of the Aphanius dispar species group (Teleostei: Aphaniidae) and description of a new species from Southern Iran. Journal of Zoological Systematics and Evolutionary Research, 56, 579–598. https://doi.org/10.1111/jzs.12228
- Teugels, G.G. (1982) Preliminary results of a morphological study of five African species of the subgenus Clarias (Clarias) (Pisces; Clariidae). Journal of Natural History, 16 (3), 439–464. https://doi.org/10.1080/00222938200770351
- Tortonese, E. (1952) On a new cyprinoid fish of the genus Acanthobrama from Palestine. Annals and Magazine of Natural History, Series 12, 5 (51) (art. 29), 271–272. https://doi.org/10.1080/00222935208654290
- Tortonese, E. (1938) Viaggio del dott. Enrico Festa in Palestina e in Siria (1893). Pesci. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino, Series 3, 46 (85), 313–358.
- Trewavas, E. (1942) The cichlid fishes of Syria and Palestine. Annals and Magazine of Natural History, Series 11, 9 (55), 526–536. https://doi.org/10.1080/03745481.1942.9755503
- Trewavas, E. (1965) Tilapia aurea (Steindachner) and the status of Tilapia nilotica exul, T. monodi and T. lemassoni. Israel Journal of Zoology, 14, 258–276.
- Trewavas, E. (1982) Tilapias: taxonomy and speciation. In: Pullin, R.S.V. & Lowe-McConnell, R.H. (Eds.), The biology and culture of tilapias. ICLARM Conference Proceedings. No. 7. International Center for Living Aquatic Resources Management, Manila, pp. 3–13.
- Tristram, H.B. (1884) Fauna and Flora of Palestine. The Survey of Western Palestine. London, xxii + 455 pp., xx pls.
- van der Laan, R., Fricke, R. & Eschmeyer, W.N. (Eds.) (2023) Eschmeyer’s Catalog of Fishes: Classification. Online. Available from: http://www.calacademy.org/scientists/catalog-of-fishes-classification/ (accessed 14 April 2023)
- Wagner, S.C. (2010) Keystone Species. Nature Education Knowledge, 3 (10), 51.
- Walbaum, J.J. (1792) Petri Artedi sueci genera piscium. In quibus systema totum ichthyologiae proponitur cum classibus, ordinibus, generum characteribus, specierum differentiis, observationibus plurimis. Redactis speciebus 242 ad genera 52. Ichthyologiae pars III. Ant. Ferdin. Rose, Grypeswaldiae [Greifswald], (viii) + 723 pp., pls. 1–3.
- Weber, C. (1991) Nouveaux taxa dans Pterygoplichthys sensu lato (Pisces, Siluriformes, Loricariidae). Revue Suisse de Zoologie, 98 (3), 637–643. https://doi.org/10.5962/bhl.part.82074
- Weinberger, G., Livshitz, Y., Givati, A., Zilberbrand, M., Tal, A., Weiss, M. & Zurieli, A. (2012) The Natural Water Resources Between the Mediterranean Sea and the Jordan River. Israel Hydrological Service, Jerusalem, 63 pp.
- Whitehead, P.J.P., Bauchot, M.-L., Hureau, J.-C., Nielsen, J.G. & Tortonese, E. (1986) n.k. In: Fishes of the North-eastern Atlantic and the Mediterranean. Vol. III. UNESCO, Paris, pp. 1015–1473. https://doi.org/10.2307/1444931
- Yashouv, A. (1969) The Bass Disentrarchus punctatus (Bloch); a potential fish for culture in fresh and brakish water. Fishery and Fishbreeding in Israel, 4, 27–29. [in Hebrew]
- Yashouv, A. & Brener, A. (1961) Report on the Research in the Lake of Kinneret. Fishermen’s Bulletin, 28 (8), 18–24. [in Hebrew]
- Yoğurtçuoğlu, B., Kaya, C., Atalay, M.A., Ekmekçi, F.G. & Freyhof, J. (2023) Two new freshwater blennies from the Eastern Mediterranean basin (Teleostei: Blenniidae). Zootaxa, 5311 (1), 85–104. https://doi.org/10.11646/zootaxa.5311.1.4