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Published: 2024-05-09
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Phylogenetic position of Kinnareemimus khonkaenensis (Dinosauria: Theropoda: Ornithomimosauria) from the Lower Cretaceous of Thailand

Biodiversity and Conservation Research Unit; Walai Rukhavej Botanical Research Institute; Mahasarakham University; Maha Sarakham; Thailand; Dinosaur Research Unit; Mahasarakham University; Maha Sarakham; Thailand
Reptilia Ornithomimosauria Southeast Asia subarctometatarsalian metatarsus Early Cretaceous Sao Khua Formation paleobiogeography

Abstract

A small-bodied ornithomimosaur, Kinnareemimus khonkaenensis Buffetaut, Suteethorn, and Tong 2009, from the Lower Cretaceous Sao Khua Formation of northeastern Thailand was first reported in 1995, then named in 2009. It is the only report of this group in Southeast Asia and was concluded to be a member of Ornithomimosauria, more derived than Harpymimus and Garudimimus but more basal than Archaeornithomimus. Since then, a few published studies have analyzed the phylogenetic position of Kinnareemimus but failed to find it within the clade Ornithomimosauria. Here, the phylogenetic analyses for assessing the relationships of Kinnareemimus within Ornithomimosauria were performed. The results suggested it might be a basal ornithomimosaur or belongs to the subclade Deinocheiridae. Kinnareemimus shares with deinocheirids the proximally projected cnemial crest and the shape of the fibula in proximal view. Its metatarsal III shows subarctometatarsalian condition with metatarsal III pinched between II and IV but visible in anterior view proximally, which might have evolved independently from other ornithomimids. However, the basal position of Kinnareemimus could also be due to the immaturity and the incomplete nature of this animal.

 

References

  1. Alifanov, V.R. & Averianov, A.O. (2006) On the finding of ornithomimid dinosaurs (Saurischia, Ornithomimosauria) in the Upper Cretaceous beds of Tajikistan. Paleontological Journal, 40 (1), 103–108. https://doi.org/10.1134/S0031030106010126
  2. Alifanov, V.R. & Saveliev, S.V. (2015) The most ancient ornithomimosaur (Theropoda, Dinosauria), with cover imprints from the Upper Jurassic of Russia. Paleontological Journal, 49 (6), 636–650. https://doi.org/10.1134/S0031030115060039
  3. Apesteguía, S., Smith, N.D., Juárez Valieri, R. & Makovicky, P.J. (2016) An unusual new theropod with a didactyl manus from the Upper Cretaceous of Patagonia, Argentina. PLoS One, 11 (e0157793), 1–41. https://doi.org/10.1371/journal.pone.0157793
  4. Allain, R., Vullo, R., Le loeuff, J. & Tournepiche, J.-F. (2014) European ornithomimosaurs (Dinosauria, Theropoda): an undetected record. Geologica Acta, 12 (2), 127–135.
  5. Averianov, A.O., Sues, H.D., Dyke, G. & Bayshashov, B. (2017) Hind limb bones of an ornithomimid dinosaur from the Upper Cretaceous Bostobe Formation, northeastern Aral Sea region, Kazakhstan. Palaeoworld, 26 (1), 194–201. https://doi.org/10.1016/j.palwor.2016.01.003
  6. Barsbold, R. (1981) Bezzubyye khishchnyye dinozavry Mongolii, Sovmestnaia Sovetsko-Mongol’skaia Paleontologicheskaia Ekspeditsiia. Trudy, 15, 28–39.
  7. Barsbold, R. (1983) Carnivorous dinosaurs from the Cretaceous of Mongolia. Transactions, Joint Soviet Mongolian Palaeontological Expedition, 19, 1–120. [in Russian].
  8. Barsbold, R. (1988b) A new Late Cretaceous ornithomimid from the Mongolian People’s Republic. Paleontological Journal, 22, 124–127.
  9. Barsbold, R. & Perle, A. (1984) [On first new find of a primitive ornithomimosaur from the Cretaceous of the MPR]. Paleontologicheskii Zhurnal, 2, 121–123.
  10. Benson, R.B.J. (2010) A description of Megalosaurus bucklandii (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods. Zoological Journal of the Linnean Society, 158, 882–935. https://doi.org/10.1111/j.1096-3642.2009.00569.x
  11. Benson, R.B.J., Rich, T.H., Vickers-Rich, P. & Hall, M. (2012) Theropod fauna from Southern Australia indicates high polar diversity and climate-driven dinosaur provinciality. PloS ONE, 7 (5), e37122. https://doi.org/10.1371/journal.pone.0037122.
  12. Bonaparte, J.F. (1986) Les dinosaures Dinosaurs (Carnosaures, Allosauridés, Sauropodes, Cétosauridés) du Jurassique Moyen de Cerro Cóndor (Chubut, Argentina). Annales de Paléontologie (Vert.-Invert.), 72, 247–289.
  13. Brochu, C.A. (2003) Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull. Journal of vertebrate Paleontology, 22 (4), 1–138. https://doi.org/10.2307/3889334
  14. Brusatte, S.L. (2013) The phylogeny of basal coelurosaurian theropods (Archosauria: Dinosauria) and patterns of morphological evolution during the dinosaur-bird transition. PhD dissertation, Columbia University, New York, New York, 944 pp.
  15. Brusatte, S.L. and Benson, R.B.J. (2013) The systematics of Late Jurassic tyrannosauroid theropods from Europe and North America. Acta Palaeontologica Polonica, 58 (1), 47–54. https://doi.org/10.4202/app.2011.0141
  16. Brusatte, S.L., Lloyd, G.T., Wang, S.C. & Norell, M.A. (2014) Gradual assembly of avian body plan culminated in rapid rates of evolution across the dinosaur-bird transition. Current Biology, 24 (20), 2386–2392. https://doi.org/10.1016/j.cub.2014.08.034
  17. Buffetaut, E. & Suteethorn, V. (1999) The dinosaur fauna of the Sao Khua Formation of Thailand and the beginning of the Cretaceous radiation of dinosaurs in Asia. Palaeogeography, Palaeoclimatology, Palaeoecology, 150, 13–23. https://doi.org/10.1016/S0031-0182(99)00004-8
  18. Buffetaut, E., Suteethorn, V. & Tong, H. (2009) An early ‘ostrich dinosaur’ (Theropoda: Ornithomimosauria) from the Early Cretaceous Sao Khua Formation of NE Thailand. In: Buffetaut, E., Cuny, G., Le Loeuff, J. & Suteethorn, V. (Eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315. Geological Society, London, pp. 229–243. https://doi.org/10.1144/SP315.16
  19. Buffetaut, E., Suteethorn, V., Martin, V., Tong, H., Chaimanee, Y. & Triamwichanon, S. (1995) New dinosaur discoveries in Thailand. In: Proceedings of the International Conference on Geology, Geotechnology and Mineral Resources of Indochina. Department of Geotechnology, Khon Kaen University, Khon Kaen, pp. 157–161.
  20. Carrano, M.T., Benson, R.B. & Sampson, S.D. (2012) The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology, 10 (2), 211–300. https://doi.org/10.1080/14772019.2011.630927
  21. Cau, A. (2018) The assembly of the avian body plan: A 160-million-year long process. Bollettino della Società Paleontologica Italiana, 57 (1), 1–25.
  22. Cerroni, M.A., Agnolin, F.L., Egli, F.B. & Novas, F.E. (2019) The phylogenetic position of Afromimus tenerensis Sereno, 2017 and its paleobiogeographical implications. Journal of African Earth Sciences, 159, 103572. https://doi.org/10.1016/j.jafrearsci.2019.103572
  23. Cerroni, M.A., Baiano, M.A., Canale, J.I., Agnolín, F.L., Otero, A. & Novas, F.E. (2022) Appendicular osteology of Skorpiovenator bustingorryi (Theropoda, Abelisauridae) with comments on phylogenetic features of abelisaurids. Journal of Systematic Palaeontology, 20 (1), 1–32. https://doi.org/10.1080/14772019.2022.2093661
  24. Choiniere, J.N., Forster, C.A. & de Klerk, W.J. (2012) New information on Nqwebasaurus thwazi, a coelurosaurian theropod from the Early Cretaceous Kirkwood Formation in South Africa. Journal of African Earth Sciences, 71, 1–17. https://doi.org/10.1016/j.jafrearsci.2012.05.005
  25. Claessens, L.P.A.M. & Loewen, M.A. (2015) A redescription of Ornithomimus velox Marsh, 1890 (Dinosauria, Theropoda), Journal of Vertebrate Paleontology, 36 (1), e1034593. https://doi.org/10.1080/02724634.2015.1034593
  26. Cuesta, E., Vidal, D., Ortega, F., Shibata, M. & Sanz, J.L. (2022) Pelecanimimus (Theropoda: Ornithomimosauria) postcranial anatomy and the evolution of the specialized manus in Ornithomimosaurs and sternum in maniraptoriforms. Zoological Journal of the Linnean Society, 194 (2), 553–591. https://doi.org/10.1093/zoolinnean/zlab013
  27. De Klerk, W., Forster, C., Sampson, S., Chinsamy-Turan, A. & Ross, C. (2000) A new coelurosaurian dinosaur from the Early Cretaceous of South Africa. Journal of Vertebrate Paleontology, 20, 324–332. https://doi.org/10.1671/0272-4634(2000)020[0324:ANCDFT]2.0.CO;2
  28. Delcourt, R. & Grillo, O.N. (2018) Tyrannosauroids from the Southern Hemisphere: implications for biogeography, evolution, and taxonomy. Palaeogeography, Palaeoclimatology, Palaeoecology, 511, 379–387. https://doi.org/10.1016/j.palaeo.2018.09.003
  29. Ezcurra, M.D. (2017) A new Early Coelophysoid neotheropod from the Late Triassic of northwestern Argentina. Ameghiniana, 54, 506–538. https://doi.org/10.5710/AMGH.04.08.2017.3100
  30. Goloboff, P.A., Farris, J.S. & Nixon, K.C. (2008) TNT, a free program for phylogenetic analysis. Cladistics, 24, 774–786. https://doi.org/10.1111/j.1096-0031.2008.00217.x.
  31. Hartman, S., Mortimer, M., Wahl, W.R., Lomax, D.R., Lippincott, J. & Lovelace, D.M. (2019) A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight. PeerJ, 7, e7247. https://doi.org/10.7717/peerj.7247
  32. Hattori, S., Shibata, M., Kawabe, S., Imai, T., Nishi, H. & Azuma, Y. (2023) New theropod dinosaur from the Lower Cretaceous of Japan provides critical implications for the early evolution of ornithomimosaurs. Scientific Reports, 13, 13842. https://doi.org/10.1038/s41598-023-40804-3
  33. Holtz, T.R. (1994) The phylogenetic position of the Tyrannosauridae: implications for theropod systematic. Journal of Paleontology, 68 (5), 1100–1117. https://doi.org/10.1017/S0022336000026706
  34. Holtz Jr., T.R., Molnar, R.E. & Currie, P.J. (2004) Basal Tetanurae. In: Weishampel, D.B., Dodson, P. & Osmólska, H. (Eds.), The Dinosauria, Second Edition. University of California Press, Berkeley/Los Angeles/London, pp. 71–110. https://doi.org/10.1525/california/9780520242098.003.0006
  35. Huene, F., von (1920) Bemerkungen zur Systematik und Stammegeschichte einiger Reptilian. Zeitschrift f¨ur Induktive Abstammungs- und Vererbungslehre, 24, 209–212. https://doi.org/10.1007/BF01785394
  36. Huene, F., von (1923) Carnivorous Saurischia in Europe since the Triassic. Bulletin of the Geological Society of America, 34, 449–458. https://doi.org/10.1130/GSAB-34-449
  37. Hunt, R.K. & Quinn, J.H. (2018) A new ornithomimosaur from the Lower Cretaceous Trinity Group of Arkansas. Journal of Vertebrate Paleontology, 38 (1), e1421209. https://doi.org/10.1080/02724634.2017.1421209
  38. Ibrahim, N., Sereno, P.C., Dal Sasso, C., Maganuco, S., Fabbri, M., Martill, D.M., Zouhri, S., Myhrvold, N. & Iurino, D.A. (2014) Semiaquatic adaptations in a giant predatory dinosaur. Science, 345, 1613–1616. https://doi.org/10.1126/science.1258750
  39. Ji, Q., Norrell, M., Makovicky, P.J., Gao, K., Ji, S. & Yuan, C. (2003) An Early Ostrich Dinosaur and Implications for Ornithomimosaur Phylogeny. American Museum Novitates, 3420, 1–19. https://doi.org/10.1206/0003-0082(2003)420%3C0001:AEODAI%3E2.0.CO;2
  40. Jin, L., Chen, J. & Godefroit, P. (2012) A New Basal Ornithomimosaur (Dinosauria: Theropoda) from the Early Cretaceous Yixian Formation, Northeast China. In: Godefroit, P. (Ed.), Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems. Indiana University Press, Bloomington, Indiana, pp. 467–487.
  41. Kirkland, J.I., Britt, B.B., Whittle, C.H., Madsen, S.K. & Burge, D.L. (1998) A small coelurosaurian theropod from The Yellow Cat Member of the Cedar Mountain Formation (Lower Cretaceous, Barremian) of eastern Utah. New Mexico Museum of Natural History and Science Bulletin, 14, 239–248.
  42. Kobayashi, Y. & Barsbold, R. (2005a) Anatomy of Harpymimus okladnikovi Barsbold and Perle 1984 (Dinosauria; Theropoda) of Mongolia. In: Carpenter, K. (Ed.), The Carnivorous Dinosaurs. Indiana University Press, Bloomington, Indiana, pp. 97–126.
  43. Kobayashi, Y. & Barsbold, R. (2005b) Reexamination of a primitive ornithomimosaur, Garudimimus brevipes Barsbold, 1981 (Dinosauria:Theropoda), from the Late Cretaceous of Mongolia. Canadian Journal of Earth Sciences, 42, 150 1–1521. https://doi.org/10.1139/e05-044
  44. Kobayashi, Y. & Barsbold, R. (2006) Ornithomimids from the Nemegt Formation of Mongolia. Journal-Paleontological Society of Korea, 22 (1), 195–207.
  45. Kobayashi, Y. & Lü, J.C. (2003) A new ornithomimid dinosaur with gregarious habits from the Late Cretaceous of China. Acta Palaeontologica Polonica, 48 (2), 235–259.
  46. Kobayashi, Y., Tsogtobaatar, K., Kubota, K., Lee, Y., Lee, H. & Barsbold, R. (2014) New ornithomimid from the Upper Cretaceous Bayanshiree Formation of Mongolia. Annual meeting, Berlin, 2014, programm and abstract.
  47. Lacerda, M.B.S., Bittencourt, J.S. & Hutchinson, J.R. (2023) Macroevolutionary patterns in the pelvis, stylopodium and zeugopodium of megalosauroid theropod dinosaurs and their importance for locomotor function. Royal Society Open Science, 10, 230481. https://doi.org/10.1098/rsos.230481
  48. Langer, M.C., Rincón, A.D., Ramezani, J., Solórzano, A. & Rauhut, O.W. (2014) New dinosaur (Theropoda, stem-Averostra) from the earliest Jurassic of the La Quinta formation, Venezuelan Andes. Royal Society open science, 1 (2), 140184. https://doi.org/10.1098/rsos.140184
  49. Lee, Y.N., Barsbold, R., Currie, P.J., Kobayashi, Y., Lee, H.J., Godefroit, P., Escuillié, F. & Chinzorig, T. (2014) Resolving the long-standing enigmas of a giant ornithomimosaur Deinocheirus mirificus. Nature, 515 (7526), 257–260. https://doi.org/10.1038/nature13874
  50. Makovicky, P.J., Kobayashi, Y. & Currie, P.J. (2004) Ornithomimosauria. In: Weishampel, D.B., Dodson, P. & Osmólska, H. (Eds.), The Dinosauria, Second Edition. University of California Press, Berkeley/Los Angeles/London, pp. 137–150. https://doi.org/10.1525/california/9780520242098.003.0008
  51. Makovicky, P.J., Li, D., Gao, K.-Q., Lewin, M., Erickson, G.M. & Norell, M.A. (2010) A giant ornithomimosaur from the Early Cretaceous of China. Proceedings of the Royal Society B: Biological Sciences, 277 (1679), 191–198. https://doi.org/10.1098/rspb.2009.0236
  52. Marsh, O.C. (1881) Principal characters of American Jurassic dinosaurs, part V. American Journal of Science, 125, 417–423. https://doi.org/10.2475/ajs.s3-21.125.417
  53. Marsh, O.C. (1890) Description of new dinosaurian reptiles. American Journal of Science, 3 (39), 81–86. https://doi.org/10.2475/ajs.s3-39.229.81
  54. McFeeters, B., Ryan, M.J., Schröder-Adams, C. & Cullen, T.M. (2016) A new ornithomimid theropod from the Dinosaur Park Formation of Alberta, Canada. Journal of Vertebrate Paleontology, 36 (6), e1221415. https://doi.org/10.1080/02724634.2016.1221415
  55. McFeeters, B., Ryan, M.J., Schröder-Adams, C. & Currie, P.J. (2017) First North American occurrences of Qiupalong (Theropoda: Ornithomimidae) and the palaeobiogeography of derived ornithomimids. Facets, 2 (1), 355–373. https://doi.org/10.1139/facets-2016-0074
  56. McFeeters, B., Ryan, M.J. & Cullen, T.M. (2018) Morphology and Variation in Ungual Phalanges of North American Ornithomimosaurs (Dinosauria, Theropoda): Implications for Interpreting Isolated Elements. Vertebrate Anatomy Morphology Palaeontology, 6, 60–67. https://doi.org/10.18435/vamp29283
  57. Naoki, I. (2016) The first record of an ornithomimosaurian dinosaur from the Upper Cretaceous of Japan. Historical Biology, 28 (1–2), 264–269. https://doi.org/10.1080/08912963.2015.1025389
  58. Nesbitt, S.J. (2011) The early evolution of Archosauria: relationships and the origin of major clades. Bulletin of the American Museum of Natural History, 352, 1–292. https://doi.org/10.1206/352.1
  59. Nesbitt, S.J., Irmis, R.B. & Parker, W.G. (2007) A critical re-evaluation of the Late Triassic dinosaur taxa of North America. Journal of Systematic Palaeontology, 5, 209–243. https://doi.org/10.1017/S1477201907002040
  60. Novas, F.E., Dalla Vecchia, F. & Pais, D.F. (2005) Theropod pedal unguals from the Late Cretaceous (Cenomanian) of Morocco, Africa. Revista del Museo Argentino de Ciencias Naturales, Nueva Serie, 7, 167–175.
  61. Novas, F.E., Agnolin, F.L., Ezcurra, M.D., Porfiri, J. & Canale, J.I. (2013) Evolution of the carnivorous dinosaurs during the Cretaceous: The evidence from Patagonia. Cretaceous Research, 45, 174–215. https://doi.org/10.1016/j.cretres.2013.04.001
  62. Osborn, H.F. (1917) Skeletal adaptations of Ornitholestes, Struthiomimus, Tyrannosaurus. Bulletin of the American Museum of Natural History, 35, 733–771.
  63. Osmólska, H. & Roniewicz, E. (1970) Deinocheiridae, a new family of theropod dinosaurs. Palaeontologica Polonica, 21, 5–19.
  64. Osmólska, H., Roniewicz, E. & Barsbold, R. (1972) A new dinosaur, Gallimimus bullatus n. gen., n. sp. (Ornithomimidae) from the Upper Cretaceous of Mongolia. Palaeontologia Polonica, 27, 103–143.
  65. Owen, R. (1842) Report on British fossil reptiles, part II. Reports of the British Association for the Advancement of Science, 11, 60–204. [for 1841]
  66. Padian, K. & May, C.L. (1993) The earliest dinosaurs. New Mexico Museum of Natural History Science Bulletin, 3, 379–381.
  67. Perez-Moreno, B.P., Sanz, J.L., Buscalioni, A.D., Moratalla, J.J., Ortega, F. & Raskin-Gutman, D. (1994) A unique multitoothed ornithomimosaur dinosaur from the Lower Cretaceous of Spain. Nature, 370, 363–367. https://doi.org/10.1038/370363a0
  68. Porfiri, J.D., Novas, F.E., Calvo, J.O., Agnolin, F.L., Ezcurra, M.D. & Cerda, I.A. (2014) Juvenile specimen of Megaraptor (Dinosauria, Theropoda) sheds light about tyrannosauroid radiation. Cretaceous Research, 51, 35–55. https://doi.org/10.1016/j.cretres.2014.04.007
  69. Rauhut, O.W.M. (2003) The interrelationships and evolution of basal theropod dinosaurs. Special Papers in Palaeontology, 69, 1–213.
  70. Rauhut, O.W.M. (2005) Osteology and relationships of a new theropod dinosaur from the Middle Jurassic of Patagonia. Palaeontology, 48 (1), 87–110. https://doi.org/10.1111/j.1475-4983.2004.00436.x
  71. Rauhut, O.W.M. (2012) A reappraisal of a putative record of abelisauroid theropod dinosaur from the Middle Jurassic of England. Proceedings of the Geologists’ Association, 123 (5), 779–786. https://doi.org/10.1016/j.pgeola.2012.05.008.
  72. Rauhut, O.W.M. & Carrano, M.T. (2016) The theropod dinosaur Elaphrosaurus bambergi Janensch, 1920, from the Late Jurassic of Tendaguru, Tanzania. Zoological Journal of the Linnean Society, 178 (3), 546–610. https://doi.org/10.1111/zoj.12425
  73. Rich, T.H. & Vickers-Rich, P. (1994) Neoceratopsians and ornithomimosaurs: dinosaurs of Gondwanan origin? National Geographic Research and Exploration, 10, 129–131.
  74. Samathi, A. (2019) Theropod dinosaurs from Thailand and Southeast Asia: phylogeny, evolution, and paleobiogeography. PhD Thesis. Universitäts-und Landesbibliothek Bonn, Bonn, 249 pp.
  75. Samathi, A., Chanthasit, P. & Sander, P.M. (2019a) Two new basal coelurosaurian theropod dinosaurs from the Early Cretaceous Sao Khua Formation of Thailand. Acta Palaeontologica Polonica, 64 (2), 239–260. https://doi.org/10.4202/app.00540.2018
  76. Samathi, A., Chanthasit, P. & Sander, P.M. (2019b) A review of theropod dinosaurs from the Late Jurassic to mid-Cretaceous of Southeast Asia. Annales de Paléontologie, 105 (3), 201–215. https://doi.org/10.1016/j.annpal.2019.03.003.
  77. Senter, P. (2007) A new look at the phylogeny of Coelurosauria (Dinosauria: Theropoda). Journal of Systematic Palaeontology, 5 (4), 429–463. https://doi.org/10.1017/S1477201907002143.
  78. Sereno, P.C. (2017) Early Cretaceous ornithomimosaurs (Dinosauria: Coelurosauria) from Africa. Ameghiniana, 54 (5), 576–617. https://doi.org/10.5710/AMGH.23.10.2017.3155
  79. Sereno, P.C., Dutheil, D.B., Iarochene, M., Larsson, H.C., Lyon, G.H., Magwene, P.M., Sidor, C.A., Varricchio, D.J. & Wilson, J.A. (1996) Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation. Science, 272 (5264), 986–991. https://doi.org/10.1126/science.272.5264.986
  80. Serrano-Branas, C.I., Torres-Rodríguez, E., Reyes-Luna, P.C., González-Ramírez, I. & González-León, C. (2016) A new ornithomimid dinosaur from the Upper Cretaceous Packard Shale formation (Cabullona Group) Sonora, México. Cretaceous Research, 58, 49–62. https://doi.org/10.1016/j.cretres.2015.08.013
  81. Serrano-Brañas, C.I., Espinosa-Chávez, B., Maccracken, S.A., Gutiérrez-Blando, C., de León-Dávila, C. & Ventura, J.F. (2020) Paraxenisaurus normalensis, a large deinocheirid ornithomimosaur from the Cerro del Pueblo Formation (Upper Cretaceous), Coahuila, Mexico. Journal of South American Earth Sciences, 101, 102610. https://doi.org/10.1016/j.jsames.2020.102610
  82. Smith, D. & Galton, P. (1990) Osteology of Archaeornithomimus asiaticus (Upper Cretaceous, Iren Dabasu Formation, People’s Republic of China). Journal of Vertebrate Paleontology, 10 (2), 255–265. https://doi.org/10.1080/02724634.1990.10011811
  83. Sues, H.D. & Averianov, A. (2016) Ornithomimidae (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan. Cretaceous Research, 57, 90–110. https://doi.org/10.1016/j.cretres.2015.07.012
  84. Tsogtbaatar, C., Kobayashi, Y., Tsogtbaatar, K., Currie, P.J., Watabe, M. & Barsbold, R. (2017) First ornithomimid (Theropoda, Ornithomimosauria) from the Upper Cretaceous Djadokhta Formation of Tögrögiin Shiree, Mongolia. Scientific Reports, 7 (1), 5835. https://doi.org/10.1038/s41598-017-05272-6
  85. Tucker, R.T., Hyland, E.G., Gates, T.A., King, M.R., Roberts, E.M., Foley, E.K., Berndt, D., Hanta, R., Khansubha, S., Aswasereelert, W. & Zanno, L.E. (2022) Age, depositional history, and paleoclimatic setting of Early Cretaceous dinosaur assemblages from the Sao Khua Formation (Khorat Group), Thailand. Palaeogeography, Palaeoclimatology, Palaeoecology, 601, 111107. https://doi.org/10.1016/j.palaeo.2022.111107
  86. Vickers-Rich, P. & Rich, T.H. (1993) Australia’s polar dinosaurs. Scientific American, 269 (1), 50–55. https://doi.org/10.1038/scientificamerican0793-50
  87. White, M.A. (2009) The subarctometatarsus: intermediate metatarsus architecture demonstrating the evolution of the arctometatarsus and advanced agility in theropod dinosaurs. Alcheringa, 33 (1), 1–21.
  88. Xu, L., Kobayashi, Y., Lü, J., Lee, Y.N., Liu, Y., Tanaka, K., Zhang, X., Jia, S. & Zhang, J. (2011) A new ornithomimid dinosaur with North American affinities from the Late Cretaceous Qiupa Formation in Henan Province of China. Cretaceous Research, 32 (2), 213–222. https://doi.org/10.1016/j.cretres.2010.12.004
  89. Yao, X., Sullivan, C., Tan, Q. & Xu, X. (2022) New ornithomimosaurian (Dinosauria: Theropoda) pelvis from the Upper Cretaceous Erlian Formation of Nei Mongol, North China. Cretaceous Research, 137, 105234. https://doi.org/10.1016/j.cretres.2022.105234
  90. Zhao, X.-J., Benson, R.B., Brusatte, S.L. & Currie, P.J. (2010) The postcranial skeleton of Monolophosaurus jiangi (Dinosauria: Theropoda) from the Middle Jurassic of Xinjiang, China, and a review of Middle Jurassic Chinese theropods. Geological Magazine, 147 (1), 13–27. https://doi.org/10.1017/S0016756809990240