Abstract
The genus Antillocladius Sæther, 1981 currently comprises 17 described species. In the present paper, seven new Brazilian species, A. atalaia sp. n., A. axitiosus sp. n., A. brazuca sp. n., A. campususp sp. n., A. gephyrus sp. n., A. plicatus sp. n., and A. ultimus sp. n. are described and illustrated as males, and Parametriocnemus subnubilus Sinharay et Chaudhuri, 1979 from India is transferred to Antillocladius. Two new species of the closely related genus, Litocladius Mendes, Andersen et Sæther, 2004, L. floripa sp. n. and L. confusus sp. n., are described and figured as males. Two new Neotropical genera are described and figured. Gravatamberus gen. n. with five included species, G. apicalus sp. n., G. curtus sp. n., G. guatemaltecus sp. n., G. nidularium sp. n., and G. pilosus sp. n., is based on larva, pupa, and male. Lyrocladius gen. n. with the single included species L. radulatus sp. n. is based on the male only. Keys to orthoclad genera with scalpellate acrostichals and to the males of the included genera are provided. According to the cladogram presented, three monophyletic groups can be recognized; one composed of Antillocladius, Litocladius and Lyrocladius; one composed of Compterosmittia Sæther, Hanocladius Wang et Sæther, Limnophyes Eaton and Paralimnophyes Brundin; and a third, composed of Gravatamberus, Genus H sensu Epler, and Gynocladius Mendes, Sæther et Andrade-Morraye.References
Andersen, T. & Contreras-Ramos, A. (1999) First record of Antillocladius Sæther from Continental South America (Chironomidae, Orthocladiinae). Acta Zoologica Academiae Scientiarum Hungarica, 45, 149–154.
Andersen, T. & Mendes, H.F. (2007) Five enigmatic new orthoclad genera from Brazil (Diptera: Chironomidae, Orthocladiinae). In: Andersen, T. (Ed.), Contributions to the Systematics and Ecology of Aquatic Diptera - A tribute to Ole A. Sæther. The Caddis Press, Columbus, Ohio, pp. 17–52.
Andersen, T. & Sæther, O.A. (2005) Onconeura, a new Neotropical orthoclad genus (Chironomidae, Orthocladiinae). Zootaxa, 957, 1–16.
Banarescu, P. (1990) Zoogeography of Fresh Waters. 1. General Distribution and Dispersal of Freshwater Animals. AULA Verlag, Wiesbaden, pp. 1–511.
Banarescu, P. (1991) Zoogeography of Fresh Waters. 2. Distribution and Dispersal of Freshwater Animals in North America and Eurasia. AULA Verlag, Wiesbaden, pp. 512–1091.
Banarescu, P. (1995) Zoogeography of Fresh Waters. 3. Distribution and Dispersal of Freshwater Animals in Africa, Pacific Areas and South America. AULA Verlag, Wiesbaden, pp. 1092–1617.
Brooks, D. (1990) Parsimony analysis in historical biogeography and evolution: methodological and theoretical update. Systematic Biology, 41, 436–445.
Brooks, D., Dowling, A.P.G., van Veller, M.G.P. & Hoberg, E.P. (2004) Ending a decade of deception: a valiant failure, a not-so-valiant failure, and a success story. Cladistics, 20, 32–46.
Brooks, D., van Veller, M.G.P. & McLennan, D.A. (2001) How to do BPA, really. Journal of Biogeography, 28, 345–358.
Chaudhuri, P.K. & Guha, D.K. (1987) A conspectus of chironomid midges (Diptera: Chironomidae) of India and Bhutan. Entomologica scandinavica, Supplement, 29, 23–33.
Chaudhuri, P.K., Hazra, N. & Alfred, J.R.B. (2001) A checklist of chironomid midges (Diptera: Chironomidae) of the Indian subcontinent. Oriental Insects, 35, 335–372.
Coffman, W.P., Cranston, P.S., Oliver, D.R. & Sæther, O.A. (1986) The pupae of Orthocladiinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. In: Wiederholm, T. (Ed.), Chironomidae of the Holarctic region. Keys and diagnoses. Part 2. Pupae. Entomologica scandinavica, Supplement, 28, 147–298.
Cranston, P.S. (2000) Parapsectrocladius: a new genus of Orthocladiinae Chironomidae (Diptera) from Patagonia, the southern Andes. Insect Systematics and Evolution, 31, 103–120.
Cranston, P.S. (2007) The identity of Dactylocladius commensalis (Diptera: Chironomidae) revealed. Aquatic Insects, 29, 103–114.
Cranston, P.S. & Edward, D.D. (1999) Botryocladius gen.n.: a new transantarctic genus of orthocladiine midge (Diptera: Chironomidae). Systematic Entomology, 24, 305–333.
Cranston, P.S., Oliver, D.R. & Sæther, O.A. (1983) The larvae of Orthocladiinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. In: Wiederholm, T. (Ed.), Chironomidae of the Holarctic region. Keys and diagnoses. Part 1. Larvae. Entomologica scandinavica, Supplement, 19, 149–291.
Cranston, P.S., Oliver, D.R. & Sæther, O.A. (1989) The adult males of Orthocladiinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. In: Wiederholm, T. (Ed.), Chironomidae of the Holarctic region. Keys and diagnoses. Part 3. Adult males. Entomologica scandinavica, Supplement, 34, 165–352.
Ebach, M., Humphries, C.J. & Williams, D.M. (2003) Phylogenetic biogeography deconstructed. Journal of Biogeography, 30, 1285–1296.
Epler, J.H. (2001) Identification Manual for the larval Chironomidae (Diptera) of North and South Carolina. A Guide to the taxonomy of the Midges of the southeastern United States, including Florida. Special Publication SJ2001-SP13, North Carolina Department of Environment and Natural Resources, Raleigh, NC, and St. Johns River Water Management District, Palatka, FL, 526 pp.
Ferrington, L.C. Jr. & Sæther, O.A. (1995) Afrotropical species of Parakiefferiella Thienemann, with a review of species with palpal projections (Diptera: Chironomidae). In: Cranston, P.S. (Ed.), Chironomids: From Genes to Ecotypes. CSIRO, Melbourne, pp. 369–377.
Humphries, C.J. & Parenti, L.R. (1999) Cladistic Biogeography. Second edition: Interpreting patterns of plant and animal distributions. Oxford University Press, New York, 187 pp.
Marcondes, C.B. & Pinho, L.C (2005) First description of emergence traps for bromeliads and preliminary results of collections for southern Brazil (Insecta: Diptera and others). Studia Dipterologica, 12, 3–7.
Mendes, H.F. (2002) Rearing Tanypodinae, Telmatogetoninae and Orthocladiinae in Brazil, an empirical approach. Chironomus Newsletter, 15, 29–32.
Mendes, H.F., Andersen, T. & Sæther, O.A. (2004) A review of Antillocladius Sæther, 1981; Compterosmittia Sæther, 1981 and Litocladius new genus (Chironomidae, Orthocladiinae). Zootaxa, 594, 1–82.
Mendes, H.F., Sæther, O.A. & Andrade-Morraye, M. (2005) Gynocladius scalpellosus gen. n., sp. n. from Brazil (Diptera: Chironomidae: Orthocladiinae). Zootaxa, 979, 1–12.
Pinho, L.C., Mendes, H.F. & Marcondes, C.B. (2005) A new Brazilian species of Stenochironomus Kieffer mining decayed leaves in bromeliads (Diptera: Chironomidae). Zootaxa, 1046, 37–47.
Sasa, M. & Suzuki, H. (1995) The chironomid species collected on the Tokara Islands, Kagoshima (Diptera). Japanese Journal of Sanitary Zoology, 46, 255–288.
Sasa, M. & Suzuki, H. (2000) Studies on the chironomid midges collected on Yakushima Island, Southwestern Japan. Tropical Medicine, 42, 53–134.
Sæther, O.A. (1968) Chironomids of the Finse Area, Norway, with special reference to their distribution in a glacier brook. Archiv für Hydrobiologie, 64, 426–483.
Sæther, O.A. (1969) Some Nearctic Podonominae, Diamesinae and Orthocladiinae (Diptera: Chironomidae). Bulletin of the Fisheries Research Board of Canada, 107, 1–154.
Sæther, O.A. (1980) Glossary of Chironomid morphology terminology (Diptera: Chironomidae). Entomologica scandinavica, Supplement, 14, 1–51.
Sæther, O.A. (1981) Orthocladiinae (Chironomidae: Diptera) from the British West Indies with descriptions of Antillocladius gen. n., Lipurometriocmemus gen. n., Compterosmittia gen. n. and Diplosmittia gen. n. Entomologica scandinavica, Supplement, 16, 1–46.
Sæther, O.A. (1982) Orthocladiinae (Diptera: Chironomidae) from SE U.S.A., with descriptions of Plhudsonia, Unniella and Platysmittia n. genera and Atelopodella n. subgen. Entomologica scandinavica, 13, 465–510.
Sæther, O.A. (1984) The immatures of Antillocladius Sæther, 1981 (Diptera: Chironomidae). Aquatic Insects, 6, 1–6.
Sæther, O.A. (2000) Zoogeographical patterns in Chironomidae (Diptera). Verhandlungen der Internationalen Vereinigung für Theoretische und Angewandte Limnologie, 27, 290–302.
Sæther, O.A., Ashe, P. & Murray, D.A. (2000) A.6. Family Chironomidae. In: Papp, L. & Darvas, B. (Eds), Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Appendix. Science Herald, Budapest, pp. 113–334.
Sæther, O.A. & Ekrem, T. (2003) Biogeography of afrotropical Chironomidae (Diptera), with special reference to Gondwanaland. Cimbebasia, 19, 123–139.
Schlee, D. (1966) Präparation und Ermittlung von Meßwerten an Chironomidae (Diptera). Gewässer und Abwässer, 41/42, 169–193.
Sinharay, D.C. & Chaudhuri, P.K. (1979) A study of Orthocladiinae. Genus Parametriocnemus Goetghebuer from India (Diptera: Chironomidae). Entomologica scandinavica, Supplement, 10, 119–123.
Spies, M. & Reiss, F. (1996) Catalog and bibliography of Neotropical and Mexican Chironomidae (Insecta, Diptera). Spixiana, Supplement, 22, 61–119.
Swofford, D.L. (1998) PAUP*. Phylogenetic analysis using parsimony (*and other methods), version 4. Sinauer Associates, Sunderland, Massachusetts [computer software.]
Swofford, D.L. & Olsen, G.L. (1990) Phylogeny reconstruction in molecular systematics. In: Hillis, D.M. & Moritz, E.C. (Eds.), Molecular Systematics. Synauer Associates, Sunderland, Massachusetts, pp. 411–501.
Tiunova, T.M., Teslenko, V.A., Arefina, T.I., Makarchenko, M.A. & Zorina, O.V. (2003) Fauna of aquatic insects in Barabashevka River Basin (Southern Primorye). Vladimir Ya. Levanidov's Biennial Memorial Meetings, 2, 61–69. [in Russian.]
van Veller, M.G.P., Zandee, M. & Kornet, D.J. (1999) Two requirements for obtaining valid common patterns under different assumptions in vicariance biogeography. Cladistics, 15, 393–406.
van Veller, M.G.P., Kornet, D.J. & Zandee, M. (2001) Methods in vicariance biogeography: assessment of the implementations of assumptions 0, 1, and 2. Cladistics, 16, 319–345.
Wang, X. (2000) A revised checklist of chironomids from China (Diptera). In: Hoffrichter, O. (Ed.), Late 20th Century Research on Chironomidae: An Anthology from the 13th International Symposium on Chironomidae. Shaker Verlag, Aachen, pp. 629–652.
Wang, X. & Sæther, O.A. (1993) First Palaearctic and Oriental records of the orthoclad genus Antillocladius Sæther (Diptera: Chironomidae). Entomologica scandinavica, 24, 227–230.
Yamamoto, M. (2004) A catalog of Japanese Orthocladiinae (Diptera: Chironomidae). Makunagi / Acta Dipterologica, 21, 1–121.