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Type: Monograph
Published: 2024-11-29
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THE SCALE INSECTS OF IRAN (HEMIPTERA: COCCOMORPHA) PART 3 THE SOFT SCALES (COCCIDAE) AND OTHER FAMILIES

Department of Insect Taxonomy Research; Iranian Research Institute of Plant Protection; Agricultural Research Education and Extension Organisation; Tehran; Iran
Science: Research; Natural History Museum; Cromwell Road; London SW7 5BD; U.K.
Hemiptera Coccomorpha Coccidae Iran

Abstract

The present work is the third and final part in a series of monographs on the Iranian scale insect fauna (Hemiptera: Coccomorpha) by the same authors: Part 1, the armoured scales (Diaspididae), was published in 2021, and Part 2, the mealybugs (Pseudococcidae and Rhizoecidae) and Putoidae, was published in 2022. Here we cover ten families: Asterolecaniidae, Cerococcidae, Coccidae, Cryptococcidae, Eriococcidae, Kermesidae, Margarodidae, Monophlebidae, Ortheziidae and Phoenicococcidae, bringing the total number of scale insect families recorded in Iran to 14. A key is provided to separate all 14 families. For each of the ten families in the present work, keys to identify the genera and species found in Iran are provided. These are accompanied by detailed descriptions, illustrations, distribution data, information on natural enemies, economic importance and the host-plants recorded in Iran. The main purpose of this work is to provide agricultural staff concerned with pest control and quarantine inspection in Iran (and elsewhere) with a reliable means of species identification. Among the recorded species in the present work, Ceroplastes elytropappi (Brain, 1920) and Mesolecanium nigrofasciatum (Pergande, 1898) (Coccidae) are considered to be doubtful records for Iran. Pulvinaria aligarhensis Avasthi & Shafee, 1985 and Pulvinaria urbicola Cockerell, 1893 are both recorded from Iran for the first time.

 

References

  1. Abolmasoumi, N., Talebi, A.A., Rakhsahni, E. & Lotfalizadeh, H. (2009) Study on faunistic, biodiversity and host rates of (Hym., Aphelinidae) in some parts of Markazai and Lorestan provinces, Iran. Journal of Entomological Research, 2 (1), 1‒12.
  2. Ahmad, R. (1975) A note on Saissetia oleae and its natural enemies in Iran. Entomophaga, 20, 221‒223. https://doi.org/10.1007/BF02371663
  3. Afchar, J. (1937) Les Insectes Nuisibles aux Arbres Fruitiers en Iran. Ministry of Agriculture, Jihad, 112 pp.
  4. Archangelskaya, A.D. (1923) Contribution to the fauna of hard or soft scales (Coccidae) of Turkestan. Transactions of the Scientific Society of Turkestan, 1, 259‒266.
  5. Archangelskaya, A.D. (1930) [New species of scale insects, Coccidae, from Central Asia]. Zashchita Rastenii, Leningrad, 7, 69‒85. [in Russian]
  6. Archangelskaya, A.D. (1931) [New species of scale insects, Coccidae, from Central Asia]. Zashchita Rastenii, Leningrad, 7, 69‒85. [in Russian]
  7. Archangelskaya, A.D. (1935) [The carmine-producing coccids (Margarodes) of Middle Asia and species of allied genus Neomargarodes]. Tashkent Publishing Committee of Sciences in Uzbek Tashkent, 38 pp. [in Russian]
  8. Archangelskaya, A.D. (1937) [The Coccidae of Middle Asia]. Izdatelstvo Komiteta Nauk UZ, Tashkent, 158 pp. [in Russian]
  9. Ashmead, W.H. (1891) A generic synopsis of the Coccidae. Family X. - Coccidae. Transactions of the Entomological Society of America, 18, 92‒102.
  10. Avasthi, R.K. & Shafee, S.A. (1985) Two new species of Pulvinaria Targ.-Tozz. (Homoptera: Coccidae) from India. Current Science, 54, 1289‒1291.
  11. Balachowsky, A.S. (1929) Contribution à l’étude des coccides de l’Afrique Mineure (6e note). Faune du Hoggar. Annales de la Société entomologique de France, 98, 301‒322. https://doi.org/10.1080/21686351.1929.12280111
  12. Balachowsky, A.S. (1953) Sur les Kermes Boitard (Hom. Coccoidea) des chênes du bassin oriental de la Méditerranée. Revue de Pathologie végétale et d’Entomologie agricole de France, 32, 181−189.
  13. Balachowsky, A.S. (1954) Sur une nouvelle espèce d’Eriococcini de la forêt de Fontainebleau avec création d’un genre nouveau. [Hom. Coccoidea]. Bulletin de la Société entomologique de France, 59, 61–64. https://doi.org/10.3406/bsef.1954.18703
  14. Bärensprung, F.V. (1849) [Observations of some native species from the family of Coccidae]. Zeitung für Zoologie, Zootomie und Palaeozoologie, 1, 165–170 + 173–176. [in German]
  15. Bazarov, B.B. (1968) [New species of Coccidae from Pamir]. Doklady Akademii Nauk Tadzhikskoi, Dushanbe, 11, 73–74. [in Russian]
  16. Ben-Dov, Y. (1969) A generic diagnosis of Bodenheimera Bodenheimer (Homoptera: Coccidae) with redescription of B. rachelae (Bodenheimer). Proceedings of the Royal Entomological Society of London (B), 38, 70‒74. https://doi.org/10.1111/j.1365-3113.1969.tb00232.x
  17. Ben-Dov, Y. (1989) Saissetia Déplanche, 1859 (Insecta, Homoptera): proposed designation of Lecanium coffeae Walker, 1852 as the type species. Bulletin of Zoological Nomenclature 46 (92), 114‒118. https://doi.org/10.5962/bhl.part.506
  18. Ben-Dov, Y. (1993) A Systematic Catalogue of the Soft Scale Insects of the World (Homoptera: Coccoidea: Coccidae). Sandhill Crane Press, Gainesville, Florida, 536 pp.
  19. Bodenheimer, F.S. (1924) The Coccidae of Palestine. First report on this family. Zionist Organization Institute of Agriculture and Natural History, Agricultural Experiment Station Bulletin, 1, 1‒100.
  20. Bodenheimer, F.S. (1926) Second note on the Coccidae of Palestine. Bulletin of Entomological Research, 17, 189‒192. https://doi.org/10.1017/S0007485300019222
  21. Bodenheimer, F.S. (1935) Studies on the zoogeography and ecology of palearctic Coccidae I-III. EOS, 10, 237‒271.
  22. Bodenheimer, F.S. (1941) Seven new species of Coccidae from Anatolia. Revue de la Faculté des Sciences de l’Université d’Istanbul, Series B, 6, 65–84.
  23. Bodenheimer, F.S. (1943) A first survey of the Coccoidea of Iraq. Government of Iraq, Ministry of Economics, Directorate General of Agriculture, Bulletin, 28, 1‒33.
  24. Bodenheimer, F.S. (1944a) Additions to the Coccoidea of Iraq, with descriptions of two new species. Bulletin de la Société Fouad 1er d’Entomologie, 28, 81‒84.
  25. Bodenheimer, F.S. (1944b) Note on the Coccoidea of Iran, with description of new species. Bulletin de la Société Fouad 1er d’Entomologie, 28, 85‒100.
  26. Bodenheimer, F.S. (1951) Description of some new genera of Coccoidea. Entomologische Berichten. Amsterdam, 13, 328‒331.
  27. Boisduval, J.B.A. (1869) Note sur deux espèces nouvelles de coccides vivant sur les bambous cultivés au jardin du Hamma. L’Insectologie Agricole, 3, 260‒262.
  28. Boitard, P. (1828) Les gallinsectes. Manuel d’entomologie ou Histoire naturelle des Insectes. Roret, Paris, 435 pp. https://doi.org/10.5962/bhl.title.122860
  29. Borchsenius, N.S. (1948) [On the revision of the genus Eriococcus Sign. (Insecta, Homoptera, Coccoidea)]. Doklady Akademii Nauk S, Moscow, New Series, 60, 501‒503. [in Russian]
  30. Borchsenius, N.S. (1949a) [A new genus and new species of hard and soft scales (Homoptera, Coccoidea) of USSR fauna]. Entomologicheskoe Obozrenye, 30, 334‒353. [in Russian]
  31. Borchsenius, N.S. (1949b) [Identification of the soft and armored scales of Armenia]. Doklady Akademii Nauk Armyanoskoi S, 1949, 1‒271. [in Russian]
  32. Borchsenius, N.S. (1950) [Mealybugs and scale insects of USSR (Coccoidea)]. Akademii Nauk S, Zoological Institute Moscow, 32, 1–250. [in Russian]
  33. Borchsenius, N.S. (1952a) [A new genus and new species of hard scales from Iran (Homoptera, Coccoidea)]. Entomologicheskoe Obozrenye, 32, 261‒263. [in Russian]
  34. Borchsenius, N.S. (1952b) [New genera and species of soft scales of the family Coccidae (=Lecaniidae) of the USSR fauna and adjacent countries (Insecta, Homoptera, Coccoidea)]. Trudy Zoologicheskogo Instituta Akademii Nauk S, 12, 269‒316. [in Russian]
  35. Borchsenius, N.S. (1953) [New genera and species of scale insects of the family Coccidae (Homoptera, Coccoidea)]. Entomologicheskoe Obozrenye, 33, 281‒290. [in Russian]
  36. Borchsenius, N.S. (1955) [New species of false-hard scales (Homoptera, Coccoidea, Coccidae) of the fauna of USSR and adjacent countries]. Trudy Zoologicheskogo Instituta Akademii Nauk S, 18, 288‒303. [in Russian]
  37. Borchsenius, N.S. (1957) [Subtribe mealybugs and scales (Coccoidea). Soft scale insects Coccidae. Vol. IX]. Fauna S. Zoologicheskii Institut Akademii Nauk S, New Series, 66, 1‒493. [in Russian]
  38. Borchsenius, N.S. (1960) [Fauna of USSR Homoptera, Kermococcidae, Asterolecaniidae, Lecanidodiaspididae, Aclerdidae]. Akademiia Nauk S, Zoologicheskii Institut (Series), Leningrad, 282 pp. [in Russian]
  39. Bosc d’Antic, L.A.G. (1784) Description de l’Orthezia characias. Journal de Physique, 1, 173.
  40. Bouché, P.F. (1844) [Contributions to the natural history of the “Scharlachläuse” (Coccina)]. Entomologische Zeitung, Stettin, 5, 293‒302. [in German]
  41. Bouché, P.F. (1851) [New species of scale insects]. Entomologische Zeitung, Stettin, 12, 110‒112. [in German]
  42. Boyer de Fonscolombe, E.L.J.H. (1834) Description des Kermes qu’on trouve aux environs d’Aix. Annales de la Société entomologique de France, 3, 201‒218.
  43. Bozorg-Amirkalaee, M., Fathi, S.A.A., Golizadeh, A. & Mahdavian, S.E. (2015) Performance of Cryptolaemus montrouzieri feeding on Pulvinaria aurantii ovisacs on citrus plants. Biocontrol Science and Technology, 25 (2), 207‒222. https://doi.org/10.1080/09583157.2014.968521
  44. Brain, C.K. (1920) The Coccidae of South Africa - V. Bulletin of Entomological Research, 11, 1‒41. https://doi.org/10.1017/S000748530004428X
  45. Brandt, J.F. & Ratzeburg, J.T.C. (1833) [Medical zoology or faithful representation and description of animals, ... published in a systematic sequence. Vol. 2]. Gedruckt bei Trowitzsch und Sohn, Berlin, IV + 364 pp. [in German]
  46. Brown, S.W. & McKenzie, H.L. (1962) Evolutionary patterns in the armored scale insects and their allies (Homoptera: Coccoidea: Diaspididae, Phoenicococcidae, and Asterolecaniidae). Hilgardia, 33, 141‒170. https://doi.org/10.3733/hilg.v33n04p141
  47. Burmeister, H. (1835) [Scarlet lice. Scale insects. Coccina. (Gallinsecta 1)]. Handbuch der Entomologie. T.F. Inslin, Berlin. [unknown pagination, in German]
  48. Cardon, D. (2007) Natural Dyes - Sources, Tradition, Technology & Science. Archetype Publications, London, 800 pp.
  49. Clausen, C.P. (1923) The citricola scale in Japan, and its synonymy. Journal of Economic Entomology, 16, 225‒226. https://doi.org/10.1093/jee/16.2.225
  50. Cockerell, T.D.A. (1892) Coccidae of Jamaica. Journal of the Institute of Jamaica, 1, 142‒143.
  51. Cockerell, T.D.A. (1893a) Coccidae, or scale insects. - II. Bulletin of the Botanical Department, Jamaica, 40, 7‒9.
  52. Cockerell, T.D.A. (1893b) Two new species of Pulvinaria from Jamaica. Transactions of the Entomological Society of London, 1893, 159‒163. https://doi.org/10.1111/j.1365-2311.1893.tb02062.x
  53. Cockerell, T.D.A. (1894) A check-list of African Coccidae. Psyche, 7, 178. https://doi.org/10.1155/1894/52465
  54. Cockerell, T.D.A. (1896) Preliminary diagnoses of new Coccidae. Psyche, Supplement 7, 18‒21. https://doi.org/10.1155/1896/92937
  55. Cockerell, T.D.A. (1897) Two forms of fluted scale. Psyche, 8, 94. https://doi.org/10.1155/1897/46231
  56. Cockerell, T.D.A. (1899a) Rhynchota, Hemiptera - Homoptera. Biologia Centrali-Americana, 2 (2), 1‒37. Available from: https://www.biodiversitylibrary.org/item/14634#page/2/mode/1up (accessed 31 January 2024).
  57. Cockerell, T.D.A. (1899b) Some notes on Coccidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 1899, 259‒275.
  58. Cockerell, T.D.A. (1901) The coccid genus Saissetia. The Entomological Student, 2, 31‒33.
  59. Cockerell, T.D.A. (1902) South African Coccidae. - II. The Entomologist, 35, 111‒114. https://doi.org/10.5962/bhl.part.19973
  60. Cockerell, T.D.A. (1910) A new wax scale from the Argentine. Canadian Entomologist, 42, 74‒76. https://doi.org/10.4039/Ent4274-3
  61. Comstock, J.H. (1881) Report of the Entomologist. Report of the Commissioner of Agriculture, United States Department of Agriculture, 1880/1881, 276‒349.
  62. Cook, L.G. & Gullan, P.J. (2004) The gall-inducing habit has evolved multiple times among the eriococcid scale insects (Sternorrhyncha: Coccoidea: Eriococcidae). Biological Journal of the Linnean Society, 83 (4), 441‒452. https://doi.org/10.1111/j.1095-8312.2004.00396.x
  63. Cook, L.G., Gullan, P.J. & Trueman, H.E. (2002) A preliminary phylogeny of the scale insects (Hemiptera: Sternorrhyncha: Coccoidea) based on nuclear small-subunit ribosomal DNA. Molecular Phylogenetics and Evolution, 25 (1), 43‒52. https://doi.org/10.1016/S1055-7903(02)00248-8
  64. Cooke, M. (1881) A Treatise, Insects Injurious to Fruit and Fruit Trees of the State of California and Remedies Recommended for Their Extermination. State Office, Sacramento, California, 71 pp. https://doi.org/10.5962/bhl.title.37277
  65. Costa, O.G. (1828) Prospetto di una nuova divisione metodica del genere Coccus, Lin. Lam. Ec. Dalla Tipografia Trani., Napoli, 8 pp.
  66. Costa, O.G. (1829) Fauna del Regno di Napoli, famiglia de coccinigliferi, o de’ gallinsetti. Emitteri, Napoli, 23 pp.
  67. Cox, J.M. & Williams, D.J. (1987) Do the Eriococcidae form a monophyletic group? Bollettino del Laboratorio di Entomologia Agraria ‘Filippo Silvestri,’ 43 (1986 Supplement), 13‒17.
  68. Danzig, E.M. (1972) [Contribution to the fauna of the white flies and scale insects (Homoptera: Aleyrodoidea, Coccoidea) of Mongolia]. Insects of Mongolia, 1, 325‒348. [in Russian]
  69. Danzig, E.M. (1975) [New species of the genus Acanthococcus Sign. (Homoptera, Coccoidea, Eriococcidae) from the Far East of USSR]. Entomologicheskoe Obozrenye, 54, 62‒81. [in Russian]
  70. Danzig, E.M. (1980) Coccoids of the Far East USSR (Homoptera, Coccinea) with phylogenetic analysis of scale insects fauna of the world. Nauka, Leningrad, 367 pp.
  71. Danzig, E.M. (1990) [New species of coccids (Homoptera, Coccinea) from Iran, Mongolia and Vietnam]. Entomologicheskoe Obozrenye, 69, 373‒376. [in Russian]
  72. Davatchi, A.G. (1946) Saissetia oleae Bernard (Homoptera--Coccidae). Ministère de l’Agriculture, Entomologie et Phytopathologie Appliquées, Tehran, 1, 1‒7 [in Persian]; 1‒2. [in French]
  73. Davoodi, A., Talebi, A.A., Rajabi, Gh.R. & Fathipour, Y. (2002) The first report of Microterys hortulanus (Hym.: Encyrtidae) from Iran. Journal of Entomological Society of Iran, 22 (1), 85‒86.
  74. Davoodi, A., Talebi, A.A., Rajabi, G.R., Fathipour, Y. & Moharramipour, S. (2004) [Report of Metaphycus claviger (Hym.: Encyrtidae) from Iran]. Journal of Entomological Society of Iran, 23 (2), Pe129‒Pe130. [in Persian]
  75. Davoodi, A., Talebi, A.A., Radjabi, G.R., Fathipour, Y. & Xu, Z. (2003) Report of Metaphycus angustifrons Compère (Hym.: Encyrtidae), parasitoid of Coccus hesperidum (L.) (Hom.: Coccidae) from Iran. Journal of Entomological Society of Iran, 22 (2), 80.
  76. De Lotto, G. (1969a) On a few old and new soft scales and mealybugs (Homoptera: Coccoidea). Journal of the Entomological Society of southern Africa, 32, 413‒422.
  77. De Lotto, G. (1969b) A new genus of wax scales (Homoptera: Coccidae). Bollettino del Laboratorio di Entomologia Agraria ‘Filippo Silvestri’, Portici, 27, 210‒218.
  78. De Lotto, G. (1971) The authorship of the Mediterranean black scale (Homoptera: Coccidae). Journal of Entomology (B), 40, 149‒150. https://doi.org/10.1111/j.1365-3113.1971.tb00117.x
  79. Del Guercio, G. (1900) [Observations on a new scale insect harmful to citrus fruits in Italy and on the way to immunize the woody part of the plants against the bite of scale insects in general and to destroy them]. Nuove Relazione della R. Stazione di Entomologia Agraria Firenze, 3, 3‒26. [in Italian]
  80. Déplanche, E. (1859) [Analysis of a memoir on a disease of coffee by M.E. Déplanche, presented by M. Études-Deslongchamps]. Bulletin de la Société Linnéenne de Normandie, 4, 203‒207. [in French]
  81. Douglas, J.W. (1890a) Notes on some British and exotic Coccidae. No. 15. Entomologist’s Monthly Magazine, 26, 79–81.
  82. Douglas, J.W. (1890b) Notes on some British and exotic Coccidae. No. 16. Entomologist’s Monthly Magazine, 26, 153–155.
  83. Ebeling, W. (1951) Citrus pests in California. California Citrograph, 37 (2), 50, 81.
  84. Ebrahimi, E. (2014) Parasitoid and hyperparasitoid wasps of scale insects in Hayk Mirzayans Insect Museum, Iran. Journal of Entomological Society of Iran, 34 (1), 73−83.
  85. Emeljanov, A.F. (1974) [Proposals on the classification and nomenclature of areals]. Entomologicheskoe Obozrenie, 53 (3), 11−26. [in Russian]
  86. Ezzat, Y.M. & Hussein, N.A. (1969) Redescription and classification of the family Coccidae in U.A.R. (Homoptera: Coccoidea). Bulletin de la Société entomologique d’Egypte, 51, 359‒426. [1967]
  87. Fabricius, J.C. (1775) Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species Adiectis Synonymis, Locis, Descriptionibus, Observationibus. VII. Ryngota. Flensburgi et Lipsiae, Korte, 832 pp. https://doi.org/10.5962/bhl.title.36510
  88. Fabricius, J.C. (1776) Genera Insectorum. Chilonii, Bartschii, 310 pp.
  89. Farahbakhsh, Gh. (1961) A Checklist of Economically Important Insects and other Enemies of Plants and Agricultural Products in Iran. Department of Plant Protection, Ministry of Agriculture, Tehran, 151 pp.
  90. Feli Kohikheili, Z. & Damavandian, M.R. (2015) First record of two parasitoid wasps (Hym.: Mymaridae) from Iran. Applied Entomology and Phytopathology, 83 (1), 73‒76.
  91. Fernald, M.E. (1903) A catalogue of the Coccidae of the world. Bulletin of the Hatch Experiment Station of the Massachusetts Agricultural College, 88, 1‒360. https://doi.org/10.5962/bhl.title.8533
  92. Ferris, G.F. (1955) Atlas of the Scale Insects of North America. Vol. 7. The Families Aclerdidae, Asterolecaniidae, Conchaspididae Dactylopiidae and Lacciferidae. iii. Stanford University Press, Palo Alto, 233 pp.
  93. Ferris, G.F. (1957) A review of the family Eriococcidae (Insecta: Coccoidea). Microentomology, 22, 81‒89.
  94. Foldi, I. (2005) The Matsucoccidae in the Mediterranean basin with a world list of species (Hemiptera: Sternorrhyncha: Coccoidea). Annales de la Société entomologique de France, 40 (2), 145–168. [2004] https://doi.org/10.1080/00379271.2004.10697412
  95. Fourcroy, A.F. (1785) Entomologia Parisiensis. s.n., Paris, 544 pp.
  96. García Morales M., Denno B.D., Miller D.R., Miller G.L., Ben-Dov Y. & Hardy N.B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http://scalenet.info (accessed 31 January 2024) https://doi.org/10.1093/database/bav118
  97. Gascard, A. (1893) Contribution À L’Étude Des Gommes Laques Des Indes & De Madagascar. Société d’Éditions Scientifiques, Paris, 87 pp. [in French]. Available from: https://archive.org/details/contributionltu00tozegoog/page/n91/mode/2up (accessed 22 November 2023)
  98. Gavrilov-Zimin, I.A. (2018) Ontogenesis, morphology and higher classification of archaeococcids (Homoptera: Coccinea: Orthezoiidea). Zoosystematica Rossica, 2018 (2), 1–264. https://doi.org/10.31610/zsr/2018.supl.2.1
  99. Ghahari, H., Abd-Rabou, Sh., Sakenin, H., Hedqvist, K.J. & Ostovan, H. (2010) A contribution to some Chalcidoidea wasps (Hymenoptera) from Iran. Journal of Biological Control, 24 (1), 17–21.
  100. Gill, R.J. (1988) The Scale Insects of California: Part 1. The Soft Scales (Homoptera: Coccoidea: Coccidae). California Department of Food & Agriculture, Sacramento, 132 pp.
  101. Gill, R.J. (1993) The Scale Insects of California: Part 2. The Minor Families (Homoptera: Coccoidea). California Department of Food & Agriculture, Sacramento, California, 241 pp.
  102. Golpayegani, Sh., Talebi, A.A., Lotfalizadeh, H. & Rakhshani, E. (2010) Faunistic study and host range of encyrtid parasitoids (Hym., Encyrtidae) collected in some parts of Lorestan and Markazi provinces of Iran. Journal of Entomological Research, 1 (4), 319‒329.
  103. Goux, L. (1989) Contribution à l’étude la morphologie des Eriococcus avec description d’une espèce nouvelle (Homoptera, Coccoidea, Eriococcidae). Bulletin de la Société Linnéenne de Provence, 40, 19–23.
  104. Goux, L. (1993) Description de trois espèces nouvelles d’Eriococcus appartenant à la faune Française (Homoptera, Coccoidea, Eriococcidae). Bulletin de la Société Linnéenne de Provence, 44, 65–69.
  105. Gray, E.J. (1828) Spicilegia Zoologica; or original figures and short systematic descriptions of new and unfigured animals. Part 1. Treuttel, Wurtz & Co., London, 12 pp.
  106. Green, E.E. (1897) Notes on Coccidae from the Royal Gardens, Kew [with additions by R. Newstead]. Entomologist’s Monthly Magazine, 33, 68‒77.
  107. Green, E.E. (1901) On some new species of Coccidae from Australia, collected by W.W. Froggatt, F.L.S. Proceedings of the Linnean Society of New South Wales, 25 (1900), 559–562. https://doi.org/10.5962/bhl.part.12174
  108. Guérin-Méneville, G.E. (1858) Communication sur un insecte, qui, en Chine, produit de la cire. Bulletin de la Société entomologique de France, 16, lxvii.
  109. Guérin-Méneville, G.E. (1867) M. ed. Morin adresse une lettre sur les ravages produits a l’Île de la Réunion par des insectes qui attaquent la canne à sucre. Revue et Magasin de Zoologie, 19, 450‒451.
  110. Gullan, P.J. & Cook, L.G. (2007) Phylogeny and higher classification of the scale insects (Hemiptera: Sternorrhyncha: Coccoidea). Zootaxa, 1668 (1), 413−425. https://doi.org/10.11646/zootaxa.1668.1.22
  111. Gwiazdowski, R.A., van Driesche R.G., Desnoyers, A., Lyon, S., Wu, S.A., Kamata, N. & Normark, B.B. (2006) Possible geographic origin of beech scale, Cryptococcus fagisuga (Hemiptera: Eriococcidae), an invasive pest in North America. Biological Control: Theory and Applications in Pest Management, 39 (1), 9–18. https://doi.org/10.1016/j.biocontrol.2006.04.009
  112. Hadzibejli, Z.K. (1960) [New species of coccids (Homoptera, Coccoidea) from Georgia]. Trudy Akademii Nauk Gruzinskoy, Instituta Zashchitii Rastenii, 13, 299‒321. [in Russian]
  113. Hadzibejli, Z.K. (1967) [Two new coccid species (Insecta, Homoptera, Coccoidea) of the fauna of Eastern Transcaucasia]. Soobshcheniya Akademii Nauk Gruzinskoy, Tbilisi, 46 (3), 715‒720. [in Russian]
  114. Hall, W.J. (1923) Further observations on the Coccidae of Egypt. Bulletin, Ministry of Agriculture, Egypt, Technical and Scientific Service, 36, 1‒61. https://doi.org/10.5962/bhl.title.19393
  115. Hall, W.J. (1925) Notes on Egyptian Coccidae with descriptions of new species. Bulletin, Ministry of Agriculture, Egypt, Technical and Scientific Service, 64, 1‒31.
  116. Hall, W.J. (1926) Contribution to the knowledge of the Coccidae of Egypt. Bulletin, Ministry of Agriculture, Egypt, Technical and Scientific Service, 72, 1‒41.
  117. Hamon, A.B. & Williams, M.L. (1984) The soft scale insects of Florida (Homoptera: Coccoidea: Coccidae). Arthropods of Florida and Neighboring Land Areas. Florida Department of Agricultural & Consumer Services, Division of Plant Industry, Gainesville, Florida, 194 pp.
  118. Hempel, A. (1920) Descripçoes de coccidas novas e pouco conhecidas. [Descriptions of new and little-known coccids]. Revista do Museu Paulista. São Paulo, 12, 329‒377. [in Portuguese]
  119. Heydari, S., Toorani, A.H., Doostdar Kalkenari, L. & Abbasipour, H. (2016) Comparison effect of botanical pesticides on first instar nymphs of the Australian mealybug, Icerya purchasi Maskell and its ladybird predator, Novius cardinalis Mulsant. 22nd Iranian Plant Protection Congress, Tehran, Iran, 27–30 August 2016. [unknown pagination]
  120. Hodgson, C.J. (1994) The Scale Insect Family Coccidae: an Identification Manual to Genera. CAB International, Wallingford, 639 pp.
  121. Hodgson, C.J. (2002) Preliminary phylogeny of some non-margarodid Coccoidea (Hemiptera) based on adult male characters. Bollettino di Zoologia Agraria e di Bachicoltura (Milano), 33 (2001‒3), 129‒137.
  122. Hodgson, C.J. (2020) A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males. Zootaxa, 4765 (1), 1–264. https://doi.org/10.11646/zootaxa.4765.1.1
  123. Hodgson, C.J. & Foldi, I. (2006) A review of the Margarodidae sensu Morrison (Hemiptera: Coccoidea) and some related taxa based on the morphology of adult males. Zootaxa, 1263 (1), 1‒250. https://doi.org/10.11646/zootaxa.1263.1.1
  124. Hodgson, C.J. & Hardy, N.B. (2013) The phylogeny of the superfamily Coccoidea (Hemiptera: Sternorrhyncha) based on the morphology of extant and extinct macropterous males. Systematic Entomology, 38, 794–804. https://doi.org/10.1111/syen.12030
  125. Hodgson, C.J. & Peronti, A.L.B.G. (2012) A revision of the wax scale insects (Hemiptera: Sternorrhyncha: Coccoidea: Ceroplastinae) of the Afrotropical Region. Zootaxa, 3372 (1), 1–265. https://doi.org/10.11646/zootaxa.3372.1.1
  126. Hodgson, C.J. & Williams, D.J. (2016) A revision of the family Cerococcidae Balachowsky (Hemiptera: Sternorrhyncha, Coccomorpha) with particular reference to species from the Afrotropical, western Palaearctic and western Oriental Regions, with the revival of Antecerococcus Green and description of a new genus and fifteen new species, and with ten new synonomies. Zootaxa, 4091 (1), 1‒175. https://doi.org/10.11646/zootaxa.4091.1.1
  127. Hoy, J.M. (1963) A catalogue of the Eriococcidae (Homoptera: Coccoidea) of the world. New Zealand Department of Scientific and Industrial Research Bulletin, 150, 1–260.
  128. ICZN (1999) International Code of Zoological Nomenclature. 4th Edition. International Commission on Zoological Nomenclature. Available from: https://code.iczn.org/?frame=1 (accessed 19 May 2024).
  129. International Commission on Zoological Nomenclature (1985) Opinion 1303. Bulletin of Zoological Nomenclature, 42, 139–141. Available from: https://www.biodiversitylibrary.org/partpdf/881 (accessed 31 January 2024)
  130. Jakubski, A.W. (1965) A critical revision of the families Margarodidae and Termitococcidae (Hemiptera, Coccoidea). Trustees of the British Museum (Natural History), London, 187 pp.
  131. Jalilvand, K., Shirazi, M., Vahedi, H.V., Fallahzadeh, M., Naghadeh & Samih, M.A. (2013) A preliminary study on natural enemies of Coccoidea (Hemiptera, Sternorrhyncha) in Kermanshah Province, Western Iran. Acta Phytopathologica et Entomologica Hungarica, 48 (2), 299‒308. https://doi.org/10.1556/APhyt.48.2013.2.11
  132. Japoshvili, G., Jalilvand, K., Vahedi, H.A. & Fallahzadeh, M. (2015) Metaphycus parthenolecanii sp. n. (Hymenoptera: Encyrtidae), a parasitoid of Parthenolecanium corni (Hemiptera: Coccidae) in Iran. Zoology in the Middle East, 2015, 1‒5. https://doi.org/10.1080/09397140.2015.1020610
  133. Jashenko, R.V. (1994) [New species of the genus Porphyrophora Brandt (Coccinea, Margarodidae) from Kazakhstan, Turkmenistan and Mongolia]. Selevinia, 2 (3), 22–38. [in Russian]
  134. Jashenko, R.V. & Danzig, E.M. (1992) [A new genus and three new species of the subfamily Monophlebinae Maskell (Homoptera, Margarodidae) from Middle Asia, Afghanistan and Vietnam]. Entomologicheskoe Obozrenie, 71 (1), 84‒90. [in Russian]
  135. Kaltenbach, J.H. (1860) [The German phytophages from the class of insects]. Verhandlungen des Naturhistorischen Vereins des Preussisches Rheinlandes, 17, 203–260. [in German]
  136. Karsch, F.A.F. (1877) [Revision of gall midges]. Müster in Westfalen, Brunn, 57 pp. [in German]
  137. Kaussari, M. (1946) Contribution à l’étude des Coccidae de l’Iran. Ministère de l’Agriculture, Département Général de la Protection Plantes, Tehran, 48 pp.
  138. Kaussari, M. (1955) La première liste des cochenilles de l’Iran. Ministère de l’Agriculture, Entomologie et Phytopathologie Appliquées, Publication du Département Général de la Protection Plantes. Tehran, 15, 14–20.
  139. Kaussari, M. (1957) La deuxième liste des cochenilles de l’Iran. Ministère de l’Agriculture, Entomologie et Phytopathologie Appliquées, Publication du Département Général de la Protection Plantes. Tehran, 16‒17, 1‒3.
  140. Kaydan, M.B. & Kozár, F. (2010) A review of the genus Neoacanthococcus Borchsenius (Hemiptera: Coccoidea: Eriococcidae) with a description of Neoacanthococcus atlihani sp. nov. in Turkey. Turkiye Entomoloji Dergisi, 34 (2), 165–177.
  141. Kaydan, M.B., Benedicty, Z.K. & Szita, É. (2015) The genus Orthezia Bosc (Hemiptera, Ortheziidae) in Turkey, with 2 new records. Turkish Journal of Zoology, 39, 160−167. https://doi.org/10.3906/zoo-1403-9
  142. Kaydan, M.B., Konczné Benedicty, Z. & Szita, É. (2014) New species of the genus Ortheziola Šulc (Hemiptera, Coccoidea, Ortheziidae), ZooKeys, 406, 65–80. https://doi.org/10.3897/zookeys.406.7596
  143. Kiritchenko, A.N. (1940) Third report on the coccid fauna of USSR. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 1940, 115–137.
  144. Kiriukhin, G. (1946) Les insectes nuisibles au pistacier en Iran. Entomologie et Phytopathologie Appliquées, 1, 1–17 (in Persian) + 2‒4 (in French).
  145. Kirkaldy, G.W. (1906) On the nomenclature of the genera of the Hemiptera. The Entomologist, 39, 253–257. https://doi.org/10.5962/bhl.part.1563
  146. Kondo, T. & Watson, G.W. (2022) 5. Collection, preservation, slide-mounting, labelling and vouchering of scale insects. In: Kondo, T. & Watson, G.W. (Eds.), Encyclopedia of Scale Insect Pests. CABI Publishing, Wallingford, pp. 548‒558. ISBN/EAN 1800620640/ 9781800620643. Available from: https://www.cabidigitallibrary.org/doi/10.1079/9781800620643.0005 (accessed 24 November 2023) https://doi.org/10.1079/9781800620643.0005
  147. Kosztarab, M.P. & Kozár, F. (1988) Scale Insects of Central Europe. Akademiai Kiado, Budapest, 456 pp. https://doi.org/10.1007/978-94-009-4045-1
  148. Koteja, J. (1974) On the phylogeny and classification of the scale insects (Homoptera, Coccinea) (discussion based on the morphology of the mouthparts). Acta Zoologica Cracoviensia, 19, 267–325.
  149. Koteja, J. (1978) An introduction to the revision of the Eriopeltini Šulc (Homoptera, Coccidae), with establishment of two new genera. Polskie Pismo Entomologiczne, 48, 311‒327.
  150. Koteja, J. (1980) Campaniform, basiconic, coeloconic, and intersegmental sensilla on the antennae in the Coccinea (Homoptera). Acta Biologica Cracoviensia, 22, 73‒88.
  151. Koteja, J. (1996) Cataloguing fossil scale insects (Homoptera, Coccinea). Inclusion Wrostek, 22, 2.
  152. Kozár, F. (1974) Mass infestation and damage of the oak scale Kermes quercus L. (Homoptera, Coccoidea). Növényvédelem, 10, 534–537.
  153. Kozár, F. (2009) Zoogeographical analysis and status of knowledge of the Eriococcidae (Hemiptera), with a world list of species. Bollettino di Zoologia Agraria e di Bachicoltura, Milano, Series II, 41 (2), 87‒121.
  154. Kozár, F., Fowjhan, M.A. & Zarrabi, M. (1996) Check-list of Coccoidea and Aleyrodoidea (Homoptera) of Afghanistan and Iran, with additional data to the scale insects of fruit trees in Iran. Acta Phytopathologica et Entomologica Hungarica, 31, 61‒74.
  155. Kozár, F., Kaydan, M.B., Konczné Benedicty, Z. & Szita, É. (2013) Acanthococcidae and Related Families of the Palaearctic Region. Hungarian Academy of Sciences, Budapest, 680 pp.
  156. Kuwana, S.I. (1914) Coccidae of Japan, V. Journal of Entomology and Zoology, 6, 1‒8.
  157. Kuwana, S.I. (1922) Studies on Japanese Monophlebinae. Contribution I: The genus Warajicoccus. Bulletin of Agriculture and Commerce, Imperial Plant Quarantine Station, Yokohama, 1, 1‒58.
  158. Lambdin, P.L. & Kosztarab, M.P. (1977) Morphology and systematics of the adult females of the genus Cerococcus (Homoptera: Coccoidea: Cerococcidae). Research Division Bulletin, Virginia Polytechnic Institute and State University, Blacksburg, 128, 1–252.
  159. Leonardi, G. (1908) [Second contribution to the knowledge of Italian scale insects]. Bollettino del R. Laboratorio di Entomologia Agraria di Portici, 3, 150‒191. [in Italian]
  160. Lindinger, L. (1907) [Identification board of German diaspidids]. Entomologische Blätter, 3, 4‒6. [in German]
  161. Lindinger, L. (1933) [Contributions to the knowledge of scale insects (Hemipt. - Homopt., Coccid.)]. Entomologischer Anzeiger, 13, 77‒166. [in German]
  162. Lindinger, L. (1936) [New contributions to the knowledge of scale insects (Coccidae)]. Entomologische Zeitschrift, 49, 444. [in German]
  163. Lindinger, L. (1957) [Another contribution to the synonymy of coccids]. Beiträge zur Entomologie. Berlin, 7, 543‒553. [in German]
  164. Linnaeus, C. (1758) Systema Naturae, per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Laurentii Salvii, Holmiae, (4) + 824 pp. https://doi.org/10.5962/bhl.title.542
  165. Lotfalizadeh, H. (2010) Some new data and corrections on Iranian encyrtid wasps (Hymenoptera: Chalcidoidea, Encyrtidae) fauna. Biharean Biologist, 4 (2), 173‒178.
  166. Lotfalizadeh, H. & Gharali, B. (2008) Pteromalidae (Hymenoptera: Chalcidoidea) of Iran: new records and a preliminary checklist. Entomofauna, 29 (6), 93‒120.
  167. Lotfalizadeh, H., Tavakoli-Korghond, G. & Mokhtari, A. (2019) On the parasitoid complex (Hymenoptera: Chalcidoidea) of Anophococcus abaii (Hemiptera: Eriococcidae), with description of Metaphycus anophococcusi Lotfalizadeh, n. sp. Annales de la Société entomologique de France, 55 (4), 317–326. https://doi.org/10.1080/00379271.2019.1621204
  168. Lotfalizadeh, H., Zargaran, M.R. & Taghizadeh, M. (2014) Species diversity of Coccoidea parasitoids wasps (Hym.: Chalcidoidea) in the northern parts of East-Azarbaijan province, Iran. North-Western Journal of Zoology, 10 (1), 60‒66.
  169. MacGillivray, A.D. (1921) The Coccidae. Tables for the Identification of the Subfamilies and Some of the More Important Genera and Species, together with Discussions. Scarab, Urbana, Illinois, 502 pp. https://doi.org/10.5962/bhl.title.8556
  170. Maleki, N. & Damavandian, M.R. (2015) Determination of economic injury level for first and second generations of Pulvinaria aurantii (Hem: Coccidae) in Thomson navel orange orchards. Arthropods, 4 (1), 13–21.
  171. Malumphy, C.P. (1991) A morphological and experimental investigation of the Pulvinaria vitis complex in Europe. Ph.D. thesis, University of London, London, 270 pp.
  172. Marchal, P. (1904) Sur une cochenille nouvelle. Annales de la Société entomologique de France, 73, 557‒561.
  173. Marchal, P. (1906) Sur deux espèces de cochenilles nouvelles (Hem. Hom.) récoltées en Algérie. Bulletin de la Société entomologique de France, 9, 143‒145. https://doi.org/10.3406/bsef.1906.23916
  174. Maskell, W.M. (1879) On some Coccidae in New Zealand. Transactions and Proceedings of the New Zealand Institute, 11 (1878), 187‒228.
  175. Maskell, W.M. (1893) Further coccid notes: with descriptions of new species from Australia, India, Sandwich Islands, Demerara, and South Pacific. Transactions and Proceedings of the New Zealand Institute, 25 (1892), 201‒252.
  176. Matile-Ferrero, D. (1978) Homoptères Coccoidea de l’Archipel des Comores. In: Faune Entomologique de l’Archipel des Comores. Matile, L. (Ed.). Mémoires du Museum national d’Histoire naturelle, New Series, Série A, Zoologie, 109, pp. 39‒70.
  177. Matov, A., Zahiri, R. & Holloway, J. (2008) The Heliothinae of Iran (Lepidoptera: Noctuidae). Zootaxa, 1763 (1), 1–37. https://doi.org/10.11646/zootaxa.1763.1.1
  178. Meyer, E.H.F. (1854) [History of Botany. Vol. III]. Velag der Gebrüder Bornträger, Konigsberg, xvi + 554 pp. [in German]
  179. Miller, D.R. & Gimpel, M.E. (1996) Nomenclatural changes in the Eriococcidae (Homoptera: Coccoidea). Proceedings of the Entomological Society of Washington, 98, 597‒606.
  180. Miller, D.R. & Gimpel, M.E. (1999) New combinations, new synonymy, and homonomy in the Eriococcidae, new homonymy and synonymy in the Cerococcidae, and transfer of Cancerococcus Koteja to the Margarodidae (Hemiptera: Coccoidea). Proceedings of the Entomological Society of Washington, 101 (1), 212‒218.
  181. Miller, D.R. & Miller, G.L. (1993a) Eriococcidae of the Eastern United States (Homoptera). Contributions of the American Entomological Institute, 27 (4), 1–91.
  182. Miller, D.R. & Miller, G.L. (1993b) Description of a new genus of scale insect with a discussion of relationships among families related to the Kermesidae (Homoptera: Coccoidea). Systematic Entomology, 18 (3), 237–251. https://doi.org/10.1111/j.1365-3113.1993.tb00664.x
  183. Miller, D.R., Rung, A. & Parikh, G. (2014) Scale Insects, edition 2, a tool for the identification of potential pest scales at U.S.A. ports-of-entry (Hemiptera, Sternorrhyncha, Coccoidea) ZooKeys, 431, 61–78. https://doi.org/10.3897/zookeys.431.7474
  184. Modeer, A. (1778) [About the fixed flight. Coccus]. Handlingar Gotheborgska Wetenskaps och Witterhets Samhallets, Wetenskaps Afdelingen, Gotheborg, Series 1, 1, 11–50. [in Sweedish]
  185. Moghaddam, M. (2009) Insects of Iran: the list of Coccoidea in the Hayk Mirzayans Insect Museum of Iranian Research Institute of Plant Protection (2). Hemiptera: Coccoidea. Publications, Plant Pests & Diseases Research Institute, Insect Taxonomy Research Department, No. 16, 1–45.
  186. Moghaddam, M. (2013) An annotated checklist of the scale insects of Iran (Hemiptera, Sternorrhyncha, Coccoidea) with new records and distribution data. ZooKeys, 334, 1–92. https://doi.org/10.3897/zookeys.334.5818
  187. Moghaddam, M. (2015) [Scale insects of ornamental plants in Iran (Hem.: Sternorrhyncha: Coccomorpha)]. Journal of Entomology Society of Iran, 34, 83–91. [in Persian]
  188. Moghaddam, M. (2017) The Scale Insects of Citrus in Iran (Hemiptera, Coccomorpha) (Identification, Biology, Natural Enemies and Distribution). Iranian Research Institute of Plant Protection, Tehran, 175 pp.
  189. Moghaddam, M. (2018) Description of a new species of Rhizococcus (Hemiptera: Coccomorpha: Acanthococcidae) from Iran. Acta Entomologica Musei Nationalis Pragae, 58 (2), 519–522. https://doi.org/10.2478/aemnp-2018-0040
  190. Moghaddam, M. (2019) Two new scale insect species (Hemiptera: Coccomorpha: Acanthococcidae and Pseudococcidae) and a new country record from protected areas in Iran. Zootaxa, 4706 (3), 451–460. https://doi.org/10.11646/zootaxa.4706.3.5
  191. Moghaddam, M. & Azad-Bakht, N.A. (2006) The first report of the genus and species of the scale insect, Exaeretopus tritici (Hem.: Cocciodea: Coccidae), from Iran. Journal of Entomological Society of Iran, 25 (2), 89–90.
  192. Moghaddam, M. & Faghih, H. (2019) A new species of soft scale insect (Hemiptera: Coccomorpha: Coccidae) on fig trees in Iran. Zootaxa, 4567 (3), 561–566. https://doi.org/10.11646/zootaxa.4567.3.7
  193. Moghaddam, M. & Mohammadi-Khoramabadi, A. (2015) The scale insects (Hemiptera: Coccomorpha) on Artemisia spp. (Asteraceae) in Iran. Journal of Insect Biodiversity and Systematics, 1 (1), 37–45.
  194. Moghaddam, M. & Nematian, M.R. (2021) New record of a scale insect pest from the genus Ceroplastes (Hemiptera, Coccomorpha, Coccidae) on Aglaonema commutatum (Araceae) from Iran. Journal of the Entomological Society of Iran, 41 (2), 175‒181.
  195. Moghaddam, M. & Tavakoli, M. (2010) Scale insects of the central Zagros region in Iran (Hemiptera: Coccoidea). Applied Entomology and Phytopathology, 77 (2), 27‒45.
  196. Moghaddam, M. & Watson, G.W. (2021) The scale insects of Iran, Part 1, The armoured scales (Hemiptera: Coccomorpha: Diaspididae). Zootaxa, 4907 (1), 1–276. https://doi.org/10.11646/zootaxa.4907.1.1
  197. Moghaddam, M. & Watson, G.W. (2022) The scale insects of Iran (Hemiptera: Coccomorpha), Part 2, The mealybugs (Pseudococcidae and Rhizoecidae) and Putoidae. Zootaxa, 5126 (1), 1–169. https://doi.org/10.11646/zootaxa.5126.1.1
  198. Moghaddam, M., Esfandiari, M. & Khosravi, M. (2015a) First record of Icerya aegyptiaca (Hemiptera: Coccoidea: Monophlebidae) from IranProceedings and Abstracts, 1st Iranian International Congress of Entomology, Tehran, Iran, 29‒31 August 2015, 42.
  199. Moghaddam, M., Sahraian, H. & Abbasi, J. (2015b) The first record of invasive soft scale of Saissetia miranda (Hemiptera: Coccoidea: Coccidae) and with an identification key to species of genus Saissetia in Iran. Proceedings and Abstracts, 1st Iranian International Congress of EntomologyTehran, Iran, 29‒31 August 2015, 228‒231.
  200. Mongue, A.J., Michaelides, S., Coombe, O., Tena, A., Kim, D.-S., Normark, B.B., Gardner, A., Hoodle, M.S. & Ross, L. (2021) Sex, males, and hermaphrodites in the scale insect Icerya purchasi. Evolution. [published online] https://doi.org/10.1101/2020.09.04.281618
  201. Morales, C.F. (1991) Margarodidae (Insecta: Hemiptera). Fauna of New Zealand / Ko te Aitanga Pepeke o Aotearoa, 21, 1‒123.
  202. Morrison, H. (1927) Descriptions of new genera and species belonging to the coccid family Margarodidae. Proceedings of the Biological Society of Washington, 40, 99‒109.
  203. Morrison, H. (1928) A classification of the higher groups and genera of the coccid family Margarodidae. United States Department of Agriculture Technical Bulletin, 52, 1‒239.
  204. Morrison, H. & Morrison, E.R. (1923) A selected bibliography of the Coccoidea. First Supplement. Miscellaneous Publication, United States Department of Agriculture, 987, 1‒44.
  205. Morrison, H. & Morrison, E.R. (1966) An annotated list of generic names of the scale insects (Homoptera: Coccoidea). Miscellaneous Publication, United States Department of Agriculture, 1015, 1‒206. https://doi.org/10.5962/bhl.title.65706
  206. Naeim Amini, S., Abbasipour, H. & Aghajanzadeh, S. (2014) Spatial distribution of the cottony camellia scale, Pulvinaria floccifera (Westwood) (Hemiptera: Coccidae) in the tea orchards. Journal of Plant Protection Research, 54 (1), 44–52. https://doi.org/10.2478/jppr-2014-0007
  207. Naeim Amini, S., Abbasipour, H., Aghajanzadeh, S. & Zamani, A. (2010) [First report of Lecanicillium lecanii and its sexual stage, from Iran]. Iran Journal Plant Pathology, 49 (3), 279‒280. [in Persian]
  208. Nan, N., Dean, G.J. & Wu, S.A. (2013) A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang and Hao) comb. n. Zootaxa, 3722 (2), 170–182. https://doi.org/10.11646/zootaxa.3722.2.3
  209. Nasonov, N.V. (1908) Steingelia gorodetskia, nov. gen. et nov. sp. nouveau genre et espèce des coccides du groupe Xylococcini. Annuaire du Musée Zoologique de l’Académie Impériale des Sciences de St. Pétersbourg, 13, 345‒352.
  210. Nast, J., Chudzicka, E., Klimaszewski, S.M., Wojciechowski, W., Czylok, A. & Koteja, J. (1990) Homoptera [Checklist of Animals of Poland]. Zaklad Narodowy im. Ossolinskich Wydawnictwo Polskiej Akademii Nauk. PAN, Kraków, 158 pp. [in Polish]
  211. Nehrangi, H. & Vahedi, H.A. (2015) Biology of Kermes quercus (Linnaeus) (Hem.: Coccoidea: Kermesidae) under field conditions of oak forests in Kermanshah district. In: Proceedings and Abstracts, 1st Iranian International Congress of Entomology, Tehran, Iran, 29‒31 August 2015, 133.
  212. Newstead, R. (1891) On some new or little known Coccidae found in England [No. 1?]. Entomologist’s Monthly Magazine, 26, 164–166.
  213. Newstead, R. (1892) On new or little known Coccidae, chiefly English (No. 2). Entomologist’s Monthly Magazine, 28, 141‒147. https://doi.org/10.5962/bhl.part.7647
  214. Newstead, R. (1894) Observations on Coccidae. No. 9. Entomologist’s Monthly Magazine, 30, 204‒207.
  215. Newstead, R. (1898) Observations on Coccidae. No. 17. Entomologist’s Monthly Magazine, 34, 92–99.
  216. Newstead, R. (1901) Observations on Coccidae. No. 19. Entomologist’s Monthly Magazine, 37, 81‒86. https://doi.org/10.5962/bhl.part.27060
  217. Newstead, R. (1903) Monograph of the Coccidae of the British Isles. Vol. 2. Ray Society, London, 270 pp.
  218. Nitsche, H. (1895) [The scale insects]. In: Lehrbuch der Mitteleuropäischen Forstinsektenkunde. Vol. 2. Holzel, Wien, pp. 737–1421. [in German]
  219. Olivier, G.A. (1791) Cochenille. Coccus. Genre d’insectes de la première section de l’ordre des Hemiptères. In: Olivier, G.A. (Ed.), Encylopédie Methodique, Histoire naturelle. Insectes. Tome 6. Pauckoucke, Paris, pp. 85–100.
  220. Ostovan, H. & Modaresi, H. (2006) Biology and natural enemies of Ceroplastes floridensis (Comstock) (Homoptera: Coccidae) in north of Iran. Abstract. 58th International Symposium on Crop Protection, Gent, Belgium, May 23, 2006. Communications in Agricultural and Applied Biological Sciences, 71 (2 Pt A), 151.
  221. Pellizzari, G. & Kozár, F. (2011) A new species of Greenisca and two new species of Ovaticoccus from Italy (Hemiptera Coccoidea Eriococcidae), with a key to European genera of Eriococcidae. Zootaxa, 3090 (1), 57–68. https://doi.org/10.11646/zootaxa.3090.1.4
  222. Pellizzari, G. & Williams, D.J. (2013) Simple rules on adjectival endings in zoological nomenclature and their use in scale insect names (Hemiptera: Sternorrhyncha: Coccoidea), with some corrections to combinations in common use. Zootaxa, 3710 (5), 401‒414. https://doi.org/10.11646/zootaxa.3710.5.1
  223. Pergande, T. (1898) The peach Lecanium. (Lecanium nigrofasciatum n. sp.). Bulletin United States Department of Agriculture, Bureau of Entomology, Washington, 18, 26‒29.
  224. Podsiadło, E. (2014) Identification characters of nymphal instars and adult female of Kermes quercus (Linnaeus, 1758) (Hemiptera: Kermesidae). Polskie Pismo Entomologiczne, 83, 259–268. https://doi.org/10.2478/pjen-2014-0020
  225. Pourjafari, K., Vahedi, H.A. & Darbemamieh, M. (2023) [The first record of Kermes spatulatus scale insect from Iran and study of morphological characteristics of adult insects and the first nymphal instars]. Phytopathology and Entomology, 91 (1), 73‒79.
  226. Ramachandra Rao, Y. (1922) The pests of the date palm in the Iraq. Mesopotamia Department of Agriculture Memoirs, 6 (1921), 1‒12.
  227. Ratzeburg, J.T.C. (1844) [The forest insects or illustration and description of those in the forests of Prussia and neighboring states as harmful or ...] Abtheilung des Königl. Nicolaische buchhandlung, Berlin, 301 pp. [in German]
  228. Rau, G.J. (1938) Four more new species of mealybugs from New York State. (Hemiptera: Kermidae and Pseudococcidae). Canadian Entomologist, 70, 157‒165. https://doi.org/10.4039/Ent70157-8
  229. Rezwani, A. & Radjabi, G. (1983) Biologie de Sphaerolecanium prunastri Boyer de Fonscolombe en Iran - Biology of Sphaerolecanium prunastri Boyer de Fonscolombe in Iran. Entomologie et Phytopathologie Appliquées, 51 (1/2), 17‒22.
  230. Riley, C.V. & Howard, L.O. (1890) Some new iceryas. Insect Life, 3, 92‒106.
  231. Rozedar, E., Vahedi, H., Mossadegh, M.S. & Samie, M.S. (2013) The scale insects (Hem.: Coccoideae) of Behbahan county and their morphological characters. Plant Protection, Scientific Journal of Agriculture, 36 (2), 41–68.
  232. Rübsaamen, E.H. (1902) [Information on the zoocecidians collected by Mr. J. Bornmüllier in the Orient]. Zoologische Jahrbücher Abteilung für Systematik, Geog., und Biol. Der Tiere, 16, 243‒336. [in German]
  233. Russell, L.M. (1941) A classification of the scale insect genus Asterolecanium. Miscellaneous Publication, United States Department of Agriculture, 424, 1‒319. https://doi.org/10.5962/bhl.title.65621
  234. Safar Alizadeh, M.H. & Bahador, M. (1987) [Presentation of a pest of wheat (Porphyrophora tritici Bod.)]. Journal of Entomological Society of Iran, 9, 1‒2 + 7‒8. [29–37 in Persian]
  235. Sanders, J.G. (1906) Catalogue of recently described Coccidae. United States Department of Agriculture, Bureau of Entomology, Technical Series, 12, 1‒18.
  236. Schmutterer, H. (1952) [The ecology of the coccids (Homoptera, Coccoidea) of Franconia. 2nd section]. Zeitschrift für Angewandte Entomologie, 33 & 34, 369–420 + 544–584 & 65–100. [in German] https://doi.org/10.1111/j.1439-0418.1953.tb00689.x
  237. Schmutterer, H. (1980) [On the status of research into scale insects (Homoptera, Coccoidea) in the Federal Republic of Germany]. Mitteilungen der Deutschen Gesellschaft für Allgemeine und Angewandte Entomologie. Bremen, 2, 49–56. [in German]
  238. Scott, H. (1907) On Cercococcus eremobius, gen. et. sp. nov., an aberrant form of Coccidae. Transactions of the Linnean Society of London, Zoology, 9, 455‒464. https://doi.org/10.1111/j.1096-3642.1907.tb00454.x
  239. Shirazi, M., Vahedi, H.A., Mirmoayedi, A.-N., Masoumi, S.M. & Jalilvand, K. (2013) Scale insects recorded on ornamental plants in urban areas of Kermanshah, Iran. African Journal of Agricultural Research, 8 (16), 1381–1383. https://doi.org/10.5897/AJAR11.1641
  240. Signoret, V. (1872a) Essai sur les cochenilles ou gallinsectes (Homopteres - Coccides), 9e partie. Annales de la Societe entomologique de France, Série 5, 2, 33‒46.
  241. Signoret, V. (1872b) Essai sur les cochenilles ou gallinsectes (Homoptères - Coccides), 8e partie. Annales de la Societe entomologique de France, Série 5, 1, 421‒434.
  242. Signoret, V. (1873a) Essai sur les cochenilles ou gallinsectes (Homoptères - Coccides), 11e partie. Annales de la Société entomologique de France, Série 5, 3, 395‒448.
  243. Signoret, V. (1873b) Essai sur les cochenilles ou gallinsectes (Homoptères - Coccides), 10e partie. Annales de la Societe entomologique de France, Série 5, 3, 27‒48.
  244. Signoret, V. (1875) Essai sur les cochenilles ou gallinsectes (Homoptères - Coccides), 14e partie. Annales de la Societe entomologique de France, Série 5, 5, 15‒40.
  245. Signoret, V. (1876a) Essai sur les cochenilles ou gallinsectes (Homoptères - Coccides), 16e partie. Annales de la Société entomologique de France, Série 5, 5, 346‒373.
  246. Signoret, V. (1876b) Essai sur les cochenilles ou gallinsectes (Homoptères - Coccides), 17e partie. Annales de la Société entomologique de France, 5, 374‒394.
  247. Signoret, V. (1877) Essai sur les cochenilles ou gallinsectes (Homoptères - Coccides), 18e partie. Annales de la Société entomologique de France, Série 5, 6 (1876), 591‒676.
  248. Silvestri, F. (1939) Compendio di Entomologia Applicata. Parte Speciale. Tipografia Bellavista, Portici, 974 pp.
  249. Spodek, M. & Ben-Dov, Y. (2014) A taxonomic revision of the Kermesidae (Hemiptera: Coccoidea) in Israel, with a description of a new species. Zootaxa, 3781 (1), 1–99. https://doi.org/10.11646/zootaxa.3781.1.1
  250. Spodek, M., Ben-Dov, Y., Ghanim, M. & Mendel, Z. (2012) Morphological and molecular taxonomy of Nidularia balachowskii Bodenheimer (Hemiptera, Coccoidea, Kermesidae) with notes on its life history in Israel. ZooKeys, 254, 23–45. https://doi.org/10.3897/zookeys.254.3959
  251. Stickney, F.S. (1934) The external anatomy of the red date scale Phoenicococcus marlatti Cockerell, and its allies. United States Department of Agriculture Technical Bulletin, 404, 1–162.
  252. Stumpf, C.F. & Lambdin, P.L. (2006) Pit scales (Sternorrhyncha: Coccoidea) of North and South America. Tennessee Agricultural Experiment Station, University of Tennessee Institute of Agriculture, Knoxville, Tennessee, xix + 231 pp.
  253. Šulc, K. (1895) On a new genus and species of Coccidae, Ortheziola vejdovskyi, n. g., n. sp. Sitzungsberichte der Königlichen Böhmischen Gesellschaft der Wissenschaften in Prag, 44, 1–9.
  254. Šulc, K. (1908) Towards the better knowledge of the genus Lecanium. Entomologist’s Monthly Magazine, 44, 36.
  255. Šulc, K. (1932) [Czechoslovak species of the genus Lecanium (gen. Lecanium, Coccidae, Homoptera)]. Acta Societatis Scientiarum Naturalium Moravicae, 7, 1‒134. [in Slovakian]
  256. Talebi, A.A., Mohammadi Khoramabadei, A. & Rakhshani, E. (2011) Checklist of eulophid wasps (Insecta: Hymenoptera: Eulophidae) of Iran. Check List, 7 (6), 708‒719. https://doi.org/10.15560/11008
  257. Tanaka, H. & Amano, H. (2007) Redescription of Pulvinaria floccifera (Insecta: Hemiptera: Coccidae). Species Diversity, 12, 211‒215. https://doi.org/10.12782/specdiv.12.211
  258. Tanaka, H. & Kamitani, S. (2022) The identities of Pulvinaria camelicola Signoret 1873, stat. nov., and the related species, Pulvinaria floccifera (Westwood 1870) (Hemiptera: Coccidae) in Japan. Zootaxa, 5115 (3), 438–450. https://doi.org/10.11646/zootaxa.5115.3.8
  259. Tang, F.T. (1977) [The scale insects of horticulture and forest of China. Vol. I]. The Institute of Gardening, Forestry Science of Shenyang, Liaoning, 259 pp. [in Chinese]
  260. Tang, F.T. (1991) [The Coccidae of China]. Shanxi United Universities Press, Taiyuan, 377 pp. [in Chinese]
  261. Tang, F.T. & Hao, J.J. (1995) [The Margarodidae and others of China]. Chinese Agricultural Science Technology Press, Beijing, 738 pp. [in Chinese]
  262. Tao, C.C.C., Wong, C.Y. & Chang, Y.C. (1983) Monograph of Coccidae of Taiwan, Republic of China (Homoptera: Coccoidea). Journal of the Taiwan Museum, 36, 57‒107.
  263. Targioni Tozzetti, A. (1866) [How certain scale insects are the cause of some honeydews on plants, and of some rusts; and how the cochineal of the fig tree gives in abundance a kind of wax]. Atti della R. Accademia dei Georgofili (New Series), 13, 115‒137; Appendix 138‒146. [in Italian]
  264. Targioni Tozzetti, A. (1868) [Introduction to the second memoir for studies on scale insects, and catalog of the genera and species of the coccid family]. Atti della Società italiana di Scienze naturali, Milano, 11, 721‒738. [in Italian]
  265. Targioni Tozzetti, A. (1869) [On two genera of scale insects (Coccidae)]. Bollettino della Società Entomologica italiana, Firenze, 1, 257‒267. [in Italian]
  266. Tereznikova, E.M. (1981) [Scale insects: Eriococcidae, Kermesidae and Coccidae]. Fauna Ukraini. Akademiya Nauk Ukrainskoi RSR. Institut Zoologii, Kiev, 20, 1‒215. [in Russian]
  267. Ter-Grigorian, T. (1983) [On the fauna of the felt scales (Homoptera, Coccinea, Eriococcidae) in Armenia]. Biologicheskii Zhurnal Armenii, 36 (10), 876‒883. [in Armenian]
  268. Torabi, M., Vahedi, H.A. & Hodgson, C.J. (2010) Preliminary survey of the scale insects fauna in Kermanshah, western Iran. Entomologia Hellenica, 19, 153‒162. https://doi.org/10.12681/eh.11584
  269. Unruh, C.M. & Gullan, P.J. (2008a) Molecular data reveal convergent reproductive strategies in iceryine scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing the re-interpretation of morphology and a revised generic classification. Systematic Entomology, 33, 8‒50.
  270. tps://doi.org/10.1111/j.1365-3113.2007.00404.x
  271. Unruh, C.M. & Gullan, P.J. (2008b) Identification guide to species in the scale insect tribe Iceryini (Coccoidea: Monophlebidae). Zootaxa, 1803 (1), 1‒106. https://doi.org/10.11646/zootaxa.1803.1.1
  272. Vahedi, H.A. (2002) A revision of the genus Porphyrophora (Hemiptera: Coccoidea: Margarodidae) with particular reference to the Middle East and with a discussion on the relationships of the hypogaeic Margarodids. Ph.D. thesis. Department of Agricultural Science, Imperial College at Wye, Wye, Ashford, Kent, 358 pp.
  273. Vahedi, H.A. (2004) Note on the biology of Porphyrophora cynodontis under field and laboratory conditions in Kermanshah, Iran. (Abstract.) In: Erkiliç, L.B. & Kaydan, M.B. (Eds.), Proceedings of the X International Symposium on Scale Insect Studies ISSIS-X, Plant Protection Research Institute, Adana, Turkey, 19‒23 April, pp. 257.
  274. Vahedi, H.A. & Hodgson, C.J. (2007) Some species of the hypogeal scale insect Porphyrophora Brandt (Hemiptera: Sternorrhyncha: Coccoidea: Margarodidae) from Europe, the Middle East and North Africa. Systematics and Biodiversity, 5 (1), 23–122. https://doi.org/10.1017/S1477200006002039
  275. Vea, I.M. & Grimaldi, D.A. (2012) Phylogeny of ensign scale insects (Hemiptera: Coccoidea: Ortheziidae) based on the morphology of recent and fossil females. Systematic Entomology, 37, 758–783. https://doi.org/10.1111/j.1365-3113.2012.00638.x
  276. Vea, I.M. & Grimaldi, D.A. (2016) Putting scales into evolutionary time: the divergence of major scale insect lineages (Hemiptera) predates the radiation of modern angiosperm hosts. Scientific Reports, 6, 23487. https://doi.org/10.1038/srep23487
  277. von Heyden, C. (1860) [From my entomological diaries. (1). On the natural history of coccids]. Korrespondenzblatt für Sammler von Insekten, 1, 89–90. [in German]
  278. Walker, F. (1852) List of the specimens of homopterous insects in the collection of the British Museum, Part IV. British Museum (Natural History), London, 1188 pp.
  279. Walker, F. (1858) List of the specimens of homopterous insects in the collection of the British Museum. British Museum (Natural History), London, 369 pp.
  280. Watson, G.W. (2022a) Towards identification of the scale insects (Hemiptera: Coccomorpha) of continental Africa: 2. Checklists and keys to six archaeococcoid families. Zootaxa, 5105 (3), 301–356. https://doi.org/10.11646/zootaxa.5105.3.1
  281. Watson, G.W. (2022b) 4.13.1 Red date scale, Phoenicococcus marlatti (Cockerell, 1899). In: Kondo, T. & Watson, G.W. (Eds.), Encyclopedia of Scale Insect Pests. CAB International, Wallingford, pp. 413‒414.
  282. Westwood, J.O. (1870) [The camellia coccus Coccus flocciferus Westw]. Gardeners’ Chronicle and Agricultural Gazette, 10, 308.
  283. Williams, D.J. (1957) The status of Coccus palmae Haworth and the identity of Lecanium coffeae Walker (Coccoidea: Homoptera). The Entomologist, 90, 314‒315.
  284. Williams, D.J. (1977) A new species of Exaeretopus Newstead (Homoptera: Coccidae) attacking wheat in Iraq. Bulletin of Entomological Research, 67, 281‒284. https://doi.org/10.1017/S0007485300011093
  285. Williams, D.J. (1985) The British and some other European Eriococcidae (Homoptera: Coccoidea). Bulletin of the British Museum (Natural History) Entomology, 51, 347–393.
  286. Williams, D.J., Gullan, P.J., Miller, D.R., Matile-Ferrero, D. & Han, S.I. (2011) A study of the scale insect genera Puto Signoret (Hemiptera: Sternorrhyncha: Coccoidea: Putoidae) and Ceroputo Šulc (Pseudococcidae) with a comparison to Phenacoccus Cockerell (Pseudococcidae). Zootaxa, 2802 (1), 1–22. https://doi.org/10.11646/zootaxa.2802.1.1
  287. Williams, D.J. & Watson, G.W. (1988) The Scale Insects of the Tropical South Pacific Region. Part 1. The Armoured Scales (Diaspididae). CAB International, Wallingford, 290 pp.
  288. Williams, D.J. & Watson, G.W. (1990) The Scale Insects of the Tropical South Pacific Region. Part. 3: The Soft Scales (Coccidae) and Other Families. CAB International, Wallingford, 267 pp.