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Issue: Vol. 5414 No. 1: 19 Feb. 2024
Type: Monograph
Published: 2024-02-19
DOI: 10.11646/zootaxa.5414.1.1
Page range: 1-83
Abstract views: 97
PDF downloaded: 5

Revision of Neotropical wolf spider genus Arctosa C.L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species

Pontifícia Universidade Católica do Rio Grande do Sul; Escola de Ciências da Saúde e da Vida; Museu de Ciências e Tecnologia; Laboratório de Aracnologia; Porto Alegre; Brazil; Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos; Laboratorio de Entomología; Av. Arenales 1256; Lima 14; Peru
Laboratório de Coleções Zoológicas; Av. Vital Brasil; 1500; Butantã; 05503-900; São Paulo; SP; Brazil
Pontifícia Universidade Católica do Rio Grande do Sul; Escola de Ciências da Saúde e da Vida; Museu de Ciências e Tecnologia; Laboratório de Aracnologia; Porto Alegre; Brazil
Araneae Diversity taxonomy wetlands Andes distribution

Abstract

The genus Arctosa C.L. Koch is redescribed and diagnosed. Seven new species are described, four of them based on both sexes (A. conflicta sp. nov., A. costenola sp. nov., A. jibarosa sp. nov. and A. villa sp. nov.), two only on females (A. ayaymama sp. nov. and A. mineira sp. nov.), and one only on the male (A. pacaya sp. nov.). We also propose the transference of Arctosa humicola (Bertkau, 1880) and Arctosa fusca (Keyserling, 1877) to Trochosa C.L. Koch, 1847, and the new synonymy of Arctosa inconspicua (Bryant, 1948) with Trochosa humicola (Bertkau, 1880) comb. nov. Besides, Arctosa andina (Chamberlin, 1916) and Arctosa pugil (Bertkau, 1880) are transferred to Hogna Simon, 1885, and Arctosa aussereri (Keyserling, 1877) to Prolycosides Mello-Leitão, 1942. Additionally, eight lycosid species are synonymized with Prolycosides aussereri: Schizocosa proletaria (Tullgren, 1905); Arctosa workmani (Strand, 1909); Hogna planithoracis (Mello-Leitão, 1938); Hogna variolosa (Mello-Leitão, 1941); Megarctosa melanostoma (Mello-Leitão, 1941); Hippasosa huachoi (Mello-Leitão, 1942); Pirata abalosi (Mello-Leitão, 1942); and Pirata soukupi (Mello-Leitão, 1942). We also transfer Trochosa tenebrosa Keyserling, 1877 to Arctosa. The males of Arctosa tenebrosa (Keyserling, 1877) comb. nov. and Trochosa humicola (Bertkau, 1880) comb. nov. are described for the first time.

 

References

  1. Alderweireldt, M. & Jocqué, R. (1993) A redescription of Tricassa deserticola Simon, 1910, representing the Tricassinae, a new subfamily of wolf spiders (Araneae, Lycosidae). Belgian Journal of Zoology, 123 (1), 27–38.
  2. Álvarez-Padilla, F. & Hormiga, G. (2007) A protocol for digesting internal soft tissues and mounting spiders for scanning electron microscopy. The Journal of Arachnology, 35 (3), 538–542. https://doi.org/10.1636/Sh06-55.1
  3. Banks, N. (1901) Some spiders and other Arachnida from Porto Rico. Proceedings of the United States National Museum, 24, 217–227. https://doi.org/10.5479/si.00963801.24-1253.217
  4. Bertkau, P. (1880) Verzeichniss der von prof. Ed. van Beneden auf seiner im auftrage der Belgischen regierung unternommen wissenschaftlichen reise nach Brasilien und La Plata im Jahren 1872–73 gesammelten Arachniden. Mémoires Couronnés et Mémoires des Savants Étrangers de l’Académie Royale des Sciences, Des Lettres et Des Beaux-Arts de Belgique, 43, 1–120.
  5. Bizuet-Flores, M.Y., Jiménez-Jiménez, M.L., Zavala-Hurtado, A. & Corcuerab, P. (2015) Diversity patterns of ground dwelling spiders (Arachnida: Araneae) in five prevailing plant communities of the Cuatro Ciénegas Basin, Coahuila, Mexico. Revista Mexicana de Biodiversidad, 86 (1), 153–163. https://doi.org/10.7550/rmb.45444
  6. Bonnet, P. (1955) Bibliographia araneorum. Analyse méthodique de toute la littérature aranéologique jusqu’en 1939. Tome II. Systématique des araignées (Étude par ordre alphabétique. 1re partie: A–B). Douladoure, Toulouse, pp. 1–918.
  7. Bonnet, P. (1957) Bibliographia araneorum. Analyse méthodique de toute la littérature aranéologique jusqu'en 1939. Tome II. Systématique des araignées (Étude par ordre alphabétique. 3e partie: G–M). Douladoure, Toulouse, pp. 1927–3026.
  8. Bonnet, P. (1959) Bibliographia araneorum. Analyse méthodique de toute la littérature aranéologique jusqu’en 1939. Tome II. Systématique des araignées (Étude par ordre alphabétique. 5me partie: T–Z). Douladoure, Toulouse, pp. 4231–5058.
  9. Bösenberg, W. & Strand, E. (1906) Japanische spinnen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 30, 93–422.
  10. Brady, A.R. (1979) Nearctic species of the wolf spider genus Trochosa (Araneae: Lycosidae). Psyche, 86, 167–212. https://doi.org/10.1155/1979/46401
  11. Brady, A.R. (2012) Nearctic species of the new genus Tigrosa (Araneae: Lycosidae). The Journal of Arachnology, 40 (2), 182–208. https://doi.org/10.1636/k11-77.1
  12. Braun, R. (1963) Das Tricca-problem (Arach., Araneae). Senckenbergiana Biologica, 44, 73–82.
  13. Bryant, E.B. (1940) Cuban spider in the museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology, 86 (7), 247–532.
  14. Bryant, E.B. (1948) The spiders of Hispaniola. Bulletin of the Museum of Comparative Zoology, 100 (4), 329–447.
  15. Buchar, J., Knoflach, B. & Thaler, K. (2006) On the identity of Arctosa varians C.L. Koch and Arctosa similis Schenkel, with notes on related species (Araneae: Lycosidae). Bulletin of the British Arachnological Society, 13 (9), 329–336.
  16. Buchholz, S. & Schröder, M. (2013) Diversity and ecology of spider assemblages of a Mediterranean wetland complex. The Journal of Arachnology, 41 (3), 364–373. https://doi.org/10.1636/P13-26.1
  17. Chamberlin, R.V. (1908) Revision of North American spiders of the family Lycosidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 60, 158–318.
  18. Chamberlin, R.V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177–299.
  19. Chamberlin, R.V. (1924) The spider fauna of the shores and islands of the Gulf of California. Proceedings of the California Academy of Sciences, 12, 561–694.
  20. Chamberlin, R.V. & Ivie, W. (1942) A hundred new species of American spiders. Bulletin of the University of Utah, 32 (13), 1–117.
  21. Dolejš, P., Kubcová, L. & Buchar, J. (2008) Subterrestrial life of Arctosa lutetiana (Araneae, Lycosidae). The Journal of Arachnology, 36 (1), 202–203. https://doi.org/10.1636/St07-33SC.1
  22. Dondale, C.D. (1986) The subfamilies of wolf spiders. Actas X Congreso Internacional de Aracnologia, Jaca/España, 1986, 327–332.
  23. Dondale, C.D. & Redner, J.H. (1983a) Revision of the wolf spiders of the genus Arctosa C.L. Koch in North and Central America (Araneae: Lycosidae). The Journal of Arachnology, 11 (1), 1–30.
  24. Dondale, C.D. & Redner, J.H. (1983b) The wolf spider genus Allocosa in North and Central America (Araneae: Lycosidae). The Canadian Entomologist, 115 (8), 933–964. https://doi.org/10.4039/Ent115933-8
  25. Dondale, C.D. & Redner, J.H. (1990) The insects and arachnids of Canada, Part 17. The wolf spiders, nurseryweb spiders, and lynx spiders of Canada and Alaska, Araneae: Lycosidae, Pisauridae, and Oxyopidae. Research Branch Agriculture Canada Publication, 1856, 1–383.
  26. Doleschall, L. (1852) Systematisches verzeichniss der im Kaiserthum Österreich vorkommenden spinnen. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematischnaturwissenschaftliche Klasse, 9, 622–651.
  27. Emerton, J.H. (1885) New England Lycosidae. Transactions of the Connecticut Academy of Arts and Sciences, 6, 481–505. https://doi.org/10.5962/bhl.part.7415
  28. Emerton, J.H. (1902) The common spiders of the United States. Ginn & Company, Boston, Massachusetts, 225 pp. https://doi.org/10.5962/bhl.title.28675
  29. Esquivel-Bobadilla, S., Adrian Lozano-Garza, O., García-De-León, F.J., Barriga-Sosa, I. de los A. & Jiménez, M.L. (2013) Development and characterization of 14 microsatellite loci in the beach wolf spider (Arctosa littoralis), using next-generation sequencing. Conservation Genetics Resources, 5 (1), 261–263. https://doi.org/10.1007/s12686-012-9782-7
  30. Fabricius, J.C. (1777) Genera insectorum eorumque characteres naturales, secundum numerum, figuram, situm et proportionem omnium partium oris; adjecta mantissa specierum nuper detectarum. Michael Friedrich Bartsch, Chilonii [Kiel], 310 pp.
  31. Framenau, V.W., Dieterich, M., Reich, M. & Plachter, H. (1996) Life cycle, habitat selection and home ranges of Arctosa cinerea (Fabricius, 1777) (Araneae: Lycosidae) in a braided section of the Upper Isar (Germany, Bavaria). Revue Suisse Zoologie, Special Edition, 1, 223–234.
  32. Gertsch, W.J. (1934) Notes on American Lycosidae. American Museum Novitates, 693, 1–26.
  33. Gertsch, W.J. & Wallace, H.K. (1935) Further notes on American Lycosidae. American Museum Novitates, 794, 1–22.
  34. Gertsch, W.J. & Wallace, H.K. (1937) New American Lycosidae with notes on other species. American Musuem Novitates, 919, 1–24.
  35. Gonnet, V., Bidegaray-Batista, L., Aisenberg, A., Laborda, Á., Hagopián, D., Izquierdo, M.A., Piacentini, L.N. & Simó, M. (2021) A wolf spider from South American grasslands: phylogenetic placement and redescription of Paratrochosina amica (Mello-Leitão 1941). Zoologischer Anzeiger, 295, 1–11. https://doi.org/10.1016/j.jcz.2021.08.009
  36. The GIMP Development Team. (2019) GIMP. Available from: https://www.gimp.org (accessed 23 July 2023)
  37. He, J. & Song, D. (1996) The Phylogenetic relationships of Chinese wolf spiders. Acta Arachnologica Sinica, 5 (1), 3–20.
  38. Hentz, N.M. (1844) Descriptions and figures of the araneides of the United States. Boston Journal of Natural History, 4, 386–396. https://doi.org/10.5962/bhl.part.16998
  39. Hippa, H. & Lehtinen, P.T. (1983) The Zantheres group of Zoicinae (Araneae, Lycosidae) and a relimitation of the subfamily. Annales Zoologici Fennici, 20, 151–156.
  40. Inkscape Project. (2020) Inkscape. Available from: https://inkscape.org (accessed 23 July 2023).
  41. Isola, S. (2007) Birds of Pacaya Samiria National Reserve. The Nature Conservancy (TNC) and United States Agency for International Development (USAID), Lima, 64 pp.
  42. Jiménez, M.L., Chamé-Vázquez, D. & Palacios-Cardiel, C. (2022) New records of spiders (Arachnida: Araneae) from the Madrean Archipelago of Sonora, Mexico. Pan-Pacific Entomologist, 98 (3), 196–204. https://doi.org/10.3956/2022-98.3.196
  43. Jiménez, M.L. & Dondale, C.D. (1984) A new species of Arctosa from Guerrero, Mexico (Araneae, Lycosidae). The Journal of Arachnology, 12 (1), 115–116.
  44. Jiménez, M.L., Palacios-Cardiel, C., Maya-Morales, J. & Berrian, J.E. (2018) Nuevos registros de arañas (Arachnida: Araneae) para la Región del Cabo, península de Baja California, México. Acta zoológica mexicana, 34 (1), 1–13. https://doi.org/10.21829/azm.2018.3412159
  45. Jocqué, R. & Alderweireldt, M. (2001) Tricassa madagascariensis n. sp., the second representative of the wolf spider subfamily Tricassinae (Araneae, Lycosidae). Annales Sciences Zoologiques, 285, 29–31.
  46. Karsch, F. (1879) Baustoffe zu einer spinnenfauna von Japan. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 36, 57–105.
  47. Karsten, R. (1923) Blood Revenge, War, and Victory Feasts Among the Jibaro Indians of Eastern Ecuador. Government printing Office, Washington, D.C., 124 pp.
  48. Kaston, B.J. (1948) Spiders of Connecticut. Bulletin of the Connecticut State Geological and Natural History Survey, 1948 (70), 1–874.
  49. Keyserling, G.E. (1877) Ueber amerikanische spinnenarten der unterordnung Citigradae. Verhandlungen Der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 26, 609–708.
  50. Koch, C.L. (1847b) Die Arachniden getreu nach der Natur abgebildet und beschrieben. Vol. 16 (“1848”). J.L. Lotzbeck, Nürnberg, 80 pp.
  51. Koch. L. (1872) Beitrag zur Kenntniss der Arachnidenfauna Tirols. Zweite Abhandlung. Zeitschrift des Ferdinandeums für Tirol und Vorarlberg, Innsbruck, Series 3, 17, 239–328.
  52. Kulczyński, W. (1885) Pająki zebrane na Kamczatce przez Dra B. Dybowskiego. Araneae in Camtschadalia a Dre B. Dybowski collectae. Pamiętnik Akademii Umiejętności w Krakowie, Wydział Matematyczno-Przyrodniczy, 11 (4), 1–60, pls. 9–11.
  53. Kulczyński, W. (1908) Fragmenta arachnologica. X. Araneae nonnullae in Cypro insula et in Palaestina a Cel. Prof. Dre G. Cecconi lectae. Bulletin International de l'Academie des Sciences de Cracovie, 1908, 49–86.
  54. Laborda, Á., Bidegaray-Batista, L., Simó, M., Brescovit, A.D., Beloso, C. & Piacentini, L.N. (2022) Abaycosa a new genus of South American wolf spiders (Lycosidae: Allocosinae). Arthropod Systematics & Phylogeny, 80, 59–74. https://doi.org/10.3897/asp.80.e76339
  55. Lambraño J.E.C. (2011) El costeñol: un dialecto con toda la barba. Pluma de Pompox S.A., Cartagena de Indias, 192 pp.
  56. Latreille, P.A. (1799) Description d’une nouvelle espèce d’araignée. Bulletin des Sciences, par la Société Philomatique, 1 (22), 170.
  57. Levi, H.W. (1964) Nineteenth century South American Arachnology. Papéis Avulsos do Departamento de Zoologia, 16 (1), 9–19.
  58. Levi, H.W. (1965) Techniques for the Study of Spider Genitalia. Psyche, 72 (2), 152–158. https://doi.org/10.1155/1965/94978
  59. Logunov, D.V. (2011) Sexual size dimorphism in burrowing wolf spiders (Araneae: Lycosidae). Proceedings of the Zoological Institute RAS, 315 (3), 274–288. https://doi.org/10.31610/trudyzin/2011.315.3.274
  60. Logunov, D.V. (2020) On three species of Hogna Simon, 1885 (Aranei: Lycosidae) from the near East and Central Asia. Arthropoda Selecta, 29 (3), 349–360. https://doi.org/10.15298/arthsel.29.3.08
  61. Lugetti, G. & Tongiorgi, P. (1965) Revisione delle specie italiane dei generi Arctosa C.L. Koch e Tricca Simon con note su una Acantholycosa delle Alpi Giulie (Araneae - Lycosidae). Redia, 49, 165–228.
  62. Lugetti, G. & Tongiorgi, P. (1966) Su alcune specie dei generi Arctosa C. L. Koch e Tricca Simon (Araneae - Lycosidae). Redia, 50, 133–150.
  63. Mello-Leitão, C.F. de (1938) Algunas arañas nuevas de la Argentina. Revista del Museo de La Plata, 1, 89–118.
  64. Mello-Leitão, C.F. de (1941a) Las Arañas de Córdoba, La Rioja, Catamarca, Tucumán, Salta y Jujuy. Colectadas por los profesores Birabén. Revista del Museo de La Plata, Nueva Serie Zoologia, 2 (12), 99–198.
  65. Mello-Leitão, C.F. de (1941b) Las arañas de la provincia de Santa Fe colectadas por el Profesor Birabén. Revista del Museo de La Plata, Nueva Serie Zoologia, 2, 199–225.
  66. Mello-Leitão, C.F. de (1942a) Cinco aranhas novas do Peru. Revista brasileira de biologia, 2 (4), 429–434.
  67. Mello-Leitão, C.F. de (1942b) Arañas del Chaco y Santiago del Estero. Revista Del Museo de La Plata, Nueva Serie Zoologia, 2 (16), 381–426.
  68. Mello-Leitão, C.F. de (1943) Catálogo das aranhas do Rio Grande do Sul. Arquivos do Museu Nacional do Rio de Janeiro, 37, 147–245.
  69. Mello-Leitão, C.F. de (1947) Aranhas do Paraná e Santa Catarina, das coleções do Museu Paranaense. Arquivos do Museu Paranaense, 6, 231–304.
  70. Mendoza Belmontes, F., Melchor, R.N. & Piacentini, L.N. (2018) Wolf spider burrows from a modern saline sandflat in central Argentina: morphology, taphonomy and clues for recognition of fossil examples. PeerJ, 6, 1–26. https://doi.org/10.7717/peerj.5054
  71. Miller, J.A., Griswold, C.E., Scharff, N., Řezáč, M., Szűts, T. & Marhabaie, M. (2012) The velvet spiders: an atlas of the Eresidae (Arachnida, Araneae). ZooKeys, 195, 1–144. https://doi.org/10.3897/zookeys.195.2342
  72. Montgomery, T.H. (1902) Descriptions of Lycosidae and Oxyopidae of Philadelphia and its vicinity. Proceedings of the Academy of Natural Sciences of Philadelphia, 54, 534–592.
  73. Montgomery, T.H. (1904) Descriptions of North American Araneae of the families Lycosidae and Pisauridae. Proceedings of the Academy of Natural Sciences of Philadelphia, 56, 261–325.
  74. Murphy, N.P., Framenau, V.W., Donnellan, S.C., Harvey, M.S., Park, Y.-C. & Austin, A.D. (2006) Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12S rRNA, 28S rRNA, and NADH1 genes: implications for classification, biogeography, and the evolution of web building behavior. Molecular Phylogenetics and Evolution, 38 (3), 583–602. https://doi.org/10.1016/j.ympev.2005.09.004
  75. Paik, K.Y. (1994) Korean Spiders of the Genus Arctosa C. L. Koch, 1848 (Araneae: Lycosidae). Korean Arachnology, 10, 36–65.
  76. Paquin, P. & Dupérré, N. (2003) Guide d'identification des araignées de Québec. Fabreries, Supplement 11, 1–251.
  77. Paredes-Munguia, W. (2012) Spiders from Pantanos de Villa Reserved Zone Lima-Peru. LAP LAMBERT Academic Publishing, Lima, 128 pp.
  78. Petrunkevitch, A. (1911) A synonymic index-catalogue of spiders of north, central and south America with all adjacent islands, Greenland, Bermuda, West Indies, Tierra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History, 29, 1178–1178. https://doi.org/10.5962/bhl.title.23819
  79. Petrunkevitch, A. (1925) Arachnida from Panama. Transactions of the Connecticut Academy of Arts and Sciences, 27, 51–248.
  80. Petrunkevitch, A. (1929) The spiders of Porto Rico. Part one. Transactions of the Connecticut Academy of Arts and Sciences, 30, 1–158.
  81. Piacentini, L.N. & Laborda, A. (2013) The South American wolf spider genus Birabenia Mello-Leitão, 1941 (Araneae: Lycosidae: Lycosinae). Zootaxa, 3716 (1), 39–52. https://doi.org/10.11646/zootaxa.3716.1.3
  82. Piacentini, L.N. & Ramírez, M.J. (2019) Hunting the wolf: A molecular phylogeny of the wolf spiders (Araneae, Lycosidae). Molecular Phylogenetics and Evolution, 136 (7), 227–240. https://doi.org/10.1016/j.ympev.2019.04.004
  83. Pickard-Cambridge, F.O. (1902) Arachnida - Araneida and Opiliones. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana. Vol. 2. Published for the editors by R. H. Porter, London, pp. 312–424.
  84. Ponomarev, A.V. (2007) New spiders (Aranei) from the south-east of Europe. Caucasian Entomological Bulletin, 3 (1), 3–7. https://doi.org/10.23885/1814-3326-2007-3-1-3-7
  85. Ramírez, M.J. (2003) The spider subfamily Amaurobioidinae (Araneae, Anyphaenidae): a phylogenetic revision at the generic level. Bulletin of the American Museum of Natural History, 277, 1–262. https://doi.org/10.1206/0003-0090(2003)277<0001:TSSAAA>2.0.CO;2
  86. Relys, V., Koponen, S. & Dapkus, D. (2009) Annual Differences and Species Turnover in Peat Bog Spider Communities. The Journal of Arachnology, 30 (2), 416–424. https://doi.org/10.1636/0161-8202(2002)030[0416:ADASTI]2.0.CO;2
  87. Roewer, V.C.F. (1955) Katalog der Araneae von 1758 bis 1940, bzw. 1954. 2. Band, Abt. a (Lycosaeformia, Dionycha [excl. Salticiformia]). 2. Band, Abt. b (Salticiformia, Cribellata) (Synonyma-Verzeichnis, Gesamtindex). Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 1751 pp.
  88. Roewer, V.C.F. (1960) Araneae Lycosaeformia II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l’Upemba, Mission G. F. de Witte, 55, 519–1040.
  89. Roth, V.D. & Brown, W.L. (1976) A new wolf spider from the Gulf of California beaches. Journal of the Arizona Academy of Science, 11 (2), 61–63.
  90. Saunier, F. (1980) Catalogue des collections arachnologiques mondiales. C.I.D.A., Paris, 70 pp.
  91. Sankaran, P.M., Caleb, J.T.D. & Sebastian, P.A. (2021) Revision of Indian wolf spiders: I. Genus Arctosa C.L. Koch, 1847 (Araneae: Lycosidae, Tricassinae). Zootaxa, 4908 (4), 489–504. https://doi.org/10.11646/zootaxa.4908.4.3
  92. Schmidt, M.H., Thies, C., Nentwig, W. & Tscharntke, T. (2008) Contrasting Responses of Arable Spiders to the Landscape Matrix at Different Spatial Scales. Journal of Biogeography, 35 (1), 157–166. https://doi.org/10.1111/j.1365-2699.2007.01774.x
  93. Sierwald, P. (2000) Description of the Male of Sosippus placidus, with notes on the subfamily Sosippinae (Araneae, Lycosidae). The Journal of Arachnology, 28 (2), 133–140. https://doi.org/10.1636/0161-8202(2000)028[0133:DOTMOS]2.0.CO;2
  94. Silva, E.L.C. da & Lise, A.A. (2009) A new species of the wolf spider genus Arctosa (Araneae, Lycosidae) from Southern Brazil. Revista Peruana de Biologia, 15 (2), 27–32. https://doi.org/10.15381/rpb.v15i2.1698
  95. Simon, E. (1876) Les arachnides de France. Vol. 3. Roret, Paris, 364 pp.
  96. Simon, E. (1885) Etudes sur les Arachnides recueillis en Tunisie en 1883 et 1884 par MM. A. Letourneux, M. Sédillot et Valéry Mayet, membres de la mission de l’Exploration scientifique de la Tunisie. Imprimerie Nationale, Paris, 55 pp.
  97. Simon, E. (1889) Etudes arachnologiques. 21e Mémoire. XXX. Descriptions de quelques arachnides du Chili et remarques synonymiques sur quelques unes des espèces décrites par Nicolet. Annales de la Société Entomologique de France, 8 (6), 217–222.
  98. Simon, E. (1910) Arachnoidea: Araneae (II.). In: Schultze, L. (Ed.), Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Südafrika. Vierter Band. Systematik und Tiergeographie. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena. Vol. 16. Verlag von Gustav Fisher, Jena, pp. 175–218.
  99. Simon, E. (1937) Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l’ordre des Araneae. Tome VI. 5e et derniére partie. Roret, Paris, pp. 979–1298.
  100. Strand, E. (1909) Neue oder wenig bekannte amerikanische Lycosiden aus der Sammlung des vestorbenen Mr Thomas Workman. Zeitschrift für Naturwissenschaften, 81, 277–286.
  101. Sundevall, C.J. (1833) Conspectus Arachnidum. C. F. Berling, Londini Gothorum [Lund], 39 pp.
  102. Tanaka, H. (1985) Descriptions of New Species of the Lycosidae (Araneae) from Japan. Acta Arachnologica, 33 (2), 51–87. https://doi.org/10.2476/asjaa.33.51
  103. Tanaka, H. (1991) Lycosid spiders of Japan VII. The genus Arctosa C. L. Koch. Sonoda Women's College Studies, 25, 289–316.
  104. Tanaka, H. (2009) Lycosidae. In: Ono, H. (Ed.), The Spiders of Japan with keys to the families and genera and illustrations of the species. Tokai University Press, Kanagawa, pp. 222–248.
  105. Thorell, T. (1875) Verzeichniss südrussischer spinnen. Horae Societatis Entomologicae Rossicae, 11, 39–122.
  106. Tullgren, A. (1905) Araneida from the swedish expedition trough the Gran Chaco and the Cordilleras. Arkiv för Zoologi, 2 (19), 1–81.
  107. Vink, C.J., Mitchell, A.D. & Paterson, A.M. (2002) A Preliminary Molecular Analysis of Phylogenetic Relationships of Australasian Wolf Spider Genera (Araneae, Lycosidae). The Journal of Arachnology, 30 (2), 227–237. https://doi.org/10.1636/0161-8202(2002)030[0227:APMAOP]2.0.CO;2
  108. Wang, L.-Y., Marusik, Y.M. & Zhang, Z.-S. (2012) Notes on three poorly known Arctosa species from China (Araneae: Lycosidae). Zootaxa, 3404 (1), 53–68. https://doi.org/10.11646/zootaxa.3404.1.2
  109. World Spider Catalog (2023) World Spider Catalog. Version 24.0. Natural History Museum Bern, Bern. Available from: http://wsc.nmbe.ch (accessed 23 July 2023) https://doi.org/10.24436/2
  110. Wunderlich, J. (1984) Zu Taxonomie und Determination europäischer spinnen-gattungen. 1. Wolfspinnen (Lycosidae) (Arachnida: Araneae). Neue Entomologische Nachrichten, 7, 21–29.
  111. Yoo, J.-S., Framenau, V.W. & Kim, J.-P. (2007) Arctosa stigmosa and A. subamylacea are two different species (Araneae, Lycosidae). The Journal of Arachnology, 35 (1), 171–180. https://doi.org/10.1636/h06-24.1
  112. Zehethofer, K. & Sturmbauer, C. (1998) Phylogenetic relationships of Central European wolf spiders (Araneae: Lycosidae) inferred from 12S ribosomal DNA sequences. Molecular Phylogenetics and Evolution, 10 (3), 391–398. https://doi.org/10.1006/mpev.1998.0536
  113. Zyuzin, A.A. (1990) Studies on burrowing spiders of the family Lycosidae (Araneae). I. Preliminary data on structural and functional features. Acta Zoologica Fennica, 190, 419–422.