Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5433 No. 1: 4 Apr. 2024
Type: Article
Published: 2024-04-04
DOI: 10.11646/zootaxa.5433.1.2
Page range: 51-95
Abstract views: 54
PDF downloaded: 0

An updated checklist for the pseudoscorpion fauna of Hungary (Arachnida: Pseudoscorpiones)

Eötvös Loránd University; Department of Systematic Zoology and Ecology; Pázmány Péter sétány 1/C; H–1117 Budapest; Hungary
Arachnida Pseudoscorpiones Central Europe false scorpion zoological collection taxonomy

Abstract

An updated checklist of the pseudoscorpion fauna of Hungary is presented. Chthonius cf. alpicola Beier, 1951 is newly reported for the country. Neobisium inaequale Chamberlin, 1930 is regarded as a junior synonym of N. sylvaticum (C.L. Koch, 1835) (syn. nov.). Chelifer tegulatus Tömösváry, 1883 is regarded as a junior synonym of Dactylochelifer latreillii (Leach, 1817) (syn. nov.). Rhacochelifer quadrimaculatus (Tömösváry, 1882) is regarded as a senior synonym of Beierochelifer peloponnesiacus (Beier, 1929) (syn. nov.), forming the combination Beierochelifer quadrimaculatus (Tömösváry, 1882) (comb. nov). Furthermore, Chthonius pygmaeus (Beier, 1934), Neobisium (Neobisium) simoni (L. Koch, 1873), Neobisium (Neobisium) validum (L. Koch, 1873), Rhacochelifer peculiaris (L. Koch, 1873), and Chernes montigenus (Simon, 1879) are excluded from the Hungarian fauna. Neobisium (N.) seminudum (Daday and Tömösváry, 1880) and Chelifer entzii Daday and Tömösváry, 1880 are designated as nomina dubia. As a consequence, the pseudoscorpion fauna of Hungary now comprises 49 species belonging to eight families with confirmed occurrence data; the occurrence of 6 further species recorded earlier for Hungary is doubtful. Phoresy is documented for Ephippiochthonius tetrachelatus (Preyssler, 1790) and Diplotemnus balcanicus (Redikorzev, 1928). Habitat preference of each species with confirmed occurrence in Hungary are assumed. Notes are given to some old data reported by Ödön Tömösváry and Eugen Daday from neighbouring countries. Microbisium manicatum (L. Koch, 1873) is excluded from the fauna list of Romania.

References

  1. Beier, M. (1928) Die Pseudoscorpione des Wiener Naturhistorischen Museums. I. Hemictenodactyli. Annalen des Naturhistorischen Museums in Wien, 42, 285–314.
  2. Beier, M. (1932a) Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich, 57, i–xx + 1–258. https://doi.org/10.1515/9783111435107.1
  3. Beier, M. (1932b) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, i–xxi + 1–294. https://doi.org/10.1515/9783111385402
  4. Beier, M. (1934) Neue cavernicole und subterrane Pseudoscorpione. Mitteilungen über Höhlen- und Karstforschung, 1934, 53–59.
  5. Beier, M. (1935) Drei neue Pseudoscorpione aus Rumänien. Bulletin de la Section Scientifique de l’Académie Roumaine, 17, 31–34.
  6. Beier, M. (1939a) Pseudoscorpionidea de Roumanie. Bulletin du Musée Royal d’Histoire Naturelle de Belgique, 15 (39), 1–21.
  7. Beier, M (1939b) Die Pseudoscorpione des oberösterreichischen Landesmuseums in Linz. Jahrbuch des Vereines für Landeskunde und Heimatpflege im Gau Oberdonau, Linz, 88, 303–312.
  8. Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296-301.
  9. Beier, M. (1951) Zur Kenntnis der ostalpinen Chthoniiden (Pseudoscorp). Entomologische Nachrichtblatt, Wien, 3, 163–166.
  10. Beier, M. (1956) Bemerkenswerte Pseudoscorpioniden-Funde aus Niederösterreich. Entomologisches Nachrichtenblatt, Wien, 8, 24–25.
  11. Beier, M. (1963a) Ordnung Pseudoscorpionidea (A­­f­ter­skorpione). Bestimmungsbücher zur Boden­fauna Europas. Vol. 1. Akademie-Verlag, Berlin, 313 pp.
  12. Beier, M. (1963b) Pseudoscorpione aus dem Museum ‘Enrico Caffi’ in Bergamo. Rendiconti dell’ Instituto Lombardo di Scienze e Lettere, 97B, 147–156.
  13. Britz, R., Hundsdörfer, A. & Fritz, U. (2020) Funding, training, permits—the three big challenges of taxonomy. Megataxa, 1 (1), 49–52. https://doi.org/10.11646/megataxa.1.1.10
  14. Canestrini, G. (1876) Osservazioni aracnologiche. Atti della Società veneto-trentina di scienze naturali resindente in Parova, 3, 206–232.
  15. Červená, M., Christophoryová, J., Purkart, A. & Krajčovičová, K. (2018) Allochernes solarii (Pseudoscorpiones: Chernetidae) newly recorded from ant nests in Slovakia. Arachnologische Mitteilungen: Arachnologische Letters, 56 (1), 40–44. https://doi.org/10.30963/aramit5607
  16. Červená, M., Krajčovičová, K. & Christophoryová, J. (2020a) An updated checklist of pseudoscorpions (Arachnida: Pseudoscorpiones) of Central Europe. Arthropoda Selecta, 29 (2), 219–228. https://doi.org/10.15298/arthsel.29.2.06
  17. Červená, M., Krajčovičová, K. & Christophoryová, J. (2020b) Pseudoscorpions (Arachnida: Pseudoscorpiones) in the nests of Formica ants in Slovakia. Klapalekiana, 56, 205–212.
  18. Červená, M., Gardini, G., Jablonski, D. & Christophoryová, J. (2021a) Checklist of pseudoscorpions (Arachnida, Pseudoscorpiones) of Albania. Zoological Studies, 60 (17), 1–25. https://doi.org/10.6620/ZS.2021.60-17
  19. Červená, M., Slenkovič, D. & Christophoryová, J. (2021b) New records of chernetid and cheliferid species (Arachnida: Pseudoscorpiones) from North North Macedonia. Natura Croatica, 30 (1), 251–256. https://doi.org/10.20302/NC.2021.30.16
  20. Chamberlin, J.C. (1929) A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part 1. The Heterosphyronida (Chthoniidae) (Arachnida-Chelonethida). Annals and Magazine of Natural History, Series 10, 4, 50–80. https://doi.org/10.1080/00222932908673028
  21. Chamberlin, J.C. (1930) A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part II. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History, Series 10, 5, 1–48 + 585–620. https://doi.org/10.1080/00222933008673173
  22. Christophoryová, J. (2013) A faunistic study on the pseudoscorpions (Arachnida: Pseudoscorpiones) of oak-hornbeam forests in SW Slovakia. Munis Entomology and Zoology, 8, 634–645.
  23. Christophoryová, J. & Krumpál, M. (2005) Communities of pseudoscorpions (Pseudoscorpiones, Arachnida) in epigeon of oak-hornbeam forests in the Malé Karpaty Mts and Trnavská pahorkatina hills (SW Slovakia). Ekológia, Bratislava, 24 (2), 76–86.
  24. Christophoryová, J., Fenďa, P. & Krištofík, J. (2011c) Chthonius hungaricus and Larca lata new to the fauna of Slovakia (Pseudoscorpiones: Chthoniidae, Larcidae). Arachnologische Mitteilungen, 41, 1–6. https://doi.org/10.5431/aramit4101
  25. Christophoryová, J., Gardini, G. & Korenko, S. (2011e) Allochernes powelli new to the fauna of Slovakia (Pseudoscorpiones: Chernetidae). Folia Faunistica Slovaca, 16, 67–70.
  26. Christophoryová, J., Krumpálová, Z., Krištofík, J. & Országhová, Z. (2011d) Association of pseudoscorpions with different types of birds nests. Biologia, 66 (4), 669–677. https://doi.org/10.2478/s11756-011-0072-8
  27. Christophoryová, J., Mock, A. & Ľuptáčik, P. (2011a) Chthonius (Chthonius) carinthiacus and Chthonius (Ephippiochthonius) tuberculatus new to the fauna of Slovakia (Pseudoscorpiones: Chthonii­dae). Arachnologische Mitteilungen, 42, 23–28. https://doi.org/10.5431/aramit4206
  28. Christophoryová, J., Šťáhlavský, F. & Fedor, P. (2011b) An updated identification key to the pseudoscorpions (Arachnida: Pseudoscorpiones) of the Czech Republic and Slovakia. Zootaxa, 2876 (1), 35–48. https://doi.org/10.11646/zootaxa.2876.1.4
  29. Christophoryová, J., Gardini, G. & Énekesová, E. (2012) Withius hispanus new to the pseudoscorpion fauna of Slovakia (Pseudoscorpiones: Withiidae). Arachnologische Mitteilungen, 44, 10–13. https://doi.org/10.5431/aramit4403
  30. Christophoryová, J., Grula, D. & Krajčovičová, K. (2017b) New records of pseudoscorpions (Arachnida: Pseudoscorpiones) associated with animals and human habitats in Slovakia and the Czech Republic. Arachnologische Mitteilungen, 53, 67–76. https://doi.org/10.5431/aramit5311
  31. Christophoryová, J., Jajcayová, D. & Krajčovičová, K. (2017c) Pseudoscorpions (Arachnida: Pseudoscorpiones) living in tree microhabitats in Slovakia. Klapalekiana, 53, 283–297. https://doi.org/10.5431/aramit5311
  32. Christophoryová, J., Kaňuchová, K. & Krajčovičová, K. (2017a) Faunistic survey of pseudoscorpions (Arachnida: Pseudoscorpiones) collected from compost heaps in Slovakia. Klapalekiana, 53, 11–19. https://doi.org/10.5431/aramit5304
  33. Christophoryová, J. & Červená, M. (2020) Apocheiridium ferum (Simon, 1879) (Arachnida, Pseudoscorpiones, Cheiridiidae), a newly recorded genus and species of pseudoscorpion for Hungary. Check List, 16, 223–228. https://doi.org/10.15560/16.2.223
  34. Christophoryová, J., Krajčovičová, K., Šťáhlavský, F., Španiel, S. & Opatova, V. (2023) Integrative taxonomy approach reveals cryptic diversity within the phoretic pseudoscorpion genus Lamprochernes (Pseudoscorpiones: Chernetidae). Insects, 14, 122. https://doi.org/10.3390/insects14020122
  35. Ćurčić, B.P.M. (1974) Arachnoidea. Pseudoscorpiones. In: Catalogus Faunae Jugoslaviae. 3 (4). Academie Slovéne, Ljubljana, pp. 1–35.
  36. Ćurčić, B.P.M., Poinar Jr., G.O. & Sarbu, S.M. (1993) New and little-known species of Chthoniidae and Neobisiidae (Pseudoscorpiones, Arachnida) from the Movile Cave in southern Dobrogea, Romania. Bijdragen tot de Dierkunde, 63, 221–241. https://doi.org/10.1163/26660644-06304002
  37. Daday, E. (1880) A magyarországi álskorpiók. Értesítő az Erdélyi Múzeum-egyesület Orvos-Természet-Tudományi Szakosztályából, 5, 191–193.
  38. Daday, E. (1889a) A Magyar Nemzeti Muzeum ál­skorpióinak áttekintése. Természetrajzi Füzetek, 11, 111–136 + 165–192.
  39. Daday, E. (1889b) Ujabb adatok a magyar-fauna ál­skorpióinak ismeretéhez. Természetrajzi Füzetek, 12, 25–28.
  40. Daday, E. (1918) Ordo Pseudoscorpiones. In: A Magyar Birodalom Allatvilága. Regia Societas Scientiarum Naturalium Hungarica, Budapest, pp. 1–2.
  41. Dashdamirov, S. (1999) New records of false scorpions in the Caucasus (Arachnida: Pseudoscorpiones). Arthropoda Selecta, 8, 79–87.
  42. Ducháč, V. & Mlejnek, R. (2000) Records of the pseudoscorpion Neobisium (Blothrus) slovacum (Neobisiidae) in caves and chasms of the Slovak Karst. In: Mock, A., Kováč, L. & Fulín, M. (Eds), Fauna jaskýň (Cave fauna): Proceedings of the workshop 20–21 October, 1999 East Slovakian Museum in Košice. East Slovakian Museum, Košice, pp. 15–20.
  43. Ebach, M.C., Valdecasas, A.G. & Wheeler, Q.D. (2011) Impediments to taxonomy and users of taxonomy: accessibility and impact evaluation. Cladistics, 27, 550–557. https://doi.org/10.1111/j.1096-0031.2011.00348.x
  44. Fabricius, J.C. (1793) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, obervationibus, descriptionibus. Vol. 2. C.G. Proft, Hafniae, viii + 519 pp. https://doi.org/10.5962/bhl.title.125869
  45. Farkas, S., Kárpáthegyi, P., Kiss, M., Novák, J. & Ujvári, Zs. (2009) Adatok a Zselic talajlakó mezo- és makrofaunájának ismeretéhez (Nematoda, Pseudoscorpiones, Acari, Chilopoda, Isopoda). Natura Somogyiensis, 13, 57–72. https://doi.org/10.24394/NatSom.2009.13.57
  46. Frivaldszky, J. (1865) Adatok a magyarhoni barlangok faunájához. Mathematikai és Természettudományi Közlemények, Vonatkozólag a Hazai Viszonyokra, 3, 17–53.
  47. Gardini, G. (1980). Identita’ di Chthonius tetrachelatus fuscimanus Simon, 1900 e redescrizione di C. (E.) nanus Beier, 1953. (Pseudoscorpionida Chthoniidae) (Pseudoscorpioni d’Italia IX). Annali del Museo Civico di Storia Naturale di Genova, 83, 261–270.
  48. Gardini, G. (1983) Redescription of Roncus lubricus L. Koch, 1873, type-species of the genus Roncus L. Koch, 1873 (Pseudoscorpionida, Neobisiidae). Bulletin of the British Arachnological Society, 6, 78–82.
  49. Gardini, G. (2009) Chthonius (C.) delmastroi n. sp. delle Alpi occidentali e del Piemonte e ridescrizione di Chthonius (C.) tenuis L. Koch, 1873 e di C. (C.) submontanus Beier, 1963 (Pseudoscorpiones Chthoniidae). Rivista Piemontese di Storia Naturale, 30, 25–51.
  50. Gardini, G. (2014) The species of the Chthonius heterodactylus group (Arachnida, Pseudoscorpiones, Chthoniidae) from the eastern Alps and the Carpathians. Zootaxa, 3887 (2), 101–137. https://doi.org/10.11646/zootaxa.3887.2.1
  51. Gardini, G. (2021) The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries. Zootaxa, 4987 (1), 1–131. https://doi.org/10.11646/zootaxa.4987.1.1
  52. Gulička, J. (1977) Neobisium (Blothrus) slovacum sp. n., eine neue Art des blinden Höhlenafterskorpions aus der Slowakei (Pseudoscorpionida). Annotationes Zoologicae et Botanicae, 117, 1–9.
  53. Hansen, H.J. (1884) Arthrogastra Danica: en monographisk fremstillning af de Danmark levende Meiere og Mosskorpioner med bidrag til sidstaevente underorders systematik. Natuurwetenschappelijk Tijdschrift, Series 3, 14, 491–554.
  54. Hadži, J. (1937) Pseudoscorpioniden aus Südserbien. Glasnik Skopsog Naucnog Drustva, 17, 151–187.
  55. Harvey, M.S. (1991) Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester and New York, 726 pp.
  56. Harvey, M.S. (2013) Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth. Available from: http://museum.wa.gov.au/catalogues-beta/pseudoscorpions (accessed 1 December 2023)
  57. Harvey, M.S. (2014) A review and redescription of the cosmopolitan pseudoscorpion Chelifer cancroides (Pseudoscorpiones: Cheliferidae). Journal of Arachnology, 42, 86–104. https://doi.org/10.1636/K13-57.1
  58. Harvey, M.S., Gardini, G. & Novák, J. (2018) Case 3736—Garypus latus Hansen, 1884 (currently Larca lata; Arachnida, Pseudoscorpiones): proposed precedence over Garypus hungaricus Tömösváry, 1882. Bulletin of Zoological Nomenclature, 75, 16–20. https://doi.org/10.21805/bzn.v75.a007
  59. Hermann, J.F. (1804) Mémoire Aptérologique. F. G. Levrault, Strasbourg, 144 pp. https://doi.org/10.5962/bhl.title.137910
  60. Hernández-Corral, J., Zaragoza, J.A. & Micó, E. (2018) New species of Pseudoscorpiones (Arachnida) from tree hollows in a Mediterranean oak forest in Spain. Zootaxa, 4497 (2), 201– 225. https://doi.org/10.11646/zootaxa.4497.2.3
  61. Horváth, G. (1884): Álskorpiók növényeken. Rovartani Lapok, 2 (1), 1–2.
  62. International Committee of Zoological Nomenclature (2020) Opinion 2462 (Case 3736) – Garypus latus Hansen, 1884 (currently Larca lata; Arachnida, Pseudoscorpiones): specific name conserved. Bulletin of Zoological Nomenclature, 77, 138–139. https://doi.org/10.21805/bzn.v77.a044
  63. Jászayová, A. & Jászay, T. (2022) New data on pseudoscorpions (Arachnida: Pseudoscorpiones) in north-east Slovakia. Arachnologische Mitteilungen, 63, 30–38. https://doi.org/10.30963/aramit6308
  64. Judson, M.L.I. (1990) On the presence of Chthonius (C.) halberti Kew and Chthonius (C.) ressli Beier in France with remarks on the status of Kewochthonius Chamberlin and Neochthonius Chamberlin (Arachnida, Chelonethida, Chthoniidae). Bulletin du Muséum National d’Histoire Naturelle, Paris, 11 (4), 593–603. https://doi.org/10.5962/p.288284
  65. Just, P., Šťáhlavský, F., Bogusch, P., Astapenková, A. & Opatova, V. (2023) Dispersal Capabilities Do Not Parallel Ecology and Cryptic Speciation in European Cheliferidae Pseudoscorpions (Pseudoscorpiones: Cheliferidae). Diversity, 15, 1040. https://doi.org/10.3390/d15101040
  66. Kaňuchová, A., Christophoryová, J. & Krajčovičá, K. (2015) Pseudoscorpions (Arachnida) collected from the heaps with decomposing material in Slovakia. Fragmenta Faunistica, 58 (2), 111–122. https://doi.org/10.3161/00159301FF2015.58.2.111
  67. Kárpáthegyi, P. (2005) Neobisidae család (Pseudoscorpions) fajainak előfordulásai Borsod-Abaúj-Zemplénmegyében. Folia Historico Naturalia Musei Matraensis, 29, 65–66.
  68. Kárpáthegyi, P. (2006) Két ritka álskorpió [Atemnus politus (Simon, 1878) és Chthonius heterodactylus Tömösvári, 1883] hazai előfordulásai. Folia Historico Naturalia Musei Matraensis, 30, 115–116.
  69. Kárpáthegyi, P. (2007) Pseudoscorpions of Hungary. Folia Historico Naturalia Musei Matraensis, 31, 81–90.
  70. Kárpáthegyi, P. & Kontschán, J. (2005) First record the Neobisium fuscimanum (C.L. Koch, 1843) in Hungary. Folia Entomologica Hungarica, 66, 5–6.
  71. Kew, H.W. (1916) A synopsos of the false-scorpions of Britain and Ireland; supplement. Proceedings of the Royal Irish Academy, 31 (38), 1–2.
  72. Koch, C.L. (1835) Deutschlands Crustaceen, Myriapoden und Arachniden. Vol. 2. Regensburg, F. Pustet, Regensburg, 90 pp.
  73. Koch, C.L. (1839) Uebersicht des Arachnidensystems. Vol. 2. Zweites Heft. C.H. Zeh, Nürnberg, 38 pp.
  74. Koch, C.L. (1843) Die Arachniden. Getreu nach der Natur Abgebildet und Beschrieben. Vol. 10. C. H. Zeh’schen Buchhandlung, Nürnberg, 220 pp.
  75. Koch, L. (1873) Uebersichtliche Dartstellung der Europäischen Chernetiden (Pseudoscorpione). Bauer & Raspe, Nürnberg, 80 pp.
  76. Krajčovičová, K., Christophoryová, J. & Krumpál, M. (2012) New faunistic and ecological data about pseudoscorpion Allochernes peregrinus Lohmander, 1939 from the Czech Republic (Pseudoscorpiones, Chernetidae). Folia Faunistica Slovaca, 17, 1–5.
  77. Krajčovičová, K. & Christophoryová, J. (2017) First record of Beierochelifer Mahnert, 1977 (Pseudoscorpiones: Cheliferidae) from Slovakia. Check List, 13 (2), 1–4. https://doi.org/10.15560/13.2.2074
  78. Krajčovičová, K., Christophoryová, K. & Mahnert, V. (2017) Rhacochelifer disjunctus (Pseudoscorpiones: Cheliferidae) new to the fauna of Slovakia. Arachnologische Mitteilungen, 53, 38–42. https://doi.org/10.5431/aramit5306
  79. Krajčovičová, K., Igondová, E., Majzlan, O. & Blažek, R. (2016) Pseudoscorpions (Arachnida: Pseudoscorpiones) of Malá Fatra National Park, Strážovské Vrchy Protected Landscape Area and Žilinská Kotlina Basin (Slovakia). Folia Faunistica Slovaca, 21, 1–7.
  80. Krajčovičová, K., Tamutis, V., Ivinskis, P., Machač, O. & Christophoryová, J. (2018) First records of pseudoscorpions (Arachnida: Pseudoscorpiones) from Lithuania. Entomologica Fennica, 29, 49–53. https://doi.org/10.33338/ef.71011
  81. Krumpál, M. & Krumpálová, Z. (2003) Šťúriky–Pseudoscorpiones. In: Mašán, P. & Svatoň, J. (Eds.), Pavúkovce Národného parku Poloniny (Arachnida: Araneae, Pseudoscorpiones, Opiliones, Acari–Parasitiformes). Štátna ochrana prírody SR Banská Bystrica a Správa Národného parku Poloniny Snina, pp. 115–126.
  82. Latifi, Z., Nassirkhani, M. & Mirshamsi, O. (2020) A new epigean pseudoscorpion species (Pseudoscorpiones: Neobisiidae) from northeast of Iran, with an identification key to the species of the family Neobisiidae from Iran. Turkish Journal of Zoology, 44, 230–237. https://doi.org/10.3906/zoo-1912-23
  83. Leach, M.D. (1817) On the characters of the genera of the family Scorpionidae, with descriptions of the British species of Chelifer and Obisium. In: The Zoological Miscellany; Being Descriptions of New or Interesting Animals. R. P. Nodder, London, pp. 48–53.
  84. Legg, G. & Jones, R.E. (1988) Synopses of the British fauna (new series). 40. Pseudoscorpions (Arthropoda; Arachnida). Brill/Backhuys, Leiden, 159 pp. https://doi.org/10.1163/9789004627642
  85. Lessert, R. de (1911) Pseudoscorpions. Catalogue des Invertébrées de la Suisse, 5, 1–50.
  86. Linnaeus, C. (1758): Systema naturae. Vol. 1. 10th Edition. L. Salvii, Holmiae, 824 pp.
  87. Lohmander, H. (1939) Zwei neue Chernetiden der nordwesteuropischen Fauna. Göteborgs Kungl. Vetenskaps- och Vitterhetssamhälles Handlingar, Series 5B, 6 (11), 1–11.
  88. Lohmander, H. (1945) Arachnologische Fragmente. 1. Über eine für die schwedische Fauna neue Pseudoskorpionart. Göteborgs Kungl. Vetenskaps- och Vitterhetssamhälles Handlingar, Series 6B, 3 (9), 3–14.
  89. Loksa, I. (1960) Faunistisch-systematische und ökologische Untersuchungen in der Lóczy-höhle bei Balatonfüred—Annales Universitatis Scientiarum Budapestiensis de Rolando Eötvös Nominatae: Sectio Biologica, 3, 253-266.
  90. Loksa, I. (1966) Die bodenzoozönolgischen Verhältnisse der Flaumeichen-Buschwälder Südostmitteleuropas. Akadémiai Kiadó, Budapest, pp. 437.
  91. Mahnert, V. (1980) Chthonius (C.) hungaricus sp. n., eine neue Afterscorpion-Art aus Ungarn (Arachnida). Folia Entomologica Hungarica, 33, 279–282.
  92. Mahnert, V. (1981) Mesochelifer ressli n. sp., eine mit Chelifer cancroides (L.) verwechselte Art aus Mitteleuropa (Pseudoscorpiones, Cheliferidae). Veröffentlichungen des Tirolor Landesmuseums Ferdinandeum, 61, 47–53.
  93. Mahnert, V. (1983) Pseudoscorpions of the Hortobágy National Park (Arachnida). In: Mahunka, S. (Ed.), The fauna of Hortobágy National Park 2. Akadémiai Kiadó, Budapest, pp. 361–363.
  94. Mahnert, V. (1988) Neobisium carcinoides (Hermann, 1804) (Pseudoscorpionida, Neobisiidae) – une espece polymorphe? Comptes Rendus Xème Colloque Europeen Arachnologie. Bulletin de la Société de Sciences de Bretagne, 59, 161–174.
  95. Mahnert, V. (1990) Pseudoscorpions of the Bátorliget Nature Reserve (NE Hungary). In: Mahunka, S. (Ed.), The Bátorliget Nature Reserves-after forty years 2. Hungarian Natural History Museum, Budapest, pp. 683–684.
  96. Mahnert, V. (2011) Pseudoscorpiones (Arachnida). In: Winkler, H. & Stuessy, T. (Eds.), Checklisten der Fauna Österreichs. No. 5. Verlag der Österreichischer Akademie der Wissenschaften, pp. 28–39.
  97. Mahnert, V. & Gardini, G. (2014) Cave-inhabiting pseudoscorpion species of the genus Roncus (Pseudoscorpiones: Neobisiidae) from western Greece, including the Ionian Islands. Arachnologische Mitteilungen, 48, 28–37. https://doi.org/10.5431/aramit4806
  98. Mock, A., Ľuptáčik, P., Fenďa, P. & Papáč, V. (2004) Biologická charakteristika jaskýň Bujanovských vrchov (Čierna hora). Aragonit, 9, 35-40.
  99. Murányi, D. & Kontschán, J. (2002) Pseudoscorpions from the Fertõ-Hanság National Park. In: Mahunka, S. (Ed.), The fauna of Fertõ-Hanság National Park 1. Hungarian Natural History Museum, Budapest, pp. 191–193.
  100. Muster, C., Korba, J., Bogusch, P., Heneberg, P. & Šťáhlavský, F. (2024) And yet they differ: reconsideration of diversity within Dactylochelifer latreillii (Arachnida: Pseudoscorpiones). Diversity, 16 (3), 137. https://doi.org/10.3390/d16030137
  101. Muster, C., Spelda, J., Rulik, B., Thormann, J., von der Mark, L & Astrin, J.J. (2021) The dark side of pseudoscorpion diversity: The German Barcode of Life campaign reveals high levels of undocumented diversity in European false scorpions. Ecology and Evolution, 11 (20), 13815–13829. https://doi.org/10.1002/ece3.8088
  102. Novák, J. (2011) Adatok a Bakony álskorpió-faunájához (Arachnida: Pseudoscorpiones). Folia Musei historico-naturalis Bakonyiensis, 28, 67–70.
  103. Novák, J. (2012). New records of pseudoscorpions for the fauna of the Bükk Mts., Northeast Hungary (Arachnida: Pseudoscorpiones). Opuscula Zoologica, Budapest, 43 (1), 57–65.
  104. Novák, J. (2013a) Adatok Magyarország álskorpió-faunkájához. Állattani Közlemények, 98 (1–2), 121–129.
  105. Novák, J. (2013b) First records of Larca lata (Hansen 1884) and Neobisium biharicum Beier 1939 from Hungary. Opuscula Zoolgica, Budapest, 44 (2), 161–166.
  106. Novák, J. (2014) Simontornya és környéke álskorpió faunája. In: Szita, É., Fetykó, K. & Kovács, T. (Eds.), Simontornya ízeltlábúi. In memoriam Pillich Ferenc. Magyar Biodiverzitás-kutató Társaság, Budapest, pp. 21–24.
  107. Novák, J. (2015) New records of the pseudoscorpion fauna of the Bakony Mts, Hungary (Arachnida: Pseudoscorpiones). Opuscula Zoologica, Budapest, 46 (2), 153–158. https://doi.org/10.18348/opzool.2015.2.153
  108. Novák, J. (2016) A Budai-hegység álskorpió-faunájának vizsgálata (Arachnida: Pseudoscorpiones). Állattani Közlemények, 101 (1–2), 3–10. https://doi.org/10.20331/AllKoz.2016.101.1-2.3
  109. Novák, J. (2017) Neobisium (N.) tothi sp. n. new species from Hungary and Romania, and first records of Neobisium (N.) noricum Beier, 1939 from Hungary (Pseudoscorpiones: Neobisiidae). Turkish Journal of Zoology, 41, 416–423. https://doi.org/10.3906/zoo-1607-19
  110. Novák, J. (2018) New data on the pseudoscorpion fauna of Hungary (Arachnida: Pseudoscorpiones). Acta Societatis Zoologicae Bohemicae, 82, 97–107.
  111. Novák, J. (2020) New data on the pseudoscorpion family Chthoniidae (Arachnida: Pseudoscorpiones) from Hungary. Opuscula Zoologica, Budapest, 51 (2), 115–121. https://doi.org/10.18348/opzool.2020.2.115
  112. Novák, J. & Kutasi, Cs. (2014) New data on the Pseudoscorpion fauna of the caves of the Bakony Mountains, Hungary. Opuscula Zoolgica, Budapest, 45 (2), 189–194.
  113. Novák, J. & Harvey, M.S. (2015) The identity of oseudoscorpions of the genus Diplotemnus (Pseudoscorpiones: Atemnidae) from Europe and Asia. North-Western Journal of Zoology, 11 (2), 316–323.
  114. Novák, J. & Hörweg, C. (2017) Redescriptions of three species of the genus Neobisium J. C. Chamberlin, 1930 from the Balkans and the Carpathians (Arachnida: Pseudoscorpiones). Acta Zoologica Academiae Scientiarum Hungaricae, 63 (3), 277–292. https://doi.org/10.17109/AZH.63.3.277.2017
  115. Novák, J. & Dányi, L. (2018) Catalogue of the type material of pseudoscorpions (Arachnida: Pseudoscorpiones) deposited in the Hungarian Natural History Museum, Budapest. Zootaxa, 4527 (3), 301–322. https://doi.org/10.11646/zootaxa.4527.3.1
  116. Novák, J., Jablonsky, D. & Christophoryová, J. (2019) Contribution to the pseudoscorpion fauna of Transylvania and the Eastern and Southern Carpathians, Romania (Arachnida: Pseudoscorpiones). North-Western Journal of Zoology, 15 (2), 127–134.
  117. Pavesi, P. (1884) Materiali per lo studio della fauna Tunisina raccolti da G. e L. Doria. Annali del Museo Civico di Storia Naturale di Genova, 20, 446–486.
  118. Pillich, F. (1914) Aus der Arthropodenwelt Simontornya’s. Simontornya Hungaria occidentalis. Komitat Tolna, 153 pp.
  119. Preyssler, J.D. (1790) Verzeichniss Böhmischer Insekten. in der von Schönfeld-Meißnerischen Buchhandlung, Prague, 108 pp.
  120. Rafalski, J. (1936) Neobisium (Neobisium) polonicum nov. spec. nowy gatunek zaleszczotka (Pseudoscorpionidea). Sprawozdania Poznańskiego Towarzystwa Przyjaciół Nauk, 9, 119.
  121. Rafalski, J. (1948) Mundochthonius carpaticus sp. nov., nowy gatunek zaleszczotka (Pseudoscorpionidea). Annales Musei Zoologici Polonici, 14, 13–20.
  122. Redikorzev, V. (1928) Beiträge zur Kenntnis der Pseudoscorpionenfauna Bulgariens. Mitteilungen aus dem Königl. Naturwissenschaftlichen Institut in Sofia, 1, 118–141.
  123. Roewer, C.F. (1937) Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (Ed.), Bronn’s Klassen und Ordnungen des Tierreichs. Bd. 5, Abt. IV: Insecta. 6: Pt. 1. Akademische Verlagsgesellschaft M.B.H., Leipzig, pp. 161–320.
  124. Růžička, A., Hajer, J. & Zacharda, M. (1995) Arachnid population patterns in underground cavities of a stony debris field (Araneae, Opiliones, Pseudoscorpionidea, Acari: Prostigmata, Rhagidiidae). Pedobiologia, 39, 42–51. https://doi.org/10.1016/S0031-4056(24)00162-8
  125. Šťáhlavský, F., Tůmová, P. & Král, J. (2003) Karyotype analysis in central European pseudoscorpions of the genus Neobisium (Pseudoscorpiones: Neobisiidae). In: Kipyatkov, V.E. & Logunov, D.V. (Eds.), 21st European Colloquium of Arachnology, St. Petersburg, 4–9 August 2003, Abstract book, pp. 80.
  126. Šťáhlavský, F. (2006) Štírci (Arachnida: Pseudoscorpiones) Národního parku Podyjí. Klapalekiana, 42, 167–178.
  127. Schawaller, W. (1989) Pseudoskorpione aus der Sowjetunion, Teil 3 (Arachnida: Pseudoscorpiones). Stuttgarter Beiträge zur Naturkunde, Series A, 440, 1–30.
  128. Schrank, F.P. (1803) Fauna Boica. Durchgedachte, Geschichte der in Baiern einheimischen und Zahmen Thiere. Vol. 3. Krüll, Landshut. [unknown pagination]
  129. Simon, E. (1878) Liste des espèces de la familie des Cheliferidae qui habitant l’Algérie et le Maroc. Annales de la Société Entomologique de France, Series 5, 8, 144–153.
  130. Simon, E. (1879) Les Arachnides de France. Vol. 7. les Ordres des Chernetes, Scorpiones et Opiliones. Librairie Encyclopédique de Rore, Paris, 332 pp.
  131. Simon, E. (1885) Étude sur les Arachnides recueillis en Tunisie en 1883 et 1884 par Mm. A. Letourneux, M. Sédillot et Valery Mayet. In: Exploration Scientifique de la Tunisie Imprimerie Nationale, Paris, pp. 1–55.
  132. Simon, E. (1900) Studio sui Chernetes Italiani conservati nel Museo Civico di Genova. 11. Annali del Museo Civico di Storia Naturale di genova, Series 2, 20, 593–595.
  133. Szalay, L. (1932) Adatok az Aggteleki barlang Arachnoidea-faunájának ismeretéhez. Állattani Közlemények, 29, 15–33.
  134. Szalay, L. (1968) Pókszabásúak I. Magyarország Állatvilága (Fauna Hungariae) LXXXIX., 18. Akadémiai Kiadó, Budapest, 122 pp.
  135. Szent-Ivány, J. (1941) Neue Angabenzur Verbreitung der Pseudoscorpione im Karpatenbecken. Fragmenta Faunistica Hungarica, 4 (1–4), 85–90.
  136. Tömösváry, Ö. (1882a) A Magyar fauna álskorpiói. Magyar Tudományos Akadémia Matematikai és Természettudományi Közlemények, 18, 135–256.
  137. Tömösváry, Ö. (1882b) Egy új alak hazánk Arach­noida faunájában Zemplén megyéböl. Természe­trajzi Füzetek, 6, 226–228 + 296–298.
  138. Tooren, D. van der (2005) The subspecies of the pseudoscorpion Dactylochelifer latreillei in the Netherlands (Pseudoscorpiones: Cheliferidae). Nederlandse Faunistische Mededelingen, 23, 77–89.
  139. Turbanov, I.S. & Kolesnikov, V.B. (2023) First records of cave-dwelling pseudoscorpions of the genus Ephippiochthonius Beier, 1930 (Arachnida: Pseudoscorpiones: Chthoniidae) from the Caucasus, with description of three new species from Abkhazia and taxonomic comments on the genus. Zootaxa, 5325 (1), 41–62. https://doi.org/10.11646/zootaxa.5325.1.3
  140. Varga, Z. (1964) Zoogeographische Einteilung Ungarns auf Grung der Makrolepidopteren–Faunakomponenten. Folia Entomologica Hungarica, 17, 119–197.
  141. Verner, P.H. (1958) Eine neue Pseudoskorpionart aus der Tschechoslowakei. Zoologischer Anzeiger, 161 (7/8), 198–199.
  142. Wägele, H., Klussmann-Kolb, A., Kuhlmann, M., Haszprunar, G., Lindberg, D., Koch, A. & Wägele, J.G. (2011) The taxonomist—an endangered race. A practical proposal for its survival. Frontiers in Zoology, 8, 25. https://doi.org/10.1186/1742-9994-8-25.
  143. World Pseudoscorpiones Catalog (2023) World Pseudoscorpiones Catalog. Natural History Museum, Bern. Available from: https://wac.nmbe.ch/order/pseudoscorpiones/3 (accessed 1 December 2023)
  144. Zaragoza, J.A. (2017) Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae). Zootaxa, 4246 (1), 1–221. https://doi.org/10.11646/zootaxa.4246.1.1