Abstract
In North America west of the Continental Divide, the milliped family Polydesmidae consists of six genera and 15 species, and occupies eight disjunct areas. Retrorsia, n. gen., is proposed for two new “micro-polydesmid” species (R. leonardi, the type species, and R. benedictae) in western Oregon and Washington, primarily in the lower Columbia River Valley. It is most closely related to Utadesmus Chamberlin and Hoffman, 1950, in Utah and New Mexico, and is characterized primarily by a retrorse gonopodal tibiotarsus that curves dorsad and counters the ventral curvature of the telopodal stem. New records are presented for the coastal species of Bidentogon Buckett and Gardner, 1968, occurring from Mendocino to Santa Cruz counties, California, and encompassing the area around San Francisco Bay. The oldest available name for this species is californicus (Chamberlin, 1918), whose type locality is Stanford, Santa Clara County. This name has been misapplied to the species occurring in Sacramento County, herein named B. expansus, n. sp., and B. helferorum Buckett and Gardner, 1968, is placed in synonymy under B. californicus.References
Attems, C. (1940) Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich, Lief. 70, 1–576.
Barr, T.C. & Reddell, J.R. (1967) The arthropod cave fauna of the Carlsbad Caverns region, New Mexico. Southwestern Naturalist, 12, 253–274.
Buckett, J.S. (1964) Annotated list of the Diplopoda of California. Simmons Publishing Co., Davis, California, 34 pp.
Buckett, J.S. & Gardner, M.R. (1968) A new genus and species of milliped from northern California (Polydesmida: Vanhoeffeniidae). Pan-Pacific Entomologist, 44, 198–202.
Causey, N.B. (1959) Two new troglodytic millipeds from Texas. Proceedings of the Biological Society of Washington, 72, 69–74.
Chamberlin, R.V. (1918) Four new western diplopods. Pomona College Journal of Entomology and Zoology, 10, 9–11.
Chamberlin, R.V. (1952) Three cave-dwelling millipeds. Entomological News, 63, 10–12.
Chamberlin, R.V., & Hoffman, R.L. (1958) Checklist of the millipeds of North America. United States National Museum Bulletin 212, 1–236.
Hoffman, R.L. (1980) Classification of the Diplopoda. Muséum d’Histoire Naturelle, Genève, 237 pp.
Hoffman, R.L. (1999) Checklist of the Millipeds of North and Middle America. VirginiaMuseum of Natural History Special Publication No. 8, 1–584.
Loomis, H.F. (1960) Millipeds of the order Polydesmida from the western states and Baja California. Journal of the Kansas Entomological Society, 33, 57–68.
Murakami, Y. (1954) Monograph on Japanese Diplopoda. 241 pp. [In Japanese, published in Japan]. Reddell, J.R. (1965) A checklist of the cave fauna of Texas. I. The Invertebrata (exclusive of Insecta). Texas Journal of Science, 17, 143–187.
Reddell, J.R. (1970) A checklist of the cave fauna of Texas. IV. Additional records of Invertebrata (exclusive of Insecta). Texas Journal of Science, 21, 389–415.
Shear, W.A. (1969) A synopsis of the cave millipeds of the United States, with an illustrated key to genera. Psyche, 76, 126–143.
Shear,W.A. (1972) The milliped genus Bidentogon (Diplopoda, Polydesmida, Trichopolydesmidae). Proceedings of the Biological Society of Washington, 85, 489–492.
Shear, W.A. (1974) North American cave millipeds. II. An unusual new species (Dorypetalidae) from southern California, and new records of Speodesmus tuganbius (Trichopolydesmidae) from New Mexico. Occasional Papers of the California Academy of Sciences No. 112, 1–9.
Shear, W.A. (1984) Cave millipeds of the United States. III. Two new species from the western states (Diplopoda, Polydesmida, Chordeumatida). Myriapodologica, 1, 95–104.
Shear, W.A. & Leonard, W.P. (2003) Microlympiidae, a new milliped family from North America, and Microlympia echina, new genus and species (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa, 243, 1–11.
Shelley, R.M. (1988) The millipeds of eastern Canada (Arthropoda: Diplopoda). Canadian Journal of Zoology, 66, 1638–1663.
Shelley, R.M. (1993) Revision of the milliped genus Scytonotus Koch (Polydesmida: Polydes-midae). Brimleyana, 19, 1–60.
Shelley, R.M. (1996) The milliped genus Utadesmus Chamberlin and Hoffman (Polydesmida: Polydesmidae). Myriapodologica, 4, 9–16.
Shelley, R.M. (1997) The identity of Polydesmus sastianus Chamberlin, proposal of a new milliped genus, and remarks on the identity of Phreatodesmus hastingsus (Chamberlin) (Polydesmida: Polydesmidae). Myriapodologica, 4, 59–67.
Shelley, R.M. (2002) Annotated checklist of the millipeds of California (Arthropoda: Diplopoda). Monographs of the Western North American Naturalist, 1, 90–115.
Shelley, R.M., Sierwald, P., Kiser, S.B. & Golovatch, S.I. (2000) Nomenclator generum et familiarum Diplopodorum II. A List of the Genus and Family-Group Names in the Class Diplopoda from 1958 through 1999. Pensoft Publishers, Sofia-Moscow. 167 pp.
Verhoeff, K.W. (1940) Zur Kenntnis ostasiatischer Diplopoden V (Polydesmoidae). Zoologischer Anzeiger, 131, 129–145.