Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5397 No. 4: 8 Jan. 2024
Type: Article
Published: 2024-01-08
DOI: 10.11646/zootaxa.5397.4.4
Page range: 521-538
Abstract views: 246
PDF downloaded: 8

Contributions to the tribes Othiini and Xantholinini (Coleoptera: Staphylinidae: Staphylininae) of Türkiye from the Aegean Region

Hacettepe University; Faculty of Science; Department of Biology; Applied Biology Section; Beytepe Çankaya Ankara; 06800; Türkiye
Hacettepe University; Faculty of Science; Department of Biology; Applied Biology Section; Beytepe Çankaya Ankara; 06800; Türkiye
Hacettepe University; Faculty of Science; Department of Biology; Applied Biology Section; Beytepe Çankaya Ankara; 06800; Türkiye
Coleoptera New species Staphylininae Faunistic Biodiversity Türkiye Aegean Region

Abstract

The main object of this study is to make contributions to the Staphylininae (Coleoptera: Staphylinidae) fauna of Türkiye with additional records from tribes Othiini and Xantholinini and a new species of Xantholinus Dejean, 1821. Examined material was composed of specimens collected from the Aegean Region between 2019–2022. A total of 18 species were recorded, belonging to nine genera and two tribes. Three of these are new records for Aegean Region. Furthermore, a new species, Xantholinus (Calolinus) manisaensis sp. n., is described from Manisa province. Description of the new species, photographs of habitus, genital segment and aedeagus as well as a differential diagnosis are also provided. The global distributions of the above species are included, and the zoogeographic status of each species is discussed. Previous records from the study area are compared to and evaluated with our findings. The majority of the collected species were European and Asian in distribution, reflecting the regional character and geographical position of Türkiye. It can be concluded that climate change has negatively affected biodiversity and with species loss as a possible result.

 

References

  1. Abacıgil, T.Ö., Varlı, S.V. & Tezcan, S. (2013) Faunistic studies on Staphylininae (Coleoptera: Staphylinidae) in Kazdağları (Balıkesir province) in Turkey. Munis Entomology and Zoology, 8 (1), 415–433.
  2. Altın, Ç. & Yağmur, E.A. (2018) Faunistic studies on the species of Paederinae and Xantholinini (Coleoptera: Staphylinidae) in Sarıkamış forest, Kars province, Turkey. Munis Entomology and Zoology, 13 (2), 458–462.
  3. Anlaş, S. (2009) Distributional checklist of the Staphylinidae (Coleoptera) of Turkey, with new and additional records. Linzer Biologische Beiträge, 41 (1), 215–342.
  4. Anlaş, S. (2014) On the genus Xantholinus Dejean of Turkey: three new species, new and additional records, with distributional checklist (Coleoptera: Staphylinidae: Staphylininae: Xantholinini). Journal of Insects Biodiversity, 2, 1–28. https://doi.org/10.12976/jib/2014.2.11
  5. Anlaş, S. (2017) The Genus Xantholinus Dejean, 1821 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) in Turkey: a New Species from South-eastern Anatolia and New Records of Known Species. Acta Zoologica Bulgarica, 69 (4), 457–464.
  6. Anlaş, S. (2019) Notes on the genus Xantholinus Dejean, 1821 (Coleoptera, Staphylinidae, Staphylininae, Xantholinini) from the western Palearctic Region. Turkiye Entomoloji Dergisi–Turkish Journal of Entomology, 43 (2), 211–225. https://doi.org/10.16970/entoted.516235
  7. Anlaş, S. & Rose, A. (2009) Some additional notes about Staphylininae (Coleoptera: Staphylinidae) fauna of Turkey. Munis Entomology and Zoology 4 (2), 346–352.
  8. Anlaş, S., Tezcan, S. & Örgel, S. (2014) Seasonal Dynamics and Species Composition of Dung-Inhabiting Staphylinids (Coleoptera, Staphylinidae) in Western Turkey. Journal of Anatolian Natural Sciences, 5 (1), 14–19.
  9. Apfelbeck, V. (1901) Bericht über eine Entomologische Forschungsreise nach der Türkei und Griechenland im jahre 1900. Wiss Mitt Bosnisch-Herzegowin Landesmus, 8, 447–469.
  10. Assing, V. (1997) A revision of Othius Stephens, 1829. III. The species of the Western Palaearctic region exclusive of the Atlantic Islands (Coleoptera, Staphylinidae, Xantholininae). Nova Supplementa Entomologica, 10, 3–130.
  11. Assing, V. (1999) A revision of Othius Stephens (Coleoptera, Staphylinidae). VIII. Further records, new species, and a new synonym. Linzer Biologische Beiträge, 31 (2), 661–691.
  12. Assing, V. (2003a) A revision of Othiini. XIII. Horizontal and vertical distribution of Othius, new species, and additional records (Coleoptera: Staphylinidae: Staphylininae). Entomological Problems, 33 (1–2), 69–88.
  13. Assing, V. (2003b) A revision of Othius Stephens. IX. New Taxa, New Synonyms, Additional Records, and a Checklist of Species (Coleoptera, Staphylinidae, Staphylininae, Othiini). In: Cuccodoro, G. & Leschen, R.A.B. (Eds.), Systematics of Coleoptera: Papers Celebrating the Retirement of Ivan Löbl, Memoirs on Entomology. International Associated Publishers, Gainsville, Florida, pp. 726–752.
  14. Assing, V. (2003c) New species and records of Staphylinidae from Turkey (Insecta: Coleoptera: Staphylinidae). Entomologische Blätter, 98 (3), 153–177.
  15. Assing, V. (2005) A revision of Othiini. XIV. New species, new synonyms, and new records (Insecta: Coleoptera: Staphylinidae). Entomological Problems, 35 (1), 51–67.
  16. Assing, V. (2006) New species and records of Staphylinidae from Turkey IV, with six new synonymies (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 76, 223–276.
  17. Assing, V. (2007a) On the Xantholinini of Turkey and adjacent regions (Coleoptera: Staphylinidae: Staphylininae). Zootaxa, 1474 (1), 1–54. https://doi.org/10.11646/zootaxa.1474.1.1
  18. Assing, V. (2007b) New species and additional records of Staphylinidae from Turkey V (Coleoptera). Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 700, 1–64.
  19. Assing, V. (2008a) A revision of Othiini. XVI. Four new species of Othius from the Himalaya and China, and additional records (Coleoptera: Staphylinidae, Staphylininae). Koleopterologische Rundschau, 78, 245–263.
  20. Assing, V. (2008b) On the taxonomy and zoogeography of some Palaearctic Paederinae and Xantholinini (Coleoptera: Staphylinidae). Linzer Biologische Beiträge, 40 (2), 1237–1294.
  21. Assing, V. (2009) On the Staphylinidae of Turkey. VI. Thirteen new species and additional records (Coleoptera). Koleopterologische Rundschau, 79, 117–172.
  22. Assing, V. (2010a) A revision of Othiini. XVII. A new species from China and additional records (Coleoptera: Staphylinidae: Staphylininae). Linzer Biologische Beiträge, 42 (2), 1077–1091.
  23. Assing, V. (2010b) On the Staphylinidae of Turkey. VII. Five new species and additional records (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 80, 71–102.
  24. Assing, V. (2013) On the Staphylinidae (Coleoptera) of Turkey IX. Five new species, a new synonymy, and additional records. Stuttgarter Beiträge zur Naturkunde A, Neue Serie, 6, 103–125.
  25. Assing, V. (2014) On the Staphylinidae of Turkey X. Two new species and additional records (Insecta: Coleoptera). Linzer Biologische Beiträge, 46 (2), 1133–1146.
  26. Assing, V. (2018) On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae). Linzer Biologische Beiträge, 50 (1), 149–182. https://doi.org/10.5281/zenodo.3985368
  27. Assing, V. (2019) On the taxonomy of some West Palaearctic Quedius species, with descriptions of new species and new synonymies (Coleoptera: Staphylinidae: Staphylininae). Linzer Biologische Beiträge, 51 (1), 189–201. https://doi.org/10.5281/zenodo.3762449
  28. Assing, V. & Schülke, M. (2012) Freude-Harde-Lohse-Klausnitzer—Die Käfer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, 560 pp.
  29. Atalay, İ. & Mortan, K. (2011) Resimli ve Haritalı Türkiye Bölgesel Coğrafyası. İnkılap Kitabevi, İstanbul, 632 pp.
  30. Bodemeyer, E. von (1900) Quer durch Klein-Asien in den Bulghar-Dagh: Eine naturwissenschaftliche Studien-Reise. Die Druck und Verlags-Aktien Gesellschaft Vormals Dölter, Emmendingen, 169 pp. https://doi.org/10.5962/bhl.title.34272
  31. Bodemeyer, B. von (1927) Über meine entomologischen Reisen nach Kleinasien (1911), Ost-Sibirien, Schilka und Amur (1912), Tunis, Oasis Gafsa, Khroumerie (1913) und Iran, das Elbursgebirge (1914). Band 1. Kleinasien. Kernen, Stuttgart, 85 pp.
  32. Bohac, J. (1999) Staphylinid beetles as bioindicators. Agriculture Ecosystems & Environment, 74 (1–3), 357–372. https://doi.org/10.1016/S0167-8809(99)00043-2
  33. Bordoni, A. (1971) Note sugli Xantholinus della Turchia e descrizione di una nuova specie (Col. Staphylinidae). Redia, 52, 679–689.
  34. Bordoni, A. (1973) Nuove specie di Xantholininae della Turchia e del Libano (Coleoptera, Staphylinidae). Fragmenta Entomologica, 9 (3), 179–194.
  35. Bordoni, A. (1976a) Quarto contributo alla conoscenza degli Staphylinidae della Turchia (Coleoptera). Fragmenta Entomologica, 12 (3), 221–239.
  36. Bordoni, A. (1976b) Studi sulla sistematica e la geonemia degli Xantholinus. IX. Xantholinus del Museo di Storia Naturale di Ginevra raccolti nel bacino del Mediterraneo ed appunti su alcuni altri Xantholinini. Prima nota (Col. Staphylinidae). Revue Suisse de Zoologie, 83 (1), 83–103. https://doi.org/10.5962/bhl.part.91434
  37. Bordoni, A. (1978) Staphilinidae dell’Asia Minore. Quinta nota: Entita’ raccolte in grotta e descrizione di nuove specie (Coleoptera), Fauna Ipogea di Turchia. Quaderni di Speleologia, Circolo Speleologico Romano, 3, 55–67.
  38. Bordoni, A. (1986) Xantholininae e Paederinae del Museo del Storia Naturale di Praga e descrizione di nuovi Lathrobiini (Col., Staphylinidae). 87 contributo alla conoscenza degli Staphylinidae. Frustula Entomologica, 7–8, 385–394.
  39. Bordoni, A. (1999) Note sulla sinonimia di alcune specie e sottogeneri del genere Xantholinus Dejan [sic], 1821 e designazione di lectotipi (Coleoptera: Staphylinidae). Opuscula Zoologica Fluminensia, 171, 1–8.
  40. Bordoni, A. (2003) Note su alcuni Xantholinini euroasiatici e descrizione di un nuovo genere della Manciuria (Coleoptera Staphylinidae). Bollettino della Società Entomologica Italiana, 134 (3), 219–228.
  41. Bordoni, A. (2004) Descrizione del primo Lepidophallus del Mediterraneo orientale (Coleoptera Staphylinidae). Bollettino della Società Entomologica Italiana, 136 (1), 33–36.
  42. Bordoni, A. (2005) Sui Gauropterus della Regione Paleartica e in particolare su quelli. descritti da Kirschenblatt, conservati nel Museo Zoologico di San Pietroburgo (Coleoptera, Staphylinidae). Bollettino della Società Entomologica Italiana, 137, 205–213.
  43. Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125–434. https://doi.org/10.5962/bhl.title.48654
  44. Coiffait, H. (1956a) Les Xantholinitae de France et des régions voisines (Col. Staphylinidae). Revue Française d’Entomologie, 23, 31–75.
  45. Coiffait, H. (1956b) Nouveaux Xantholininae d’Europe centrale et de la région méditerranéenne (Col. Staphylinidae). Bulletin de la Société entomologique de France, 61, 139–143. https://doi.org/10.3406/bsef.1956.18849
  46. Coiffait, H. (1965) Sur quelques Xantholininae de l’Asie Mineure (Coleoptera, Staphylinidae). Reichenbachia, 5 (12), 119–123.
  47. Coiffait, H. (1966) Anadolu’nun Xantholininae (Col. Staphylinidae) leri II. Istanbul Üniversitesi Fen Fakültesi Mecmuasi B: Sciences Naturelles, 31 (1–2), 21–24.
  48. Coiffait, H. (1972) Coleopteres Staphylinidae de la region Palearctique occidentale. I. Generalites. Sous-familles: Xantholininae et Leptotyphlinae. Nouvelle Revue d'Entomologie, Toulouse, Supplement 2, 1–651.
  49. Coiffait, H. (1978) Staphylinides récoltés par T. Deuve en Anatolie septentrionale (Col. Staph.). Nouvelle Revue d’Entomologie, 8 (2), 163–175.
  50. Çiftçi, D. & Hasbenli, A. (2016) Faunistic Studies on Steninae, Paederinae and Staphylinae (Coleoptera: Staphylinidae) from Sündiken Mountains (Turkey). Entomofauna, 37 (6), 101–136.
  51. Dejean, P. F. M. A. (1821) Catalogue de la collection de coléoptères de M. le Baron Dejean. Crevot, Libraire, Paris, 136 pp. https://doi.org/10.5962/bhl.title.11259
  52. Demirsoy, A. (2003) Yaşamın Temel Kuralları, “Omurgasızlar/Böcekler-Entomoloji” Cilt-II/Kısım-II. 8th Edition. Meteksan A.Ş., Ankara, 941 pp.
  53. Erichson, W.F. (1839a) Erster Band. In: Genera et species Staphylinorum insectorum coleopterorum familiae. F.H. Morin, Berlin, pp. 1–400. https://doi.org/10.5962/bhl.title.59644
  54. Erichson, W.F. (1839b) Die Käfer der Mark Brandenburg. Erster Band. Zweite Abtheilung. Morin, Berlin, pp. 385–740.
  55. Fabricius, J.C. (1787) Mantissa insectorvm sistens eorvm species nvper detectas adiectis characteribvs genericis, differentiis specificis, emendationibvs, observationibvs. Tom. I. C.G. Proft, Hafniae, 348 pp. https://doi.org/10.5962/bhl.title.11657
  56. Fauvel, A. (1873) Faune Gallo-Rhénane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhénanes et la Valais avec tableaux synoptiques et planches gravées. Bulletin de la Société Linnéenne de Normandie, 7, 8–131.
  57. Fauvel, A. (1900) Staphylinides paléarctiques nouveaux. Revue d’Entomologie, 19, 218–253.
  58. Fırat, S. & Sert, O. (2016) Faunistic and zoogeographical composition and preliminary evaluations of some ecological features of the subfamily Staphylininae (Coleoptera: Staphylinidae) of the Central Anatolian Region of Turkey. Turkish Journal of Zoology, 40 (2), 164–185. https://doi.org/10.3906/zoo-1501-46
  59. Frank, J.H. & Thomas, M.C. (2010) Rove Beetles of the World, Staphylinidae (Insecta: Coleoptera: Staphylinidae). Featured Creatures from the Entomology and Nematology Department, Florida Cooperative Extension Service, Institute of Food and Agricultural Sciences, University of Florida, EENY-114 (IN271). Available from: http://edis.ifas.ufl.edu/in271 (accessed 4 September 2023)
  60. Ganglbauer, L. (1895) Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des französischen und italienischen Alpengebietes. Zweiter Band. Famielienreihe Staphylinoidea. 1. Theil. Staphylinidae, Pselaphidae. Carl Gerold’s Sohn, Wien, 881 pp.
  61. Goeze, J.A.E. (1777) Entomologische Beyträge zu des Ritter Linné zwölften Ausgabe des Natursystems. Erster Theil. Weidmanns Erben & Reich, Leipzig, 736 pp. https://doi.org/10.5962/bhl.title.45974
  62. Gravenhorst, J.L.C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. Carolus Reichard, Brunsuigae, 206 pp. https://doi.org/10.5962/bhl.title.9568
  63. Gutiérrez-Chacon, C., Zúñiga, M.D.C., Van Bodegom, P.M., Chará, J. & Giraldo, L.P. (2009) Rove beetles (Coleoptera: Staphylinidae) in Neotropical riverine landscapes: characterising their distribution. Insect Conservation and Diversity, 2 (2), 106–115. https://doi.org/10.1111/j.1752-4598.2009.00050.x
  64. Gyllenhal, L. (1827) Insecta Suecica descripta. Classis I. Coleoptera sive Eleuterata. Tom I. Pars IV. Cum appendice ad partes priores. F. Fleischer, Lipsiae, 762 pp.
  65. Hochhuth, J.H. (1849) Die Staphylinen-Fauna des Kaukasus und Transkaukasiens. Bulletin de la Société Impériale des Naturalistes de Moscou, 22 (1), 18–214.
  66. Horion, A. (1965) Faunistik der mitteleuropäischen Käfer. Staphylinidae. 2. Paederinae bis Staphylininae. Verlagsdruckerei Ph. C. W. Schmidt, Überlingen-Bodensee, 335 pp.
  67. Jacquelin du Val, P.N.C. (1856) Famille des staphylinides. In: Manuel Entomologique. Genera des coléoptères d’Europe comprenant leur classification en familles naturelles, la description de tous les genres, des tableaux synoptiques destinés à faciliter l’étude, le catalogue de toutes les espèces de nombreux au trait de caractères Tome Deuxième. A. Deyrolle, Paris, pp. 1–40. https://doi.org/10.5962/t.173866
  68. Japoshvili, G. & Anlaş, S. (2011) Notes on the family staphylinidae (coleoptera) collected by pitfall traps in Gölcük Natural Park, Isparta province of Turkey. Journal of the Entomological Research Society, 13 (1), 41–48.
  69. Johnson, S.N. & Jones, T.H. (2017) Introduction to global climate change and terrestrial invertebrates. In: Johnson, S.N. & Jones, T.H. (Eds.), Global climate change and terrestrial invertebrates. John Wiley & Sons, Ltd., Chichester, pp. 1–8. https://doi.org/10.1002/9781119070894.ch1
  70. Kesdek, M., Yıldırım, E., Anlaş, S. & Tezcan, S. (2009) Contribution to the knowledge of Staphylinidae (Coleoptera) fauna of Turkey. Munis Entomology & Zoology, 4 (2), 355–364.
  71. Kolenati, F.A.R. (1846) Meletemata entomologica. Fascicule III. Brachelytra Caucasi cum distributione geographica adnexis pselaphinis, scydmaenis, notoxidibus et xylophagis. Typis Imperialis Academiae Scientiarum, Petropoli, 44 pp.
  72. Kraatz, G. (1858) Beitrag zur Käferfauna Griechenlands. Zweites Stück: Palpicornia, Silphales, Scydmaenidae, Pselaphidae, Staphylinidae. Berliner Entomologische Zeitschrift, 2, 37–67. https://doi.org/10.1002/mmnd.18580020106
  73. Leach, W.E. (1819) [new genera]. In: Samouelle, G. (Ed.), The Entomologist’s useful compendium; or an introduction to the knowledge of British insects, comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linné, and the modern method of arranging the classes Crustacea, Myriapoda, spiders, mites, and insects from their affinities and structure, according to the views of Dr. Leach. Also an explanation of the terms used in entomology; a calendar of the times of appearance, and usual situations of near 3000 species of British insects; with instructions for collecting and fitting up objects for the microscope. Thomas Boys, London, pp. 496.
  74. Löbl, I. & Löbl, D. (2015) Catalogue of Palaearctic Coleoptera Hydrophiloidea- Staphylinoidea. Vol. 2. Revised and Updated Edition. Vol. 2. Brill, Leiden, 1702 pp. https://doi.org/10.1163/9789004296855
  75. Märkel, F. & Kiesenwetter, E.A.H. von (1848) Bericht über eine entomologische Excursion in die Kärnthner Alpen im Jahre 1847. Entomologische Zeitung, Stettin, 9 (11), 321–329.
  76. Mulsant, E. & Rey, C. (1877) Histoire naturelle des Coléoptères de France. Tribu des brévipennes. Deuxième famille. Xantholiniens. Mémoires de l’Académie des Sciences, Belles-Lettres et Arts de Lyon, 22, 217–344. https://doi.org/10.5962/bhl.title.51574
  77. Müller, O.F. (1776) Zoologia Danicae prodromus, seu animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. Hallager, Hafniae, 282 pp. https://doi.org/10.5962/bhl.title.13268
  78. Newton, A. (2022) Staphyliniformia world catalog database. In: Bánki, O., Roskov, Y., Döring, M.., Ower, G., Vandepitte, L., Hobern, D., Remsen, D., Schalk, P., DeWalt, R.E., Keping, M., Miller, J., Orrell, T., Aalbu, R., Adlard, R., Adriaenssens, E.M., Aedo, C., Aescht, E., Akkari, N., Alexander, S. et al., Catalogue of Life Checklist. August 2022. Available from: https://doi.org/10.48580/dfqf-3gk (accessed 11 December 2023)
  79. Olivier, A.G. (1795) Entomologie, ou histoire naturelle des insectes, avec leur caractères génériques et spécifiques, leur description, leur synonymie, et leur figure illuminée. Coléoptères. Tome troisième. Lanneau, Paris, 557 pp.
  80. Öncüer, C. (1991) Türkiye bitki zararlısı böceklerin parazit ve predatör kataloğu-A Catalogue of the Parasites and Predators of Insect Pests of Turkey. Vol. 505. Ege Üniversitesi Ziraat Fakültesi Yayınları, İzmir, 354 pp.
  81. Özdemir, S. (2021) Faunistic Contributions, and Ecological and Zoogeographical Evaluations on Staphylininae (Coleoptera: Staphylinidae) Fauna of Eastern Black Sea Region of Turkey. Transactions of the American Entomological Society, 147 (4), 929–959. https://doi.org/10.3157/061.147.0408
  82. Özgen, İ. (2017) Contributions to the knowledge fauna of staphylininae (Coleoptera: Staphylinidae) in Turkey. Journal of Entomology and Zoology Studies, 5 (2), 1488–1490.
  83. Özgen, İ. (2019) New and additional records of the family Staphylinidae (Insecta: Coleoptera) from Eastern Anatolia, Turkey. International Journal of Fauna and Biological Studies, 6 (4), 119–122.
  84. Özgen, İ., Anlaş, S. & Eren, S. (2010) Contribution to the knowledge of Staphylinidae (Coleoptera) fauna of cotton and pistachio fields in Southeastern Anatolia. Journal of Anatolian Natural Sciences, 1 (1), 20–26.
  85. Peyron, E. (1858) Catalogue des Coléoptères des environs de Tarsous (Caramanie), avec la description des espèces nouvelles. Annales De La Societe Entomologique De France, 3, 353–434.
  86. Reitter, E. (1889) Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen über bekannte Arten. Siebenter Theil. Deutsche Entomologische Zeitschrift, 2, 273–288.
  87. Reitter, E. (1908) Bestimmungs-Tabelle der Staphyliniden-Gruppen der Othiini und Xantholinini aus Europa und den angrenzenden Ländern. Verhandlungen des Naturforschenden Vereines in Brünn, 46, 100–124. [1907]
  88. Sahlberg, J. (1913) Coleoptera mediterranea orientalia, queae in Aegypto, Palaestina, Syria, Caramanis atque in Anatolia occidentali anno 1904 collegerunt John Sahlberg et Unio Saalas. Öfversigt af Finska Vetenskaps-Societetens Förhandlinger (A), 55 (19), 1–281.
  89. Schülke, M. & Smetana, A. (2015) Staphylinidae. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea—Staphylinoidea. Revised and updated edition. Brill, Leiden, pp. 304–1134.
  90. Solodovnikov, A. Y. (2000) On the Othius Stephens in the fauna of the Caucasus and Northeastern Anatolia (Coleoptera: Staphylinidae: Xantholininae). Zoosystematica Rossica, 8 (2), 324–328. [1999]
  91. Stephens, J.F. (1829) The nomenclature of British insects; being a compendious list of such species as are contained in the Systematic Catalogue of British Insects, and forming a guide to their classification, &c. &c. Baldwin and Cradock, London, 68 pp. https://doi.org/10.5962/bhl.title.51800
  92. Stephens, J.F. (1833) n.k. In: Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V. Baldwin and Cradock, London, pp. 241–304.
  93. Taylan, D. & Toker, F. (2023) Hydrological drought analysis for Uşak province surrounding. Journal of Engineering Sciences and Design, 11 (1), 1–21. https://doi.org/10.21923/jesd.1194882
  94. Tezcan, S. & Amiryan, J. (2003) The rove beetles (Coleoptera, Staphylinidae) of the ecologically managed cherry orchards of western Turkey. Materials of the IV. Republican Youth Scientific Conference, 2003, 83–92.
  95. Tihelka, E., Thayer, M.K., Newton, A.F. & Cai, C. (2020) New Data, Old Story: Molecular Data Illuminate the Tribal Relationships among Rove Beetles of the Subfamily Staphylininae (Coleoptera: Staphylinidae). Insects, 11 (3), 164. https://doi.org/10.3390/insects11030164
  96. Thomson, C.G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Berlingska Boktryckeriet, Lund, 290 pp. https://doi.org/10.5962/bhl.title.138677
  97. Thomson, C.G. (1860) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom. II. Berlingska Boktryckeriet, Lund, 304 pp. https://doi.org/10.5962/bhl.title.138677