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Type: Monograph
Published: 2026-01-09
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First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines

Department of Biological Sciences; College of Science and Mathematics; Mindanao State University-Iligan Institute of Technology; Tibanga; Iligan City 9200; Philippines.; Premier Research Institute of Science and Mathematics; Mindanao State University-Iligan Institute of Technology. Andres Bonifacio Avenue; Tibanga; 9200 Iligan City; Philippines.
Department of Biological Sciences; College of Science and Mathematics; Mindanao State University-Iligan Institute of Technology; Tibanga; Iligan City 9200; Philippines.; Premier Research Institute of Science and Mathematics; Mindanao State University-Iligan Institute of Technology. Andres Bonifacio Avenue; Tibanga; 9200 Iligan City; Philippines.
Department of Biological Sciences; College of Science and Mathematics; Mindanao State University-Iligan Institute of Technology; Tibanga; Iligan City 9200; Philippines.; Premier Research Institute of Science and Mathematics; Mindanao State University-Iligan Institute of Technology. Andres Bonifacio Avenue; Tibanga; 9200 Iligan City; Philippines.
Coleoptera Bibliography biodiversity systematic records endemic taxonomy

Abstract

More than a century ago, Wilhelm Schultze published his seminal work, A Catalogue of Philippine Coleoptera, documenting over 290 species of scarab beetles. Since then, no comprehensive updates have been made. This catalog seeks to provide an exhaustive record of all scarab species reported in the Philippines from the early 1800s to the present. Currently, scarab beetles in the country are represented by seven subfamilies, 104 genera, and 787 species. Notably, numerous species listed here are known solely from their original descriptions. With the growing interest in the taxonomy and biodiversity of Philippine scarab beetles, we anticipate the potential rediscovery of certain species and significant taxonomic revisions through the combination or splitting of the country’s currently known genera and species of scarab beetles. Ultimately, this compilation should be regarded as an evolving reference rather than a definitive work, serving as a foundation for future research and discoveries in Philippine scarab biodiversity.

 

References

  1. Ahrens, D. & Fabrizi, S. (2016) A monograph of the Sericini of India (Coleoptera: Scarabaeidae). Zoologisches Forschungsmuseum Alexander Koenig-Leibniz-Institut für Biodiversität der Tiere, Bonn, 355 pp.
  2. Alexis, R. & Delpont, M. (2000) 27e contribution à l’étude des Cetoniidae: Description de sept nouvelles espèces et de deux sous-espèces appartenant à la faune orientale et australo-papoue (Coleoptera, Scarabaeoidea, Cetoniidae, Cetoniinae, Schizorhinini). Lambillionea, 100 (1), 53–68.
  3. Allard, V. (1995) Schizorhinini 2. Les Coléoptères du Monde, 24. Sciences Nat., Venette, 136 pp.
  4. Antoine, P. (1993) Révision des Microvalgini orientaux (Coleoptera: Cetoniidae: Valginae). Annales de la Société Entomologique de France, New Series, 29 (3), 329–340. https://doi.org/10.1080/21686351.1993.12277694
  5. Antoine, P. (1996) Quelques espèces nouvelles ou peu connues de la famille des Cetoniidae V (Coleoptera, Cetoniidae). Coléoptères, 2 (6), 53–68.
  6. Antoine, P. (2002) Glycyphana (Glycyphaniola) delponti, nouvelle espèce de Cetoniidae (Coleoptera, Scarabaeoidea). Cetoniimania, 2 (1), 12–13.
  7. Anichtchenko, A., Minkina, Ł., Vasiljeva, A. & Medina, M.N.D. (2021) A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae, 61 (1), 99–111. https://doi.org/10.37520/aemnp.2021.006
  8. Arnaud, P. (1980) Description d’un nouveau Phaedimus des Îles Philippines (Coleoptera Cetoniidae). Revue Française d’Entomologie, New Series, 2 (3), 123–125.
  9. Arnaud, P. (1982) Description de trois espèces nouvelles de Cetonides asiatiques (Col. Scarabaeidae). Miscellanea Entomologica, 49, 117–120.
  10. Arnaud, P. (1985) Description de deux espèces nouvelles de Cetonidae de l’île Palawan (Philippines). Bulletin de la Société des Sciences Naturelles, 46, 18–19.
  11. Arnaud, P. (1987a) Diagnose préliminaire à la description de Cetonides. Bulletin de la Société des Sciences Naturelles, 56, 27.
  12. Arnaud, P. (1987b) Description de deux espèces nouvelles de Cetonidae des Philippines. Bulletin de la Société des Sciences Naturelles, 55, 24–25.
  13. Arnaud, P. (1987c) Diagnose preliminaire de cinq nouvelles espèces de Cetonidae et d’une nouvelle espèce de Lucanidae des îles Phlippines. Bulletin de la Société des Sciences Naturelles, 53, 4.
  14. Arnaud, P. (1989a) Nouvelles Cétoines des Philippines. Bulletin de la Société des Sciences Naturelles, 62, 1–3.
  15. Arnaud, P. (1989b) Nouvelles Cétoines du sud-est asiatique. Bulletin de la Société des Sciences Naturelles, 64, 23–25.
  16. Arnaud, P. (1990) Description de Cétoines des Philippines. Bulletin de la Société des Sciences Naturelles, 66, 8.
  17. Arnaud, P. (1992) Description de nouvelles Cétoines des Philippines. Bulletin de la Société des Sciences Naturelles, 75–76, 29–32.
  18. Arnaud, P. (1996) Description d’une nouvelle espèce d’Euglypta (Col Cetonidae). Besoiro, 2, 5.
  19. Arnaud, P. (2000) Description de Cetoniidae nouveaux des Philippines. Besoiro, 5, 4–5.
  20. Arnaud, P. (2004) Description de nouvelles espèces de Cetoniidae des Philippines, suivie de notes synonymiques ou complémentaires sur quelques espèces peu connues. Besoiro, 11, 2–7.
  21. Arnaud, P. (2019) Description de nouvelles espèces de Cetoniidae des Philippines. Besoiro, 27, 3–8.
  22. Arnaud, P. & Delpont, M. (2001) Première contribution à l’étude du genre Thaumastopeus Kraatz 1885 aux Philippines. Cetoniimania, 1 (3), 73–80.
  23. Arrow, G.J. (1899) Notes on the rutelid genera Anomala, Mimela, Popillia, and Strigoderma. In: Entomological Society of London (Ed.), Transactions of the Entomological Society of London for the year 1899. Richard Clay & Sons, Limited, London and Bungay, London, pp. 271–276. https://doi.org/10.1111/j.1365-2311.1899.tb00668.x
  24. Arrow, G.J. (1905) On some Oriental Aphodiid Coleoptera of the Rhyparus group, with description of a new genus. The Annals and Magazine of Natural History, 7 (15), 534–540. https://doi.org/10.1080/03745480509442847
  25. Arrow, G.J. (1910) The Fauna of British India, including Ceylon and Burma. Coleoptera: Lamellicornia (Cetoniinae and Dynastinae). Taylor and Francis, London, 322 pp.
  26. Arrow, G.J. (1915) Note on the nomenclature of certain species of Ruteline Coleoptera. The Annals and Magazine of Natural History, Series 8, 16 (93), 231–232. https://doi.org/10.1080/00222931508693709
  27. Arrow, G.J. (1931a) Fauna of British India, including Ceylon and Burma. Coleoptera: Lamellicornia. Part III (Coprinae).Taylor and Francis, London, 428 pp.
  28. Arrow, G.J. (1931b) Two new species of lamellicorn beetles belonging to the genus Peltonotus. The Annals and Magazine of Natural History, Series 10, 8 (48), 611–612. https://doi.org/10.1080/00222933108673453
  29. Arrow, G.J. (1937a) Scarabaeidae: Dynastinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 156. W. Junk, Berlin, pp. 1–124.
  30. Arrow, G.J. (1937b) Systematic notes on beetles of the subfamily Dynastinae, with descriptions of a few new species in the British Museum collection (Coleoptera). Transactions of the Entomological Society of London, 86, 35–58. https://doi.org/10.1111/j.1365-2311.1937.tb00246.x
  31. Baer, G.A. (1886) Catalogue des Coléoptères des Iles Philippines. Annales de la Société Entomologique de France, 6 (6), 97–200.
  32. Balthasar, V. (1944) Vier neue Arten der Gattung Onthophagus Latr. 78. Beitrag zür Kenntnis der Scarabaeiden (Col.). Sborník Entomologického Oddělení Národního Musea v Praze, 21–22, 90–94.
  33. Balthasar, V. (1959) Neue Onthophagus-Arten (97. Beitrag zur Kenntnis der Scarabaeidae, Col.). Entomologische Blätter, 55, 186–195.
  34. Balthasar, V. (1963a) Monographie der Scarabaeidae und Aphodidae der palaearktischen und orientalischen Region. Band 1. Tschechoslowakische Akademie der Wissenschaften, Prag, 391 pp.
  35. Balthasar, V. (1963b) Monographie der Scarabaeidae und Aphodidae der palaearktischen und orientalischen Region. Band 2. Coprinae (Onitini, Oniticellini, Onthophagini). Tschechoslowakische Akademie der Wissenschaften, Prag, 627 pp.
  36. Balthasar, V. (1964) Monographie der Scarabaeidae und Aphodidae der palaearktischen und orientalischen Region. Band 3. Coleoptera: Lamellicornia Aphodiidae. Tschechoslowakische Akademie der Wissenschaften, Prag, 652 pp.
  37. Barbero, E., Palestrini, C. & Roggero, A. (2011) Tibiodrepanus tagliaferrii - a new Afrotropical Drepanocerina species (Coleoptera: Scarabaeidae: Oniticellini), with notes on phylogeny and distribution of the genus. Zootaxa, 2923 (1), 27–47. https://doi.org/10.11646/zootaxa.2923.1.2
  38. Bezděk, A. (2004) Catalogue of Diplotaxini (Coleoptera: Scarabaeidae: Melolonthinae) of the Old World. Zootaxa, 463 (1), 1–90. https://doi.org/10.11646/zootaxa.463.1.1
  39. Bezděk, A. & Hájek, J. (2010a) Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic. Scarabaeidae: Dynamopodinae, Dynastinae, Melolonthinae, and Rutelinae. Acta Entomologica Musei Nationalis Pragae, 50 (1), 279–320.
  40. https://doi.org/10.1515/aemnp-2017-0075
  41. Bezděk, A. & Hájek, J. (2010b) Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic. Scarabaeidae: Cetoniinae (including Trichiini) and Valginae. Acta Entomologica Musei Nationalis Pragae, 50 (2), 629–655.
  42. Blanchard, É. (1842) Catalogue des Cétonides de la collection du Muséum d’Histoire Naturelle à Paris. Poussielgue, Paris, 23 pp.
  43. Blanchard, É. (1850–1851) Catalogue des collections entomologiques du Muséum d’Histoire naturelle de Paris. Classe des Insectes, Ordre des Coléoptères. Tome I. Gide et Baudry, Paris, 240 pp.
  44. Blanchard, É. (1853) Voyage au Pôle Sud et dans l’Océanie, sur les corvettes “l’Astrolabe” et “la Zélée”; exécuté par ordre du Roi pendant les années 1837–1838–1839–1840, sous le commandement de M.J. Dumont d’Urville, capitaine de vaisseau; publié par ordre du gouvernement, sous la direction supérieure de M. Jacquinot, capitaine de vaisseau, commandant de la Zélée. Zoologie, par MM. Hombron et Jacquinot. Tome quatrième. Description des Insectes. Gide et J. Baudry, Paris, 422 pp.
  45. Boheman, C.H. (1858) Coleoptera. In: Virgin, C.A. (Ed.), Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden under Befäl af C.A. Virgin, åren 1851–1853. Vetenskapliga Iakttagelser på H.M. Konung Oscar den Förstes Befallning utgifna af K. Svenska Vetenskaps Akademien. Andra Delen, Zoologi. 1. Insecta. P.A. Norstedt & Söner, Stockholm, pp. 1–617.
  46. Bordat, P., Paulian, R. & Pittino, R. (1990) Insectes Coléoptères Aphodiidae. Faune de Madagascar, 74, 1–255.
  47. Boucomont, A. (1913) Note synonymique sur quelques Onthophagus (Col. Scarabaeidae) de Malaisie. Bulletin de la Société Entomologique de France, 18 (17), 418.
  48. https://doi.org/10.3406/bsef.1913.25417
  49. Boucomont, A. (1914) Les Coprophages de l’Archipel Malais (Coléopt.). Annales de la Société Entomologique de France, 83, 238–350.
  50. https://doi.org/10.1080/21686351.1914.12279408
  51. Boucomont, A. (1920) Coléoptères coprophages nouveaux d’Asie et de Malaisie (Scarabaeidae). Annales de la Société Entomologique de France, 88, 307–320.
  52. https://doi.org/10.1080/21686351.1919.12279931
  53. Boucomont, A. (1921) Description de Coprophages (Col.) des Îles Philippines et de Bornéo. Bulletin de la Société Entomologique de France, 26 (7), 88–92.
  54. https://doi.org/10.3406/bsef.1921.26754
  55. Boucomont, A. (1924a) Les Onthophagus (Coleoptera, Scarabaeidae) des Iles Philippines. The Philippine Journal of Science, 24, 669–681.
  56. Boucomont, A. (1924b) Synonymies et homonymies de Lamellicornes Coprophages (Col.). Bulletin de la Société Entomologique de France, 29 (10), 114–115.
  57. https://doi.org/10.3406/bsef.1924.27327
  58. Boucomont, A. (1925) Lamellicornes coprophages nouveaux des Îles Philippines. Bulletin de la Société Entomologique de France, 30 (9), 151–154.
  59. https://doi.org/10.3406/bsef.1925.27496
  60. Bourgoin, A. (1929) Note sur Cetonia dubia Wallace (Col. Scarabaeidae). Bulletin de la Société Entomologique de France, 34 (17), 270–272.
  61. https://doi.org/10.3406/bsef.1929.28201
  62. Bourgoin, A. (1930) Communications. Description de deux Cétonides nouveaux (Col. Scarabaeidae). Bulletin de la Société Entomologique de France, 35 (19), 295–298.
  63. https://doi.org/10.3406/bsef.1930.28353
  64. Bourgoin, A. (1931) Deux Astraea nouveaux (Col. Scarabaeidae). Bulletin de la Société Entomologique de France, 36 (11), 176–178.
  65. https://doi.org/10.3406/bsef.1931.28442
  66. Brenske, E. (1892a) Beitrag zur Kenntnis der Gattungen Lepidiota und Leucopholis. Berliner Entomologische Zeitschrift, 37 (1), 33–62.
  67. https://doi.org/10.1002/mmnd.18920370109
  68. Brenske, E. (1892b) Neue Arten der Coleopteren-Gattung Holotrichia (Lachnosterna). Berliner Entomologische Zeitschrift, 37, 159–192.
  69. https://doi.org/10.1002/mmnd.18920370205
  70. Brenske, E. (1892c) Ueber einige neue Gattungen und Arten der Melolonthiden. Entomologische Nachrichten, 18, 151–159.
  71. https://doi.org/10.1002/mmnd.47918920205
  72. Brenske, E. (1894a) Die Melolonthiden der palaearktischen und orientalischen Region im Königlichen naturhistorischen Museum zu Brüssel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Mémoires de la Société Entomologique de Belgique, 2, 3–87.
  73. Brenske, E. (1894b) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38 (1893), 347–358.
  74. https://doi.org/10.1002/mmnd.18940380310
  75. Brenske, E. (1896) Neue Melolonthiden aus Africa und Asien. Stettiner Entomologische Zeitschrift, 57, 178–205.
  76. https://doi.org/10.1002/mmnd.48018960708
  77. Brenske, E. (1899) Die Serica-Arten der Erde. III. Berliner Entomologische Zeitschrift, 44, 161–272.
  78. https://doi.org/10.1002/mmnd.18990440303
  79. Brown, R.M., Siler, C.D., Oliveros, C.H., Esselstyn, J.A., Diesmos, A.C., Hosner, P.A., Linkem, C.W., Barley, A.J., Oaks, J.R., Sanguila, M.B., Welton, L.J., Blackburn, D.C., Moyle, R.G., Peterson, A.T. & Alcala, A.C. (2013) Evolutionary processes of diversification in a model island archipelago. Annual Review of Ecology, Evolution, and Systematics, 44 (1), 411–435.
  80. https://doi.org/10.1146/annurev-ecolsys-110411-160323
  81. Bunalski, M. (2022) Melolonthidae (Coleoptera: Scarabaeoidea) of the Palaearctic and Oriental Regions. II. Annotated catalogue of the genus Sophrops Fairmaire, 1887. Polish Journal of Entomology, 91 (3), 119–136.
  82. https://doi.org/10.5604/01.3001.0015.9948
  83. Burmeister, H.C.C. (1842) Handbuch der Entomologie. Dritter Band. Coleoptera Lamellicornia, Melitophila. Enslin, Berlin, 828 pp.
  84. Burmeister, H.C.C. (1844) Handbuch der Entomologie. Vierter Band. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia, Anthobia et Phyllophaga systellochela. Enslin, Berlin, xii + 587 + [1] pp.
  85. Burmeister, H.C.C. (1847) Handbuch der Entomologie. Fünfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Enslin, Berlin, viii + 584 pp.
  86. Burmeister, H.C.C. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie, Fortsetzung. Zweite Abtheilung: Coleoptera Lamellicornia Phyllophaga chaenochela. Enslin, Berlin, x + 569 + [1] pp.
  87. Burmeister, H.C.C. & Schaum, H.R. (1841) Kritische Revision der Lamellicornia melitophila (second part). Zeitschrift für die Entomologie, 3, 226–282.
  88. Calcetas, O.A. (2019) Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines. Zootaxa, 4619 (2), 330–346. https://doi.org/10.11646/zootaxa.4619.2.8
  89. Calcetas, O.A. (2023) Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896. European Journal of Taxonomy, 890, 184–203.
  90. https://doi.org/10.5852/ejt.2023.890.2261
  91. Calcetas, O.A. & Adorada, J.R. (2017) Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines. Zootaxa, 4232 (1), 85–103. https://doi.org/10.11646/zootaxa.4232.1.6
  92. Calcetas, O.A., Adorada, J.L. & Amalin, D. (2024) First Record of Infestation of Amphitrichia flachi (Brenske, 1892) (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini) on Cacao (Theobroma cacao L.) in the Philippines. The Philippine Journal of Science, 153 (3), 965–971.
  93. https://doi.org/10.56899/153.03.19
  94. Calcetas, O.A., Adorada, J.L., Adorada, J.R., Caoili, B.L., Rosales, A.M. & Dimapilis, E.F. (2021) New Records of Scarab Insect Pests of Cacao (Theobroma cacao L.) in the Philippines. The Philippine Journal of Science, 150 (5), 1197–1206.
  95. https://doi.org/10.56899/150.05.30
  96. Calcetas, O.A., Adorada, J.R. & Navasero, M.V. (2020) Review of the genus Exopholis Motschulsky, 1859 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines. Zootaxa, 4859 (4), 527–536. https://doi.org/10.11646/zootaxa.4859.4.4
  97. Candèze, E.C.A. (1869) Diagnoses de quelques Rutélides nouvelles. Coleopterologische Hefte, 5, 41–45.
  98. Cartwright, O.L. & Gordon, R.D. (1971) Coleoptera: Scarabaeidae. Insects of Micronesia, Series 17, 17 (4), 257–296.
  99. Červenka, R. (1994) Contribution to the knowledge of Aphodiidae II. New subgenus and two new species of Genus Aphodius Illiger (Coleoptera: Scarabaeoidea: Aphodiidae). Folia Heyrovskyana, 2 (8), 111–115.
  100. Chandra, K. & Gupta, D. (2013) Taxonomic studies on dung beetles (Coleoptera: Scarabaeidae, Geotrupidae, Hybosoridae) of Chhattisgarh, India. Munis Entomology & Zoology, 8 (1), 331–360.
  101. Chapin, E.A. (1931a) Adoretus luridus Blanchard and its near relatives in the Philippine Islands. Journal of the Washington Academy of Sciences, 21, 305–308.
  102. Chapin, E.A. (1931b) New species of Melolonthinae Scarabaeidae from the Philippine Islands. Journal of the Washington Academy of Sciences, 21, 309–314.
  103. Chen, K.P., Yang, M.M. & Tsai, J.F. (2002) Dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Lanyu. Collection and Research, 15, 53–62.
  104. Cheung, K., Bai, M., Leung, M., Lee, Y., Chan, K. & Yang, X. (2018) Scarabaeinae (Coleoptera: Scarabaeidae) from Hong Kong. Zoological Systematics, 43 (3), 233–267.
  105. Chevrolat, L.A.A. (1841) Description de trente-quatre espèces de Coléoptères de Manille et d’un Tricondyle de Ceylan. Revue Zoologique, par La Société Cuvierienne, 4, 221–228.
  106. Dalla Torre, K.W. (1912a) Melolonthinae I. Coleopterorum Catalogus. Vol. 45. W. Junk & S. Schenkling, Berlin, 84 pp.
  107. Dalla Torre, K.W. (1912b) Melolonthinae II. Coleopterorum Catalogus. Vol. 47. W. Junk & S. Schenkling, Berlin, 50 pp.
  108. Dalla Torre, K.W. (1912c) Melolonthinae III. Coleopterorum Catalogus. Vol. 49. W. Junk & S. Schenkling, Berlin, 156 pp.
  109. Dechambre, R.P. (1975) Désignation de types et note synonymique sur des Dynastinae (Col. Scarabaeidae). Bulletin de la Société Entomologique de France, 80 (3–4), 84–89.
  110. https://doi.org/10.3406/bsef.1975.21443
  111. Dellacasa, G. (1983) Taxonomic studies on Aphodiinae: X. Revision of subgenus Loboparius A. Schmidt, with description of a new species (Coleoptera Scarabaeidae). Annali del Museo civico di Storia Naturale Giacomo Doria, 84, 245–268.
  112. Dellacasa, M. & Dellacasa, G. (2006a) Aphodius reichei Harold, 1859: neotype designation and placement in the genus Gilletianus Balthasar, 1933. Kogane, 7, 27–30.
  113. Devecis, J. (2001) Plectrone delponti nov. sp. des Philippines. Observations sur le genre Plectrone Wallace 1867. Cetoniimania, 1 (4), 81–85.
  114. Devecis, J. (2004) Agestrata alexisi, nouvelle espèce des Philippines. Cetoniimania, 1 (1), 8–12.
  115. Devecis, J. (2009) Contribution à la connaissance des genres Plectrone Wallace 1867 et Paraplectrone Miksic 1985. Catalogue descriptif illustré et chorologie de leurs espèces. Cetoniimania, 6 (2), 52–88.
  116. Devecis, J. (2019) Contribution à l’étude du genre Agestrata Eschscholtz, 1829 (Coleoptera, Scarabaeoidea, Cetoniidae). Cetoniimania, New Series, 14, 24–80.
  117. Dohrn, C.A. (1872) Exotisches. Stettiner Entomologische Zeitung, 33, 143–165.
  118. Drumont, A., Ponchel, Y., Silvestre, G., Dechambre, R.P. & Jingke, L. (2006) Note sur la distribution geographique de Blabephorus pinguis Fairmaire, 1898 (Coleoptera, Dynastidae). Lambillionea, 106 (3), 443–446.
  119. Dunlap, J.B., Jameson, M.L., Engasser, E.L., Skelley, P.E. & Redford, A.J. (2016) Anomala sulcatula. In: Hawaiian Scarab ID. University of Nebraska–Lincoln, Lincoln, Nebraska. Available from: https://digitalcommons.unl.edu/hawaiianscarabid (accessed 16 October 2025)
  120. Dupuis, F. (2002) Un nouvel Oryctini des Philippines, Clyster rosaemariae (Coleoptera, Dynastidae). Coléoptères, 8 (10), 169–173.
  121. Endrődi, S. (1957) Zur Kenntnis der Dynastinen (Coleopt.). Revision des Dynastinen-Materials des Zoologischen Forschungsinstitutes und Museums Alexander Koenig, Bonn. Bonner Zoologische Beiträge, 1 (8), 64–70.
  122. Endrődi, S. (1959) Neue und bemerkenswerte Käferformen. Zur Kenntnis der Dynastien des Zoologischen Forschungsinstituts und Museums Alexander Koenig, Bonn. Bonner Zoologische Beiträge, 10, 116–125.
  123. Endrődi, S. (1969a) Monographie der Dynastinae (Coleoptera: Lamellicornia, Melolonthidae). 4. Tribus Pentodontini der orientalischen Region. Entomologische Arbeiten aus dem Museum G. Frey, 20, 403–479.
  124. Endrődi, S. (1969b) Neue orientalische und papuanische Pentodontini (Col. Melolonthidae). Folia Entomologica Hungarica, 22 (1), 5–19.
  125. Endrődi, S. (1971a) Monographie der Dynastinae 2 Tribus Oryctoderini (Coleoptera: Lamellicornia: Melolonthidae). Pacific Insects, 13 (2), 207–241.
  126. Endrődi, S. (1971b) Monographie der Dynastinae 4 Tribus Pentodontini (papuanische und pazifische Inselwelt) (Coleoptera: Lamellicornia: Melolonthidae). Pacific Insects, 13 (2), 243–320.
  127. Endrődi, S. (1973) Monographie der Dynastinae 5 Tribus Oryctini (Coleoptera, Lamellicornia, Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 24, 1–87.
  128. Endrődi, S. (1974) Monographie de Dynastinae 5 Tribus Oryctini 3 Teil Die Arten der orientalischen australischen und pazifischen Regionen. Acta Zoologica Academiae Scientiarum Hungaricae, 20 (3–4), 309–358.
  129. Endrődi, S. (1976) Monographie de Dynastinae (Coleoptera) 6 Tribus Dynastini I. Acta Zoologica Academiae Scientiarum Hungaricae, 22 (3–4), 217–269.
  130. Endrődi, S. (1977) Monographie de Dynastinae 6 (Coleoptera) Tribus Dynastini II. Acta Zoologica Academiae Scientiarum Hungaricae, 23 (1–2), 37–86.
  131. Endrődi, S. (1985) The Dynastinae of the World. Monographiae Biologicae. Vol. 28. Dr. W. Junk Publishers, Dordrecht, 800 pp.
  132. Erichson, W.F. (1834) Beiträge zur Zoologie, gesammelt auf einer Reise um die Erde, von Dr. F. J. F. Meyen. Sechste Abhandlung. Insekten: Coleoptera. Nova Acta Physico-Medica Academiae Caesarea Leopoldina-Carolinae, 16 (1), 219–276.
  133. Eschscholtz, J.F.G. von. (1822) Entomographien. Erste Lieferung. G. Reimer, Berlin, 128 pp.
  134. https://doi.org/10.1515/9783112513941
  135. Eydoux, J.F.T. (1839) Descriptions sommaires de quelques Coléoptères nouveaux, provenant de Manille et destinés à être publiés dans le Voyage autour du monde de la Corvette la Bonité. Revue Zoologique, par la Société Cuvierienne, 2, 264–267.
  136. Eydoux, J.F.T. & Souleyet, L.F.A. (1841–1842) Voyage autour du monde exécuté pendant les années 1836 et 1837 sur la corvette La Bonite, commandée par M. Vaillant. Zoologie. Tome Premier. Seconde Partie. Arthus Bertrand, Paris, 334 pp.
  137. Fabricius, J.C. (1775) Systema Entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Officina Libraria Kortii, Flensburgi et Lipsiae, [xxxii] + 832 pp.
  138. https://doi.org/10.5962/bhl.title.36510
  139. Fabrizi, S., Liu, W. G., Bai, M., Yang, X. & Ahrens, D. (2021) A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa, 4922 (1), 1–400. https://doi.org/10.11646/zootaxa.4922.1.1
  140. Fairmaire, L. (1883) Essai sur les Coléoptères de l’archipel de la Nouvelle-Bretagne. Annales de la Société Entomologique de Belgique, 27 (2), 1–58.
  141. Fairmaire, L. (1898) Descriptions de coléoptères d’Asie et de Malaisie. Annales de la Société Entomologique de France, 67, 382–400.
  142. https://doi.org/10.3406/bsef.1898.22155
  143. Frey, G. (1963) Neue Coprophagen und Melolonthiden mit Bestimmungstabellen (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 14 (2), 550–559.
  144. Frey, G. (1971) Neue afrikanische und asiatische Onthophagus-Arten (Col., Scarab., Coprinae). Entomologische Arbeiten aus dem Museum G. Frey, 22, 93–98.
  145. Frey, G. (1972a) Ein neuer Onthophagus von den Philippinen. Entomologische Arbeiten aus dem Museum G. Frey, 23, 268–269.
  146. Frey, G. (1972b) Katalog aller in die Gattung Sophrops Fairm. eingereihten Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 23, 354–356.
  147. Frey, G. (1976) Zwei neue Anomala-Arten aus den Philippinen (Col. Melolonthidae, Rutelinae). Entomologische Arbeiten aus dem Museum G. Frey, 27, 297–298.
  148. Gapud, V. (2005) The status of insect biodiversity in the Philippines. Asian Journal of Biodiversity, 2, 2–5.
  149. Gemminger, M. & Harold, E. (1869) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tomus IV. Scarabaeidae. Monachii, sumptu E. H. Gummi, Monachii, 368 pp. [pp. 979–1346]
  150. https://doi.org/10.5962/bhl.title.9089
  151. Gerstaecker, C.E.A. (1862) Phaedimus jagori, eine neue Goliathide von Luzon. Archiv für Naturgeschichte, 28 (1), 362–365.
  152. Geshev, G.A. (2024). A peculiar new genus and species of rhinoceros beetle, Ceratohyphus luzonicus, from the Philippines (Coleoptera: Scarabaeidae: Dynastinae). Faunitaxys, 12 (30), 1–10.
  153. Geshev, Q.Y.G. & Pathomwattananurak, W. (2023) A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa, 5361 (1), 114–124. https://doi.org/10.11646/zootaxa.5361.1.6
  154. Gillet, J.J.E. (1910) Espèces nouvelles du genre Copris et relevé synonymique des espèces decrites. Notes from the Leyden Museum, 32 (1), 1–31.
  155. Gillet, J.J.E. (1924) Scarabéides coprophages de Sumatra. Annales de la Société Entomologique de Belgique, 64, 102–104.
  156. Gory, H.L. & Percheron, A. (1833) Monographie des Cétoines et genres voisins, formant dans les familles naturelles de Latreille, la division des Scarabées Mélitophiles. J.B. Baillière, Paris, 410 pp.
  157. https://doi.org/10.5962/bhl.title.8957
  158. Guérin-Méneville, F.É. (1843) Animaux articulés, Coléoptères. In: Delessert, A. (Ed.), Souvenirs d’un voyage dans l’Inde exécuté de 1834 à 1839. Vol. 2. A. Delessert, Paris, pp. 33–50.
  159. Haller, G. (1884) Ueber Aberrationen, Varietäten und Arten einiger exotischer Cetonien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 7 (1), 30–34.
  160. https://doi.org/10.5962/bhl.part.4551
  161. Han, G.Y., Choi, J.B. & Park, J.K. (2021) A checklist of subfamily Scarabaeinae (Coleoptera: Scarabaeidae) from Cambodia. Journal of Asia-Pacific Biodiversity, 14 (1), 132–139.
  162. https://doi.org/10.1016/j.japb.2020.11.004
  163. Hao, M., Aidoo, O.F., Qian, Y., Wang, D., Ding, F., Ma, T., Tettey, E., Ninsin, K.D., Osabutey, A.F. & Borgemeister, C. (2022) Global potential distribution of Oryctes rhinoceros, as predicted by Boosted Regression Tree model. Global Ecology and Conservation, 37, e02175.
  164. https://doi.org/10.1016/j.gecco.2022.e02175
  165. Harold, E. von (1860). Diagnoses de nouvelles espèces d’Aphodiens. Annales de la Société Entomologique de France, 8 (3), 614–616.
  166. Harold, E. von. (1862a) Beiträge zur Kenntnis einiger coprophagen Lamellicornien (Viertes Stück). Berliner Entomologische Zeitschrift, 6, 379–403.
  167. https://doi.org/10.1002/mmnd.47918620310
  168. Harold, E. von. (1862b) Beiträge zur Kenntnis einiger coprophagen Lamellicornien (Drittes Stück). Berliner Entomologische Zeitschrift, 6, 138–171.
  169. https://doi.org/10.1002/mmnd.47918620115
  170. Harold, E. von. (1868) Abänderungen vergebener Namen. Coleopterologische Hefte, 4, 104.
  171. Harold, E. von. (1869) Abänderungen vergebener Namen. Coleopterologische Hefte, 5, 122–125.
  172. Harold, E. von. (1877) Enumeration des Lamellicorn coprophages rapportés de l’Archipel Malais, de la Nouvelle-Guinée et de l’Australie boréale par M.M.J. Doria, O. Beccari et L.M.D’Albertis. Annali del Museo Civico di Storia Naturale di Genova, 10, 38–110.
  173. Heller, K.M. (1893) Über einige Anomala-Arten aus Luzon. Entomologische Nachrichten, 19, 321–325.
  174. Heller, K.M. (1895) Zwei neue Cetoniden von den Philippinen. Deutsche Entomologische Zeitschrift, 1895, 281–285.
  175. https://doi.org/10.1002/mmnd.48018950203
  176. Heller, K.M. (1896) Neue Käfer von Celebes gesammelt von den Herren Dr. P. und Dr. F. Sarasin. Abhandlungen der Berichte des Königlichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden, 6 (3), 1–25.
  177. Heller, K.M. (1897a) Neue Käfer von Celebes II. Abhandlungen der Berichte des Königlichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden, 6 (10), 1–36.
  178. Heller, K.M. (1897b) Note XXI. Zwei neue Apogonien von den Philippinen. Notes from the Leyden Museum, 19, 189–193.
  179. Heller, K.M. (1897c) Über Camelonotus Fairm., Horonotus Burm., Neodipelicus Rits. und Palmerstonia Blackb. Notes from the Leyden Museum, 19, 163–173.
  180. Heller K.M. (1897d) Zwei neue Arten der Coleopteren-Gattung Callynomes Westw. Notes from the Leyden Museum, 19, 177–183.
  181. Heller, K.M. (1899a) Neue Käfer von den Philippinen. Abhandlungen der Berichte des Königlichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden, 7 (8), 1–8.
  182. Heller, K.M. (1899b) Neue und wenig gekannte Thaumastopaeus-Arten. Deutsche Entomologische Zeitschrift, 1899 (Heft II), 353–362.
  183. https://doi.org/10.1002/mmnd.48018990220
  184. Heller, K.M. (1921) Systematische und faunistische Notizen über Käfer. Entomologische Mitteilungen, 10 (6), 195–198.
  185. https://doi.org/10.5962/bhl.part.3326
  186. Heller K.M. (1934) New and little-known Philippine Coleoptera. The Philippine Journal of Science, 54 (2), 279–308.
  187. Herbst, J.F.W. (1785) Der Käfer. In: Jablonsky, C.G. (Ed.), Natursystem aller bekannten in- und ausländischen Insecten, als eine Fortsetzung der von Büffonschen Naturgeschichte. Nach dem System des Ritters von Linné und Fabricius. Vol. 1. Ben Joachim Pauli, Berlin, pp. 1–310.
  188. Hope, F.W. (1835) Monograph on Mimela, a genus of Coleopterous Insects. Transactions of the Entomological Society of London, 1, 108–117.
  189. https://doi.org/10.1111/j.1365-2311.1838.tb00150.x
  190. Hope, F.W. (1841) Descriptions of some nondescript lamellicorn beetles in his collection. The Annals and Magazine of Natural History, 8, 302–303.
  191. Hope, F.W. (1846) Descriptions of new Coleoptera. Transactions of the Entomological Society of London, 4, 146–147.
  192. https://doi.org/10.1111/j.1365-2311.1846.tb01354.x
  193. Hope, F.W. (1847) Descriptions of some new species of exotic Cetoniidae. Transactions of the Entomological Society of London, 5, 32–35.
  194. https://doi.org/10.1111/j.1365-2311.1847.tb01694.x
  195. Howden, H. (1995) New Rhyparini from Fiji and the Philippines (Coleoptera: Scarabaeidae: Aphodiinae). The Coleopterists Bulletin, 49 (1), 23–27.
  196. Huijbregts, J. & Krikken, J. (2009) A remarkable new two-horned species of Onthophagus from Borneo (Coleoptera: Scarabaeidae: Scarabaeinae). Malayan Nature Journal, 61, 1–9.
  197. Huijbregts, J. & Krikken, J. (2011) Taxonomy of Onthophagus javanensis and its closest relatives in Sundaland (Coleoptera: Scarabaeidae: Scarabaeinae). Tijdschrift voor Entomologie, 154 (1), 33–59.
  198. https://doi.org/10.1163/22119434-900000307
  199. Itoh, T. (1992) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae, Melolonthidae) I. Lamellicornia, 7, 7–14.
  200. Itoh, T. (1993a) A new genus and species of melolonthine beetle (Coleoptera, Melolonthidae) from Borneo. Elytra, 21 (1), 77–81.
  201. Itoh, T. (1993b) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae, Melolonthidae) II. Lamellicornia, 9, 3–20.
  202. Itoh, T. (1996) A Taxonomic Study on the genus Octoplasia Brenske (Coleoptera, Melolonthidae). Japanese Journal of Entomology, 64 (4), 772–778.
  203. Itoh, T. (2003a) Notes on Holotrichia constricta group, with description of a new species and redescription of two species (Scarabaeidae, Melolonthinae, Melolonthini). Kogane, 4, 7–15.
  204. Itoh, T. (2003b) Two new species and a new record of the Holotrichia constricta group (Scarabaeidae, Melolonthinae, Melolonthini). Elytra, 31 (2), 371–378.
  205. Jákl, S. (2008) Three new Oriental and Australian species of Thaumastopeus Kraatz, 1885 (Coleoptera: Scarabaeidae: Cetoniinae). Studies and Reports, Taxonomical Series, 4 (1–2), 111–117.
  206. Jákl, S. (2011) A contribution to knowledge of the Cetoniinae beetles from Philippines with descriptions of five new species (Coleoptera: Scarabaeidae: Cetoniinae). Studies and Reports, Taxonomical Series, 7 (1–2), 209–220.
  207. Jákl, S. (2014) Contribution to the fauna of Taenioderini with descriptions of five new species and two new subspecies (Coleoptera: Scarabaeidae: Cetoniinae). Studies and Reports, Taxonomical Series, 10 (2), 433 –452.
  208. Jákl, S. (2017a) A contribution to the knowledge of genus Clinterocera Motschulsky, 1858 with descriptions of new species (Coleoptera, Scarabaeoidea, Cetoniinae). Cetoniimania, New Series, 10, 3–25.
  209. Jákl, S. (2017b) Revision of Netociomima, Gonoprotaetia and Autoprotaetia, subgenera of the genus Protaetia (Coleoptera, Scarabaeidae, Cetoniinae). Cetoniimania, New Series, 11, 3–104.
  210. Jákl, S. (2020) A contribution to knowledge of the Oriental and Australian subgenus Mecinonota Kraatz, 1892 (Coleoptera: Scarabaeidae: Cetoniinae: Taenioderini). Studies and Reports, Taxonomical Series, 16 (2), 387–408.
  211. Jákl, S. (2021) Taxonomical notes about Miksicus Özdikmen & Turgut, 2009 and Miksicoprotaetia Legrand & Chew Kea Foo, 2010, subgenera of Protaetia Burmeister, 1842, with description of new species (Coleoptera, Scarabaeidae, Cetoniinae). Studies and Reports, Taxonomical Series, 17 (2), 297–313.
  212. Jákl, S. (2022) Description of two new species of Protaetia Burmeister, 1842 from the Philippines from rarely collected subgenera Lawangia Schenkling, 1921 and Protaetiola Mikšič, 1963 (Coleoptera: Scarabaeidae: Cetoniinae). Studies and Reports, Taxonomical Series, 18 (1), 95–103.
  213. Jákl, S. (2023) A contribution to knowledge of Chalcoprotaetia Mikšič, 1963 and Hemiprotaetia Mikšič, 1963 (Coleoptera: Scarabaeidae: Cetoniinae). Studies and Reports, Taxonomical Series, 19 (2), 309–324.
  214. Jákl, S. (2024) Review of the genus Protaetiomorpha Miksic, 1968 (Coleoptera: Scarabaeidae: Cetoniinae). Studies and Reports, Taxonomical Series, 20 (2), 365–374.
  215. Jákl, S. & Legrand, J. P. (2018) New species of Ixorida J. Thomson from south Vietnam (Coleoptera: Scarabaeidae: Cetoniinae). Studies and Reports, Taxonomical Series, 14 (1), 81–88.
  216. Jákl, S. & Zídek, J. (2017) A contribution to the knowledge of Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae). Folia Heyrovskyana, Series A, 25 (2), 19–30.
  217. Janson, O.E. (1881) Notices of new or little known Cetoniidae Nr. 7. Cistula Entomologica, 2 (24), 603–611.
  218. Janson, O.E. (1888) Description of a new Cetoniid. Notes from the Leyden Museum, 10, 207–208.
  219. Jordana y Morera, R. (1885) Bosquejo geográfico e histórico-natural del Archipiélago Filipino. Imprenta de Moreno y Rojas, Madrid, 461 pp.
  220. https://doi.org/10.5962/bhl.title.58482
  221. Kaneko, N. & Wada, K. (2020) Catalogue of the type material of Scarabaeoidea (Coleoptera) deposited in the Research Institute of Evolutionary Biology, Tokyo, Japan. ZooKeys, 958, 35–89. https://doi.org/10.3897/zookeys.958.52799
  222. Keith, D. (2006) Reflexions sur le genre Engertia et description d’une nouvelle espèce des Philippines (Col. Scarabaeoidea Melolonthidae). Lambillionea, 106 (1), 90–94.
  223. Keith, D., Pham, P.V. & Sabatinelli, G. (2023) Exolontha permirabilis, a new species from Vietnam with notes on the genus (Col. Scarabaeidae Melolonthinae Melolonthini). Faunitaxys, 11 (73), 1–5.
  224. Kim, J.I. (2011) Arthropoda: Insecta: Coleoptera: Scarabaeoidea. Pleurosticti. National Institute of Biological Resources, Incheon, 252 pp.
  225. Kirby, W. (1818) A century of insects, including several new genera described from his cabinet. Transactions of the Linnean Society, 12 (2), 375–453.
  226. https://doi.org/10.1111/j.1095-8339.1817.tb00239.x
  227. Kobayashi, H. (2013) Ten new species of the genus Apogonia (Coleoptera, Scarabaeidae, Melolonthinae) from the Philippines. Kogane, 14, 57–73.
  228. Kobayashi, H. (2014) Notes on the genus Apogonia (Coleoptera, Scarabaeidae, Melolonthinae) from the Philippines. Kogane, 16, 50–60.
  229. Kobayashi, H. & Fujioka, M. (2014) Some new species of the genus Hoplia (Coleoptera, Scarabaeidae, Melolonthinae, Hopliini) from the Philippines. Kogane, 16, 25–31.
  230. Kobayashi, H. & Medina, M.N. (2024) Three new species related to Apogonia rizali Heller, 1897 (Coleoptera, Scarabaeidae, Melolonthinae) from Mindanao Is., the Philippines. Kogane, 27, 22–33.
  231. Kobayashi, H. & Sasaki, N. (2016) On the genus Callynomes (Coleoptera, Scarabaeidae, Cetoniinae) from the Philippines and Sulawesi Is. Kogane, 18, 49–66.
  232. Kraatz, G. (1880) Synonymische Bemerkungen über exotische Cetoniden. Entomologische Monatsblätter, 11, 31–32.
  233. https://doi.org/10.1002/mmnd.4800240305
  234. Kraatz, G. (1890) Ueber die Varietäten der Eudicella morgani Westw. und Coelorrhina hornimani Westw. (aus West-Africa). Deutsche Entomologische Zeitschrift, 1890, 215–218.
  235. https://doi.org/10.1002/mmnd.48018900334
  236. Kraatz, G. (1892) Monographische Revision der Ruteliden-Gattung Popillia Serville. Deutsche Entomologische Zeitschrift, 1892 (2), 225–306.
  237. https://doi.org/10.1002/mmnd.48018920107
  238. Kraatz, G. (1893) Zwei neue Arten der Goliathiden-Gattung Phaedimus. Deutsche Entomologische Zeitschrift, 1893 (1), 75–77.
  239. https://doi.org/10.1002/mmnd.48018930111
  240. Kraatz, G. (1894a) Phaedimus mohnikei Kraatz var. minor? Deutsche Entomologische Zeitschrift, 1894 (2), 107–108.
  241. https://doi.org/10.1002/mmnd.48018950122
  242. Kraatz, G. (1894b) Protaetia mindoroensis Kraatz n. sp. Deutsche Entomologische Zeitschrift, 1894 (2), 298.
  243. https://doi.org/10.1002/mmnd.48018940538
  244. Kraatz, G. (1895) Poecilophana nov. gen. Cetonidarum errichtet auf Cetonia ochroplagiata Heller. Deutsche Entomologische Zeitschrift, 34 (2), 285–287.
  245. https://doi.org/10.1002/mmnd.48018950204
  246. Kraatz, G. (1899) Glycyphana bipustulata (anstatt binotata). Deutsche Entomologische Zeitschrift, 1899 (2), 399.
  247. https://doi.org/10.1002/mmnd.48018990232
  248. Krajčik, M. (1998) Cetoniidae of the World. Catalogue—Part 1. M. Krajčik, Most, Czech Republic, 96 pp.
  249. Krajčik, M. (1999) Cetoniidae of the World. Catalogue—Part 2. M. Krajčik, Most, Czech Republic, 72 pp.
  250. Krajčik, M. (2006) Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma.x Supplement, 3, 1–189.
  251. Král, D., Rakovič, M. & Mencl, L. (2014) Two new Gilletianus species (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini) from Sulawesi, Indonesia. Studies and Reports, Taxonomical Series, 10 (1), 113–126.
  252. Král, D. & Šípek, P. (2013) Aphodiinae (Coleoptera: Scarabaeidae) of the Goa, Maharashtra and Rajasthan (India) with description of Aphodius (Gilletianus) rajawatorum sp. nov. Acta Entomologica Musei Nationalis Pragae, 53 (2), 633–648.
  253. Krikken, J. (1977a) Some new and otherwise noteworthy species of Onthophagus Latreille from the Indo-Australian archipelago. Zoologische Mededelingen, 52 (13), 169–184.
  254. Krikken, J. (1977b) The Asian genus Pleuronota Kraatz and allied forms a clarification (Coleoptera: Cetoniidae). Zoologische Mededelingen, 51 (13), 199–209.
  255. Krikken, J. (1978) A new Plectrone from the Philippine Islands (Coleoptera: Cetoniidae). Entomologische Berichten, 38, 30–32.
  256. Krikken, J. (1979) Taxonomic review of the Southeast Asian genus Rhinacosmus Kraatz (Coleoptera: Cetoniidae). Zoologische Mededelingen, 54 (20), 281–289.
  257. Krikken, J. (1980) Taxonomic reviews of three small Cremastochiliform genera from Asia and Africa (Coleoptera: Cetoniidae). Zoologische Mededelingen, 56 (2), 7–25.
  258. Krikken, J. (1986) Two new species of Onthophagus Latreille from the Philippine Island of Samar (Coleoptera: Scarabaeidae). Zoologische Mededelingen, 60 (19), 277–283.
  259. Krikken, J. (2009) Drepanocerine dung beetles: a group overview, with description of new taxa (Coleoptera: Scarabaeidae: Scarabaeinae). Haroldius, 4, 3–30.
  260. Krikken, J. (2018) Two new genera of Australian flower chafers, with an annotated overview of their relatives (Coleoptera: Scarabaeidae: Cetoniinae). Haroldius, 5, 3–34.
  261. Krikken, J. & Huijbregts, J. (2007) Taxonomic diversity of the genus Ochicanthon in Sundaland (Coleoptera: Scarabaeidae: Scarabaeinae). Tijdschrift voor Entomologie, 150 (2), 421.
  262. https://doi.org/10.1163/22119434-900000241
  263. Krikken, J. & Huijbregts, J. (2009) Onthophagus pilularius and its close relatives in Sundaland: a taxonomic reappraisal (Coleoptera, Scarabaeidae, Scarabaeinae). Deutsche Entomologische Zeitschrift, 56 (1), 41–55.
  264. https://doi.org/10.1002/mmnd.200900004
  265. Krikken, J. & Huijbregts, J. (2011) New Southeast Asian Onthophagus species in the deflexicollis group (Coleoptera: Scarabaeidae: Scarabaeinae). Tijdschrift voor Entomologie, 154 (2), 3–213.
  266. https://doi.org/10.1163/22119434-900000320
  267. Kühbandner, M. (1984) Beschreibung der bisher unbekannten Weibchens von Protaetia kuehbandneri Mikšić, 1983 (Insel Romblon, Philippinen) (Coleoptera, Scarabaeidae, Cetoniinae). Entomofauna, 5 (16), 209–212.
  268. Kuijten, P.J. (1975) Beitrag zur Kenntnis der Dynastinae (Coleoptera: Melolonthidae). Folia Entomologica Hungarica, Series Nova, 28 (1), 105–108.
  269. Kuijten, P.J. (1988) Rutelarcha, Lutera, and Cyphelytra: notes on taxonomy and nomenclature, diagnoses, and keys (Coleoptera: Rutelinae, Rutelini). Zoologische Mededelingen, 62 (6), 75–90.
  270. Kuijten, P.J. (1992) A Revision of the genus Parastasia in the Indo-Australian Region (Coleoptera: Scarabaeidae: Rutelinae). Zoologische Verhandelingen, 275, 1–207.
  271. Kuijten, P.J. (1994) Notes on the Dynastid genus Neohyphus Heller, 1896 (Coleoptera, Lamellicornia). Deutsche Entomologische Zeitschrift, 41 (1), 57–61.
  272. https://doi.org/10.1002/mmnd.19940410103
  273. Lansberge, G.W. van. (1875) Monographie des Onitides. Annales de la Société Entomologique de Belgique, 18, 1–148.
  274. Laporte, F.L.N. de C. (Comte de Castelnau) (1840). Histoire naturelle des insectes coléoptères, avec une introduction renfermant l’anatomie et la physiologie des animaux articulés, par M. Brullé. Vol. 2. P. Duménil, Paris, 563 pp.
  275. Legrand, J.P. (2004) Cetoniidae du Sabah (Malaisie orientale) et des Philippines, nouveaux genre, nouvelles espèces (Coleoptera, Cetoniidae). Les Cahiers Magellanes, Hors Série, 9, 1–10.
  276. Legrand, J.P. (2017) Notes sur le genre Ixorida J. Thomson 1880, et son sous-genre Mecinonota Kraatz 1892, aux Philippines et description d’espèces nouvelles (Coleoptera, Scarabaeoidea, Cetoniidae, Cetoniinae, Taenioderini, Taenioderina). Cetoniimania, New Series, 10, 53–60.
  277. Legrand, J.P. (2018a) Précisions sur le genre Glycyphana Burmeister 1842, sous-genre Caloglycyphana Miksic 1968, aux Philippines et description d’une nouvelle sous-espèce et d’une nouvelle espèce (Coleoptera, Scarabaeoidea, Cetoniinae). Cetoniimania, New Series, 13, 36–42.
  278. Legrand, J.P. (2018b) Une nouvelle espèce du genre Protaetia Burmeister 1842, sous-genre Miksicoprotaetia Legrand et Chew Kea Foo 2010, des Philippines (Coleoptera, Scarabaeoidea, Cetoniinae). Cetoniimania, New Series, 13, 3–6.
  279. Legrand, J.P. (2020) Note sur l’identité de Cetonia confusa Gory et Percheron 1833 et d’Euglypta intermedia Miksic 1979 et replacement de ces espèces dans le genre Glycosia (Schoch) (Coleoptera, Scarabaeoidea, Cetoniidae). Cetoniimania, New Series, 15, 48–52.
  280. Legrand, J.P. (2022). Deux nouvelles espèces du genre Clinteria (Burmeister) du Vietnam et des Philippines (Palawan) (Coleoptera, Scarabaeoidea, Cetoniidae, Gymnetini). Cetoniimania, New Series, 16, 7–11.
  281. Legrand, J.P. (2025). Description d’une nouvelle espèce de Pleuronota Kraatz, 1892 de l’île de Palawan, Philippines (Coleoptera, Scarabaeoidea, Cetoniidae, Taeniodernini). Cetoniimania, New Series, 19, 1–3.
  282. Legrand, J.P. & Chew Kea Foo, S. (2004) Cetoniidae du Sabah (Malaisie orientale) et des Philippines nouveau genre, nouveau sous-genre, nouvelles espèces, nouvelle sous-espèce (Coleoptera Cetoniidae). Les Cahiers Magellanes, Hors Série, 10, 1–11.
  283. Legrand, J.P. & Flutsch, G. (2016) Une nouvelle espèce de Reineria Miksic 1968, sous-genre Euglyptica Arnaud, 2004, de Mindanao, Philippines (Coleoptera, Scarabaeoidea, Cetoniidae, Cetoniinae). Cetoniimania, New Series, 9, 79–81.
  284. Legrand, J.P. & Flutsch, G. (2017) Illustration du mâle de Reineria (Euglyptica) thieuleuxi Legrand et Flutsch, 2016 et reclassement de l’espèce dans le sous-genre Reineria Miksic (Coleoptera, Scarabaeoidea, Cetoniidae, Cetoniinae). Cetoniimania, New Series, 12, 27–28.
  285. Litsinger, J.A., Apostol, R.F. & Obusan, M.B. (1983) White grub, Leucopholis irrorata (Coleoptera: Scarabaeidae): pest status, population dynamics, and chemical control in a rice-maize cropping pattern in the Philippines. Journal of Economic Entomology, 76 (5), 1133–1138.
  286. https://doi.org/10.1093/jee/76.5.1133
  287. Löbl, I. & Smetana, A. (Eds.) (2006) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea–Scirtoidea– Dascilloidea–Buprestoidea–Byrrhoidea. Apollo Books, Stenstrup, 690 pp.
  288. Masumoto, K. (1980) Description of a new species of Rhyssemus (Scarabaeidae) from the Philippines. Elytra, 8 (1), 21–22.
  289. Masumoto, K., Ochi, T. & Ho, B.H. (2019) A new Rhyparus species from Taiwan (Scarabaeidae, Aphodiinae, Rhyparini). Kogane, 22, 13–18.
  290. Machatschke, J.W. (1972) Scarabaeoidea: Melolonthidae Rutelinae. In: Wilcox, J.A. (Ed.), Coleopterorum Catalogus. Supplementa. Pars 66. Fascicle I. Editio Secunda. W. Junk, The Hague, pp. 1–361.
  291. Matsumoto, T. (2005) A new discovery of the genus Latipalpus (Scarabaeidae, Melolonthinae, Melolonthini) from the Philippines. Elytra, 33 (1), 219–224.
  292. Matsumoto, T. (2008a) A taxonomic note on a melolonthine species (Coleoptera, Scarabaeidae, Melolonthinae, Melolonthini) from Palua. Kogane, 9, 11–14.
  293. Matsumoto, T. (2008b) Notes on the Holotrichia constricta group, with description of a new species (Coleoptera, Scarabaeidae, Melolonthinae, Melolonthini). Kogane, 9, 15–19.
  294. Matsumoto, T. (2008c) Three new species of the Holotrichia constricta group (Scarabaeidae, Melolonthinae, Melolonthini) from the Philippines. Elytra, 36 (1), 101–108.
  295. Matsumoto, T. (2010a) Four new species of the Holotrichia constricta group (Scarabaeidae, Melolonthinae, Melolonthini) from the Philippines. Elytra, 38 (1), 43–52.
  296. Matsumoto, T. (2010b) Taxonomic study on the genus Psilopholis Brenske (Scarabaeidae, Melolonthinae, Melolonthini) with considerations of divisions of the subtribe Melolonthina. Elytra, 38 (2), 239–247.
  297. Matsumoto, T. (2015) Separation of Amphitrichia from Holotrichia (Scarabaeidae, Melolonthinae, Melolonthini) with description of a new species and new records of two known species. Kogane, 17, 11–17.
  298. Matsumoto, T. (2022) Notes on Amphitrichia stylifer (Chapin) and its related species (Coleoptera, Scarabaeidae, Melolonthinae) from the Philippines. Kogane, 25, 29–34.
  299. Mencl, L., Rakovič, M. & Král, D. (2013) A new species of the genus Rhyparus (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) from the Oriental Region having an accessory costa on each elytron. Studies and Reports, Taxonomical Series, 9 (2), 487–498.
  300. Mikšić, R. (1962) Erster Beitrag zur Kenntnis der Protaetia-Arten (38. Beitrag zur Kenntnis der Scarabaeiden). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden, 28 (2), 5–35.
  301. Mikšić, R. (1963a) Zweiter Beitrag zur Kenntnis der Protaetia-Arten die Protaetien der Philippinischen Inseln (39. Beitrag zur Kenntnis der Scarabaeiden). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden, 29 (4), 333–452.
  302. Mikšić, R. (1963b) Vierter Beitrag zur Kenntnis der Protaetia-Arten. (41. Beitrag zur Kenntnis der Scarabaeiden). Deutsche Entomologische Zeitschrift, 10 (3–5), 415–436.
  303. https://doi.org/10.1002/mmnd.4800100317
  304. Mikšić, R. (1967) Beschreibung neuer und Bemerkungen über bekannte Cetoniiden aus der orientalischen und australischen Region (54. Beitrag zur Kenntnis der Scarabaeiden). Bollettino dell’Associazione Romana di Entomologia, 22 (9), 21–31.
  305. Mikšić, R. (1968) Eine neue Art und zwei neue Rassen der Gattung Glycyphana Burm. von den Philippinen (58. Beitrag zur Kenntnis der Scarabaeiden). Bulletin Scientifique Jugoslavie, Sectio A, 13 (9–10), 306–307.
  306. Mikšić, R. (1970a) Contribution à la connaissance des Cetoniinae des regions paléarctique et orientale (62e Contribution à la connaissance des Scarabaeidae). Bulletin Mensuel de la Société Linnéenne de Lyon, 39 (2), 54–59.
  307. https://doi.org/10.3406/linly.1970.6059
  308. Mikšić, R. (1970b) Revision der Gattung Glycyphana Burmeister (Coleoptera, Cetoniinae) I. Teil (56. Beitrag zur Kenntnis der Scarabaeiden). Zoologische Verhandelingen, 107, 1–112.
  309. Mikšić R. (1970c) Katalog der Gattung Glycyphana Burm. (Col.) 60. Beitrag zur Kenntnis der Scarabaeiden. Folia Entomologica Hungarica, Series Nova, 23 (6), 141–147.
  310. Mikšić, R. (1970d) Revision der Gattung Ixorida J. Thomson (71. Beitrag zur Kenntnis der Scarabaeiden). Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen, 22 (4), 119–125.
  311. Mikšić R. (1971a) Einige Bemerkungen uber die systematische Stellung und Einteilung der asiatischen Goliathina (78. Beitrag zur Kenntnis der Scarabaeiden). Acta Entomologica Jugoslavica, 7 (2), 37–47.
  312. Mikšić, R. (1971b) Revision der Gattung Glycyphana Burmeister (Coleoptera, Cetoniinae) II. Teil (64. Beitrag zur Kenntnis der Scarabaeiden). Zoologische Verhandelingen, 118, 3–153.
  313. Mikšić R. (1971c) Revision der Gattung Meroloba Thomson (Col., Scarab.) (75. Beitrag zur Kenntnis der Scarabaeiden). Folia Entomologica Hungarica, Series Nova, 24 (36), 389–402.
  314. Mikšić R. (1972a) Revision der Gattung Mecinonota Kraatz (Col. Scarab.) (69. Beitrag zur Kenntnis der Scarabaeiden). Mitteilungen der Münchener Entomologischen Gesellschaft, 62, 87–128.
  315. Mikšić, R. (1972b) Ein Beitrag zur Kenntnis der Gattung Phaedimus Waterh. (Coleoptera-Scarabaeidae) (76. Beitrag zur Kenntnis der Scarabaeiden). Bulletin Scientifique Jugoslavie, Sectio A, 17 (3–4), 73–74.
  316. Mikšić, R. (1972c) Macronotiola nov. gen.- eine neue Gattung der Gymnetini (74. Beitrag zur Kenntnis der Scarabaeiden). Acta Entomologica Jugoslavica, 8 (1–2), 123–134.
  317. Mikšić, R. (1973) Drei neue Arten der Gattung Plectrone Wall. (Coleoptera-Scarabaeiden) (80. Beitrag zur Kenntnis der Scarabaeiden). Bulletin Scientifique Jugoslavie Sect. A, 18 (4–6), 78–79.
  318. Mikšić, R. (1974a) Die orientalischen und palaearktischen Gattungen der Heterorrhinini (83. Beitrag zur Kenntnis der Scarabaeiden). Revue Suisse de Zoologie, 81 (4), 737–783.
  319. https://doi.org/10.5962/bhl.part.76040
  320. Mikšić, R. (1974b) Eine neue Art der Gattung Plectrone von den Philippinen (87. Beitrag zur Kenntnis der Scarabaeiden). Acta Entomologica Jugoslavica, 10 (1–2), 161–164.
  321. Mikšić, R. (1974c) Neue Unterarten der Cetoniinae (Coleoptera-Scarabaeidae) (86. Beitrag zur Kenntnis der Scarabaeiden). Bulletin Scientifique Jugoslavie, Sectio A, 19 (9–10), 236–237.
  322. Mikšić, R. (1974d) Revision der Gattung Euselates Thomson (78. Beitrag zur Kenntnis der Scarabaeiden). Mitteilungen aus dem zoologischen Museum in Berlin, 50 (1), 55–129.
  323. Mikšić, R. (1976) Monographie der Cetoniinae der paläarktischen und orientalischen Region. Coleoptera: Lamellicornia. Band I. Allgemeiner Teil. Systematischer Teil. Gymnetini (Taenioderina, Chalcotheina). Forstinstitut, Sarajevo, 444 pp.
  324. Mikšić, R. (1977) Monographie der Cetoniinae der paläarktischen und orientalischen Region. Coleoptera: Lamellicornia, Band 2: Systematischer Teil: Gymnetini (Lomapterina, Clinteriina), Phaedimini, Gnathocerini, Heterorrhinini. Forstinstitut, Sarajevo, 399 pp.
  325. Mikšić, R. (1979) Drei neue Arten der Gattung Euglypta Mohn. (Coleoptera: Scarabaeidae, Cetoniinae). Acta Entomologica Jugoslavica, 15 (1–2), 27–34.
  326. Mikšić, R. (1981) Revision der Gattung Astraea Mohn. (Coleoptera, Scarabaeidae, Cetoniinae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden, 44 (9), 203–233.
  327. Mikšić, R. (1982a) Monographie der Cetoniinae der paläarktischen und orientalischen Region Coleoptera: Lamellicornia. Band 3. Systematischer Teil. Cetoniini I.Teil. Forstinstitut, Sarajevo, 529 pp.
  328. Mikšić, R. (1982b) Neue Beiträge zur Kenntnis der paläarktischen und orientalischen Cetoniinae (Coleoptera, Scarabaeidae). Reichenbachia, Zeitschrift für Taxonomische Entomologie des Staatlichen Museums für Tierkunde in Dresden, 20 (12), 99–105.
  329. Mikšić, R. (1983) Beschreibungen neuer und Bemerkungen über bekannte Cetoniinae aus dem tropischen Asien (Coleoptera, Scarabaeidae). Entomofauna, 4 (24), 381–397.
  330. Mikšić, R. (1984) Weitere Beiträge zur Kenntnis der Cetoniinae-Fauna der Orientalischen Region (Coleoptera, Scarabaeidae). Entomofauna, 5 (26), 299–315.
  331. Mikšić, R. (1985) Ein Beitrag zur Kenntnis der Chalcotheina (Coleoptera, Cetoniinae, Gymnetini). Articulata, 2 (6), 141–144.
  332. Mikšić, R. (1986) Neue Beiträge zur Kenntnis der Cetoniinae der Philippinen und von Malaysien (Coleoptera, Scarabaeidae). Entomofauna, 7 (5), 65–77.
  333. Mikšić, R. (1987) Monographie der Cetoniinae der paläarktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 4. Systematischer Teil. Cetoniini II. Teil. Forstinstitut, Sarajevo, 607 pp.
  334. Minkina, Ł. & Kakizoe, S. (2020) Termitodiellus mindanaoensis, a new species of Rhyparini Schmidt, 1910 (Scarabaeidae: Aphodiinae) from the Philippines. Insecta Mundi, 762, 1–6.
  335. Minkina, Ł., Anichtchenko, A., Vasiljeva, A. & Skelley, P.E. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81–101.
  336. Minkina, Ł., Jákl, S. & Bezděk, A. (2023) A new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species with modified pygidium and last abdominal ventrite in females. Faunitaxys, 11 (66), 1–5.
  337. Miyake, Y. & Yamaya, S. (1997) New or little known Scarabaeid beetles from Southeast Asia, preserved in Nagaoka Municipal Science Museum. Bulletin of the Nagaoka Municipal Science Museum, 32, 5–20.
  338. Miyake, Y. & Yamaya, S. (1999) Descriptions of some new dynastid beetles from Southeast Asia, preserved in Nagaoka Municipal Science Museum. Bulletin of the Nagaoka Municipal Science Museum, 34, 101–112.
  339. Miyake, Y. & Yamaya, S. (2000) Descriptions of some new dynastid beetles from Southeast Asia, preserved in Nagaoka Municipal Science Museum (II). Bulletin of the Nagaoka Municipal Science Museum, 35, 7–12.
  340. Mohamedsaid, M.S., Zaidi, M.I. & Salleh, I. (1997) The Scarabaeidae (Coleoptera) from Banggi Island, Sabah. Serangga, 2 (2), 213–215.
  341. Mohnike, O. (1873) Die Cetoniden der Philippinischen Inseln. Archiv für Naturgeschichte, 39 (1), 109–247.
  342. https://doi.org/10.5962/bhl.part.12371
  343. Mohnike, O. (1874) Neue und seltene Käfer aus den Familien Cetoniden und Dynastiden, Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 31, 261–262.
  344. Moore, M.R., Cave, R.D. & Branham, M.A. (2018) Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys, 745, 101–378. https://doi.org/10.3897/zookeys.745.23685
  345. Moser, J. (1906) Beschreibung neuer Cetoniden Arten. Annales de la Société Entomologique de Belgique, 50, 273–280.
  346. Moser, J. (1907) Beitrag zur Kenntnis der Cetoniden II. Annales de la Société Entomologique de Belgique, 51, 141–151.
  347. Moser, J. (1909) Beitrag zur Kenntnis der Cetoniden VI. Annales de la Société Entomologique de Belgique, 53 (7), 313–323.
  348. Moser, J. (1910) Neue Coleoptera Lamellicornia von den Philippinen. The Philippine Journal of Science, 5, 183–189.
  349. Moser, J. (1911a) Beitrag zur Coleopteren-Fauna der Philippinen. The Philippine Journal of Science, 6, 331–333.
  350. Moser, J. (1911b) Beitrag zur Kenntnis der Cetoniden IX. Annales de la Société entomologique de Belgique, 55 (4), 119–129.
  351. Moser, J. (1912) Neue Arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Société Entomologique de Belgique, 56, 420–449.
  352. Moser, J. (1913) Beitrag zur Kenntnis der Melolonthiden II. Annales de la Société Entomologique de Belgique, 57, 331–345.
  353. https://doi.org/10.5962/bhl.part.4592
  354. Moser, J. (1914) Beitrag zur Kenntnis der Cetoniden (Col.) XIV. Deutsche Entomologische Zeitschrift, 1914 (6), 573–614.
  355. https://doi.org/10.1002/mmnd.48019140603
  356. Moser, J. (1915a) Beitrag zur Kenntnis der Melolonthiden. (Col.) IV. Deutsche Entomologische Zeitschrift, 1915, 113–151.
  357. https://doi.org/10.1002/mmnd.48019150204
  358. Moser, J. (1915b) Neue Melolonthiden und Cetoniden (Col.). Deutsche Entomologische Zeitschrift, 1915, 579–605.
  359. https://doi.org/10.1002/mmnd.48019150603
  360. Moser, J. (1915c) Weitere neue Serica-Arten (Col.). Stettiner Entomologische Zeitung, 76, 144–202.
  361. https://doi.org/10.1002/mmnd.191519150402
  362. Moser, J. (1916a) Beitrag zur Kenntnis der Melolonthiden. (Col.) V. Deutsche Entomologische Zeitschrift, 1916, 129–188.
  363. Moser, J. (1916b) Beitrag zur Kenntnis der Cetoniden (Col.) XV. Stettiner Entomologische Zeitung, 77, 139–157.
  364. Moser, J. (1916c) Eine neue Phaedimus-Art von Luzon (Col.). Deutsche Entomologische Zeitschrift, 1916–1917, 385–386.
  365. Moser, J. (1917) Beitrag zur Kenntnis der Cetoniden (Col.) XVI. Deutsche Entomologische Zeitschrift, 1917 (1/2), 1–22.
  366. Moser, J. (1917b) Beitrag zur Kenntnis der Melolonthiden (Col.) VI. Stettiner Entomologische Zeitung, 78, 54–97.
  367. Moser, J. (1918) Beitrag zur Kenntnis der Cetoniden (CoAl.) XVII. Stettiner Entomologische Zeitung, 79, 168–190.
  368. Moser, J. (1921a) Beitrag zur Kenntnis der Melolonthiden (Col.) XII. Stettiner Entomologische Zeitung, 82, 48–73.
  369. Moser, J. (1921b) Neue Cetoniden Arten (Col.). Stettiner Entomologische Zeitung, 82, 183–187.
  370. https://doi.org/10.1002/mmnd.48019210312
  371. Moser, J. (1922a) Drei neue Cetoniden-Arten von den Philippinen (Col.). Stettiner Entomologische Zeitung, 83, 113–114.
  372. Moser, J. (1922b) Neue Serica-Arten aus den orientalischen Region (Col.). Stettiner Entomologische Zeitung, 83, 101–112.
  373. Moser, J. (1924a) Beitrag zur Kenntnis der Melolonthiden (Col.) XIV. Stettiner Entomologische Zeitung, 84, 137–164.
  374. Moser, J. (1924b) New Species of Melolonthid Beetles from the Philippine Islands. The Philippine Journal of Science, 25, 349–357.
  375. Moser, J. (1925) New Philippine Cetonidae. The Philippine Journal of Science, 27, 69.
  376. Moser, J. (1926) Beitrag zur Kenntnis der Melolonthiden (Col.) XIV. Stettiner Entomologische Zeitung, 87, 198–208.
  377. Motschulsky, V. de (1853) Diagnoses de Coléoptères nouveaux, trouvés par MM. Tatarinoff et Gaschkéwitsch aux environs de Pékin. Études Entomologiques, 2, 44–51.
  378. Muramoto, R. (1995) Two new species of the genus Eophileurus Arrow (Coleoptera, Scarabaeidae, Dynastinae) from Southeast Asia. Japanese Journal of Systematic Entomology, 1 (2), 223–226.
  379. Nagai, S. (1980) A new species of the genus Theodosia J. Thomson from Palawan Island, Philippines (Coleoptera: Scarabaeidae-Cetoniinae). Transactions of the Shikoku Entomological Society, 15 (1–2), 49–51.
  380. Nagai, S. (1984) On some new cetoniid beetles from the Philippines (Coleoptera: Scarabaeidae). Transactions of the Shikoku Entomological Society, 16 (3), 1–9.
  381. Nagai, S. (1985) Studies on the subfamily Cetoniinae of Asia (4) (Coleoptera, Scarabaeidae). Gekkan-Mushi, 174, 2–7.
  382. Nagai, S. (1986) Studies on the subfamily Cetoniinae (Coleoptera, Scarabaeidae) of Asia, V. In: Ueno, S.I. (Ed.), Entomological Papers Presented to Yoshihiko Kurosawa on the Occasion of His Retirement. Coleopterists’ Association of Japan, Tokyo, pp. 184–187.
  383. Nagai, S. (1998) Studies on the Subfamily Cetoniinae of Asia (7) (Coleoptera, Scarabaeidae). Gekkan-Mushi, 324, 4–9.
  384. Nakamura, H. (1974) List of the subfamily Dynastinae (Scarabaeidae s. lat.) from Malaysia, Indonesia and Philippines. Miscellaneous Publications of the Entomological Research Institute, Yamanashi, 5, 1–6.
  385. Newman, E. (1838) New Species of Popillia Leach. Magazine of Natural History and Journal of Zoology, Botany, Mineralogy, Geology and Meteorology, New Series, 2, 336–338 + 392.
  386. Newman, E. (1841) Entomological notes. The Entomologist, 1, 169–171.
  387. https://doi.org/10.5962/bhl.part.9070
  388. Nomura, S. (1973) Notes on the Japanese Scarabaeidae III. The Entomological Review of Japan, 25 (1–2), 13–20.
  389. Ochi, T. & Kon, M. (1994) Dung beetles (Coleoptera, Scarabaeoidea) collected from Sabah, Borneo (I). Elytra, 22 (2), 281–298.
  390. Ochi, T., Kon, M. & Barclay, M.V.L. (2008) A review of the subgenus Parascatonomus of the genus Onthophagus (Coleoptera: Scarabaeidae) of Borneo. Taichius, Special publication of the Japan Coleopterological Society, 2, 191–234.
  391. Ochi, T. (1990) Two new coprophagous beetles (Coleoptera, Scarabaeidae) from the Philippines. Elytra, 18 (2), 215–219.
  392. Ochi, T. & Araya, K. (1992) Studies on the Coprophagous Scarab Beetles from East Asia II (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 6, 79–108.
  393. Ochi, T. & Araya, K. (1996) Studies on the coprophagous scarab beetles from East Asia III (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 8, 1–15.
  394. Ochi, T. & Kon, M. (1995) Dung beetles (Coleoptera, Scarabaeoidea) collected from Sabah, Borneo (II). Elytra, 23 (1), 43–60.
  395. Ochi, T. & Kon, M. (1997) Studies on the Coprophagous Scarab beetles from East Asia. VI Descriptions of new taxa of the genus Paragymnopleurus (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 8, 235–238.
  396. Ochi, T. & Kon, M. (1999) Notes on the coprophagous Scarab beetles (Coleoptera, Scarabaeidae) from Southeast Asia (II). A new species of the genus Onthophagus from Philippines. The Entomological Review of Japan, 54 (2), 137–141.
  397. Ochi, T. & Kon, M. (2001) Notes on the Coprophagous Scarab-Beetles (Coleoptera, Scarabaeidae) from South-East Asia (III) A new species of the genus Onthophagus from the Philippines. Special Publication of the Japanese Coleopterological Society, 1, 371–374.
  398. Ochi, T. & Kon, M. (2004) Notes on the coprophagous scarab-beetles (Coleoptera, Scarabaeidae) from Southeast Asia (V). Two new species of the genus Onthophagus from Philippines. Kogane, 5, 31–36.
  399. Ochi, T. & Kon, M. (2006) Notes on the coprophagous scarab beetles (Coleoptera, Scarabaeidae) from Southeast Asia (XI). Four new species and a new subspecies of Onthophagus from Luzon, the Philippines. Kogane, 7, 41–48.
  400. Ochi, T. & Kon, M. (2007) Notes on the coprophagous scarab beetles (Coleoptera, Scarabaeidae) from Southeast Asia (XV). Three new species of the genus Onthophagus from Palawan and Borneo. Kogane, 8, 89–95.
  401. Ochi, T. & Kon, M. (2025a) A new species of Phaedimus (Coleoptera, Scarabeidae, Cetoniinae) from the Philippines. Kogane, 28, 59–62.
  402. Ochi, T. & Kon, M. (2025b) A new species of the genus Setylaides (Coleoptera, Scarabaeidae, Aphodiinae, Eupariini) from Palawan, Philippines. Kogane, 28, 32–35.
  403. Ochi, T. & Kon, M. (2025c) Five new species of the genus Onthophagus with notes on several known Onthophagus and Parascatanomus species (Coleoptera, Scarabaeidae) from the Philippines. Kogane, 28, 44–58.
  404. Ochi, T. & Sakai, K. (2021) A revisional study of the genus Phaedimus Waterhouse, 1841 from the Philippines (Coleoptera, Scarabaeidae, Cetoniinae). Giornale Italiano di Entomologia, 16 (66), 25–48.
  405. Ochi, T., Kakizoe, S., Minkina, Ł., Kawahara, M. & Cabras, A.A. (2021) A revisional study of the genus Rhyparus from the Philippines (Coleoptera, Scarabaeidae, Aphodiinae, Rhyparini). Kogane, 24, 17–42.
  406. Ochi, T., Kawahara, M. & Cabras, A.A. (2021) Five new species of the genera Onthophagus and Parascatonomus from the Philippines (Coleoptera, Scarabaeidae). Kogane, 24, 57–68.
  407. Ochi, T., Kon, M. & Barclay, M.V.L. (2016) A Review of the Subgenus Pseudophanaeomorphus Ochi of the genus Onthophagus (Coleoptera: Scarabaeidae). Giornale Italiano di Entomologia, 14 (61), 415–448.
  408. Ochi, T., Kon, M. & Cabras, A.A. (2023a) A revisional study of the subgenus Indachorius of the genus Onthophagus from the Philippines (Coleoptera, Scarabaeidae, Scarabaeinae) with descriptions of six new species. Kogane, 26, 37–51.
  409. Ochi, T., Kon, M. & Cabras, A.A. (2023b) Six new Onthophagus species from the Philippines (Coleoptera, Scarabaeidae, Scarabaeinae). Kogane, 26, 52–63.
  410. Ochi, T., Kon, M. & Cabras, A.A. (2023c) Three new species of the subgenus Matashia Matsumura, 1938 of the genus Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae, Onthophagini). Giornale Italiano di Entomologia, 16 (68), 647–658.
  411. Ochi, T., Kon, M. & Cabras, A.A. (2024) Description of five new taxa of Onthophagus and Parascatonomus (Coleoptera, Scarabaeidae, Scarabaeinae) from the Philippines, with a redescription and a new distributional record. Kogane, 27, 47–60.
  412. Ochi, T., Kon, M. & Cabras, A.A. (2025a) Notes on the coprophagous scarab-beetles (Coleoptera, Scarabaeidae) from Southeast Asia. XXXVII. Descriptions of a new species of Panelus and a new species of Cyobius from the Philippines. Giornale Italiano di Entomologia, 17 (70), 285–290.
  413. Ochi, T., Kon, M. & Cabras, A.A. (2025b) Two new species of the subgenus Onthophagiellus of the genus Onthophagus (Coleoptera: Scarabaeidae) from the Philippines. Kogane, 28, 36–43.
  414. Ochi, T., Kon, M. & Kakizoe, S. (2016) Nine New Taxa of the Genus Gilletianus from Southeast Asia (Coleoptera: Scarabaeidae). Japanese Journal of Systematic Entomology, 22 (2), 307–320.
  415. Ochi, T., Kon, M. & Kawahara, M. (2010) Four new species of the genus Gilletianus from Southeast Asia (Coleoptera, Scarabaeidae). Kogane, 11, 111–120.
  416. Ohaus, F. (1898) Phaenomeridae. Stettiner Entomologische Zeitung, 59 (1–2), 3–41.
  417. Ohaus, F. (1900) Revision der Parastasiiden. Deutsche Entomologische Zeitschrift, 2, 225–266.
  418. https://doi.org/10.1002/mmnd.48019000210
  419. Ohaus, F. (1901) Ruteliden der alten Welt. Deutsche Entomologische Zeitschrift, 1, 125–134.
  420. Ohaus, F. (1905) Beiträge zur Kenntnis der Ruteliden. Deutsche Entomologische Zeitschrift, 1, 81–99.
  421. https://doi.org/10.1002/mmnd.48119050214
  422. Ohaus, F. (1908) Beiträge zur Kenntnis der Ruteliden (Col.). Deutsche Entomologische Zeitschrift, 5 (5), 634–644.
  423. https://doi.org/10.1002/mmnd.48019080517
  424. Ohaus, F. (1910) Die Ruteliden der philippinischen Inseln. The Philippine Journal of Science, 5 (4), 233–262.
  425. Ohaus, F. (1912a) Nachtraege und Berichtigungen zu: “Die Ruteliden der Philippinischen Inseln”. The Philippine Journal of Science, 7 (4), 255–269.
  426. Ohaus, F. (1912b) Über einige mit Kulturpflanzen verschleppte Adoretiden. Entomologische Blätter, 8–9, 218–227.
  427. Ohaus, F. (1913) Anomala (Euchlora) micholitzi n. sp. (Col. lamell. Rutelin.). Entomologische Rundschau, 30, 69–70.
  428. Ohaus, F. (1914a) Revision der Adoretini (Col. lamell. Rutelin.). Deutsche Entomologische Zeitschrift, 58, 471–514.
  429. https://doi.org/10.1002/mmnd.48019140502
  430. Ohaus, F. (1914b) Zweiter Nachtrag zur Kenntnis der Ruteliden der Philippin. Inseln. Stettiner Entomologische Zeitung, 75, 185–193.
  431. Ohaus, F. (1915a) XVII Beitrag zur Kenntnis der Ruteliden (Col. lamell.). Stettiner Entomologische Zeitung, 76, 88–143.
  432. https://doi.org/10.1002/mmnd.191519150308
  433. Ohaus, F. (1915b) Dritter Nachtrag zur Kenntnis der Philippiner Ruteliden (Col. lamell.). Stettiner Entomologische Zeitung, 76, 339–344.
  434. https://doi.org/10.1002/mmnd.191519150110
  435. Ohaus, F. (1916) IV. Nachtrag zur Kenntnis der Philippin. Ruteliden (Col. lamell.). Stettiner Entomologische Zeitung, 77, 353–359.
  436. https://doi.org/10.1002/mmnd.48019160306
  437. Ohaus, F. (1923) V. Nachtrag zur Kenntnis der philippinischen Ruteliden (Coleoptera, Lamellicornia). The Philippine Journal of Science, 22 (16), 581–590.
  438. Ohaus, F. (1924a) Eine neue philippinische Euchlora-Art (Coleoptera-Rutelidae). The Philippine Journal of Science, 24 (1), 103.
  439. Ohaus, F. (1924b) Einige neue Anomala-Arten der Philippinen. The Philippines Journal of Science, 24 (1), 53–55.
  440. Ohaus, F. (1927) Zwei neue Anomala-Arten der Philippinen (Coleoptera, Lamellicornia, Ruteliden). The Philippine Journal of Science, 32, 61–62.
  441. Ohaus, F. (1930a) VI. Nachtrag zur Kenntnis der philippinischen Ruteliden (Coleoptera, Lamellicornia). The Philippine Journal of Science, 43 (4), 555–558.
  442. Ohaus, F. (1930b) XXVI. Beitrag zur Kenntnis der Ruteliden (Col. lamell.) Deutsche Entomologische Zeitschrift, 2–3, 138–158.
  443. https://doi.org/10.1002/mmnd.48019300205
  444. Ohaus, F. (1934) Coleoptera: Lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. Erster Teil. In: Wytsman, P. (Ed.), Genera Insectorum. Coleoptera, Fascicule 199A. Louis Desmet-Verteneuil, Bruxelles, pp. 1–172.
  445. Ohaus, F. (1943) Revision der Gattung Mimela Kirby (Col. Scarab. Rutelin.). Deutsche Entomologische Zeitschrift, 75, 65–88.
  446. Özdikmen, H. & Turgut, S. (2009) Miksicus nom. nov., a replacement name for the preoccupied beetle genus Urbania Miksic, 1963 (Coleoptera: Scarabaeoidea: Cetoniidae). Entomological News, 120, 227–229.
  447. https://doi.org/10.3157/021.120.0217
  448. Paulian, R. (1944) Sur quelques coléoptères coprophages récoltés par le Dr. E. Mjöberg à Bornéo et en Australie. Revue Française d’Entomologie, 10, 64–75.
  449. Paulian, R. (1981) Sur quelques Coléoptères Scarabaeoidea indo-malais nouveaux ou peu connus. Bulletin de la Société Entomologique de France, 86 (3–4), 110–112.
  450. https://doi.org/10.3406/bsef.1981.17952
  451. Pavićević, D. (1986) Eine neue Protaetia-Art von den Philippinen (Coleoptera, Scarabaeidae, Cetoniinae). Entomofauna, 7 (8), 117–120.
  452. Pavićević, D. (1988). Beitrag zur Kenntnis der philippinischen Cetoniinae (Coleoptera, Scarabaeidae). Entomofauna, 9 (6), 169–179.
  453. Pereira, F.S. (1954) A new myrmecophilous scarabaeid beetle from the Philippine Islands with a review of Haroldius. Psyche, 61 (1), 1–8.
  454. https://doi.org/10.1155/1954/72174
  455. Perissinotto, R. & Clennell, L. (2021) Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys, 1026, 17–43. https://doi.org/10.3897/zookeys.1026.60036
  456. Petrovitz, R. (1961a) Neue und verkannte Aphodiinae aus allen Erdteilen (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 12 (1), 99–135.
  457. Petrovitz, R. (1961b) Neue und verkannte Aphodiinae aus allen erdteilen (Col. Scarab.) II. Teil. Entomologische Arbeiten aus dem Museum G. Frey, 12, 344–356.
  458. Petrovitz, R. (1962) Neue und verkannte Aphodiinae aus allen Erdteilen (Col. Scarab.) III. Teil. Entomologische Arbeiten aus dem Museum G. Frey, 13, 101–131.
  459. Petrovitz, R. (1972) Neue laparostikte Scarabaeidae aus der orientalischen and neotropischen Region (Coleoptera). Memorie della Società Entomologica Italiana, 51, 161–168.
  460. Pic, M. (1905) Note sur deux Lamellicornes exotiques. Le Naturaliste, 27, 131.
  461. https://doi.org/10.5962/bhl.part.26096
  462. Pittino, R. (1984) Taxonomic considerations on, types revisions, lectotypes designations and descriptions of new or little known Psammodiini from Palaearctic, Oriental and Ethiopian regions (Coleoptera Aphodiidae) (XXVI contribution to the knowledge of Coleoptera Scarabaeoidea). Giornale Italiano di Entomologia, 2 (6), 13–98.
  463. Pittino, R. & Mariani, G. (1986) A revision of the Old World species of the genus Diastictus Muls. and its allies (Platytomus Muls., Pleurophorus Muls., Afrodiastictus n. gen., Bordatius n. gen.) (Coleoptera, Aphodiidae, Psammodiinae). Giornale Italiano di Entomologia, 3 (12), 1–165.
  464. Pittino, R., Rakovič, M. & Mencl, L. (2013) A contribution to knowledge of Rhyssemus Mulsant, 1892 species from the Indochinese subregion with description of a new species from Laos and proposal of two new synonyms (Coleoptera: Aphodiidae: Psammodiinae: Rhyssemini). Studies and Reports, Taxonomical Series, 9 (1), 193–200.
  465. Pokorny, S. & Zídek, J. (2014) Review of the Gymnopleurini (Coleoptera: Scarabaeidae: Scarabaeinae). II. The genus Paragymnopleurus Shipp. Insecta Mundi, 0395, 1–10.
  466. Pope, R.D. (1960) Aserica, Autoserica, Neoserica, or Maladera? (Coleoptera: Melolonthidae). The Annals and Magazine of Natural History, Series 13, 3 (33), 545–550.
  467. https://doi.org/10.1080/00222936008651055
  468. Prell, H. (1912) Beiträge zur Kenntnis der Dynastien (V). Entomologische Blätter, 8, 179–187.
  469. Prell, H. (1934) Beiträge zur Kenntnis der Dynastien (XII–XIII). Beschreibungen und Bemerkungen. Entomologische Zeitschrift, Frankfurt, 47, 162–164, 186–188, 194–195.
  470. Prokofiev, A.M. (2019) Two new species of Engertia Dalla Torre, 1913 and Philacelota Heller, 1900 from Indonesia and the Philippines with a revised identification key (Coleoptera: Scarabaeidae: Melolonthinae). European Journal of Taxonomy, 561, 1–13.
  471. https://doi.org/10.5852/ejt.2019.561
  472. Rakovič, M. (1987) A revision of the genus Odochilus Harold with remarks on the tribal classification of the subfamily Aphodiinae (Coleoptera, Scarabaeidae). Acta Entomologica Bohemoslovaca, 84, 27–44.
  473. Rakovič, M. (1997) A review of the genus Odochilus (Coleoptera: Aphodiidae: Odochilinae). Acta Societatis Zoologicae Bohemicae, 61 (2), 157–167.
  474. Rakovič, M. & Anichtchenko, A. (2021) Two new species of the genus Odochilus Harold, 1877 (Coleoptera: Scarabaeidae: Aphodiinae: Odochilini). Zootaxa, 4942 (3), 447–450.
  475. https://doi.org/10.11646/zootaxa.4942.3.8
  476. Rakovič, M., Mencl, L. & Král, D. (2023) Neotrichiorhyssemus ghati sp. nov. (Coleoptera: Scarabaeidae: Aphodiinae: Psammodiini) from southwest India. Studies and Reports, Taxonomical Series, 19 (1), 125–137.
  477. Redtenbacher, L. (1867 [1868]) Reise der Österreichischen Fregatte “Novara” um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Zweiter Band 1. Abtheilung: Coleopteren. Carl Gerold’s Sohn, Wien, 249 pp.
  478. Reichenbach, A. (1995) Protaetia anneliae nov. spec. von den Philippinen (Insecta: Coleoptera: Lamellicornia: Cetoniinae). Reichenbachia: Zeitschrift für Taxonomische Entomologie des Staatlichen Museums für Tierkunde in Dresden, 31 (1), 45–48.
  479. Ricchiardi, E. (1996) Notes on Philippines Dasyvalgus (Coleoptera, Cetoniidae, Valginae, Valgini). Bollettino del Museo Regionale di Scienze Naturali di Torino, 14 (2), 365–377.
  480. Ritsema, C. (1897) Note VII. Six new species of the Melolonthid genus Apogonia. Notes from the Leyden Museum, 19, 117–124.
  481. Ritsema, C. (1904) Note VII. Second supplementary list of the described species of the Melolonthid genus Apogonia. Notes from the Leyden Museum, 25, 111–116.
  482. Ritsema, C. (1912) Third supplementary list of the described Apogonia-species, with an alternation in nomenclature. Notes from the Leyden Museum, 34 (2), 128.
  483. Rowland, J.M. (2003) Male horn dimorphism, phylogeny and systematics of rhinoceros beetles of the genus Xylotrupes (Scarabaeidae: Coleoptera). Australian Journal of Zoology, 51, 213–258.
  484. https://doi.org/10.1071/ZO02013
  485. Rowland, J.M. (2006a) Partial revision of the systematics of rhinoceros beetles of the genus Xylotrupes (Scarabaeidae, Coleoptera). Privately published, Albuquerque, 41 pp.
  486. Rowland, J.M. (2006b) Continued revision of the systematics of rhinoceros beetles of the genus Xylotrupes (Scarabaeidae, Coleoptera). Privately published, Albuquerque, 49 pp.
  487. Rowland, J.M. (2011) Notes on nomenclature in Xylotrupes Hope (Scarabaeidae: Dynastinae: Dynastini). Insecta Mundi, 0176, 1–10.
  488. Sakai, K. (1995) Notes on the Genus Glycosia Schoch (Coleoptera, Scarabaeidae, Cetoniinae) from Southeast Asia. Gekkan-Mushi, 295, 3–11.
  489. Sakai, K. (1996) Notes on the Genus Taeniodera from the Philippines (Coleoptera, Scarabaeidae, Cetoniinae). The Nature and Insects, 31 (9), 25–28.
  490. Sakai, K. (1997) Two New species of the genus Callynomes Mohnike (Coleoptera, Scarabaeidae, Cetoniinae) from the Philippines. Gekkan-Mushi, 321, 10–15.
  491. Sakai, K. & Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha’s Iconographic Series of Insects. Vol. 3. Shizawa Printing Co. Ltd., Tokyo, 421 pp.
  492. Sasaki, N. (2019) A new species of the genus Callynomes (Coleoptera, Scarabaeidae, Cetoniinae) from the Philippines with a key to the species of Callynomes. Kogane, 22, 67–70.
  493. Saylor, L.W. (1937) Necessary changes in status of important rhizotrogid genera (Col. Scarabaeidae). Revista de Entomologia, 7, 318–322.
  494. Schaufuss, L.W. (1885) Beitrag zur Fauna der niederländischen Besitzungen auf den Sunda-Inseln. Horae Societatis Entomologicae Rossicae, 19, 183–209.
  495. Schaum, H.R. (1844) Observations critiques sur la famille des Lamellicornes mélitophiles. Annales de la Société Entomologique de France, 2 (2), 333–426.
  496. Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Coleopterorum Catalogus. Vol. XXI. Pars 72. W. Junk, Berlin, 431 pp.
  497. Schmidt, A. (1909) Neue Aphodiinen und einige synonymische Bemerkungen. Notes from the Leyden Museum, 31, 101–124.
  498. Schmidt, A. (1913) Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv für Naturgeschichte, Abt. A, 79A (11), 117–178.
  499. Schmidt, A. (1922) Scarabaeidae: Coprinae, Aphodiinae, Hybosorinae. In: Die von Dr. K. Friederichs in Samoa und Indochina gesammelten Käfer. Archiv für Naturgeschichte, 88 (10), 147–159.
  500. Schoch, G. (1895) Die Genera und Species meiner Cetoniden-Sammlung. II. Teil. Tribus Cetoniadae, Diplognathidae und Cremastochilidae. E. Zwingli, Zürich, 148 pp.
  501. Schoch, G. (1896) Catalogus systematicus Cetonidarum et Trichiidarum adhuc cognitarum. W. Jun, Zürich, 95 pp.
  502. Schoch, G. (1897) Nachtrag IV zu Schoch: Die Genera und Species meiner Cetonidensammlung. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 9 (10), 451–477.
  503. https://doi.org/10.5962/bhl.part.2749
  504. Schönherr, C.J. (1817) Appendix ad C.J. Schönherr Synonymiam Insectorum. Tom. I. Part. 3. Sistens descriptiones novarum specierum. Lewerentz, Scaris, 266 pp.
  505. Schultze, W. (1915) I. Beitrag zur Coleopteren Fauna der Philippinen. The Philippine Journal of Science, Series D, 10, 271–279.
  506. Schultze, W. (1916a) A catalogue of Philippine Coleoptera. The Philippine Journal of Science, 11, 1–194 (154–180).
  507. Schultze, W. (1916b) II. Beitrag zur Coleopteren Fauna der Philippinen. The Philippine Journal of Science, 11 (4), 291–298.
  508. Schultze, W. (1916c) III. Beitrag zur Coleopteren Fauna der Philippinen. The Philippine Journal of Science, 11, 347–350.
  509. Schultze, W. (1920) Eight contribution to the Coleoptera fauna of the Philippines. The Philippine Journal of Science, 16 (2), 191–201.
  510. Schürhoff, P.N. (1933a) Beiträge zur Kenntnis der Cetoniden (Col.). Mitteilungen der Deutschen Entomologischen Gesellschaft, 4 (1), 26–32.
  511. Schürhoff, P.N. (1933b) Beiträge zur Kenntnis der Cetoniden III. Entomologisches Nachrichtenblatt, 7 (3), 89–96.
  512. Schürhoff, P.N. (1933c) Weitere Beiträge zur Kenntnis der Cetoniden (Col.). Mitteilungen der Deutschen Entomologischen Gesellschaft, 4 (7), 97–102.
  513. Schürhoff, P.N. (1935) Beiträge zur Kenntnis der Cetoniden (Col.) VII. Mitteilungen der Deutschen Entomologischen Gesellschaft, 6 (3–4), 21–28.
  514. Schürhoff, P.N. (1938) Beiträge zur Kenntnis der Cetoniden (Col.) X. Revision der Gattung Pachnoda Burm. Annales de la Société Entomologique de Belgique, 78, 187–194.
  515. Schürhoff, P.N. (1942) Beiträge zur Kenntnis der Cetoniden (Col.) IX. Mitteilungen der Münchner Entomologischen Gesellschaft, 32 (1), 279–293.
  516. Sharp, D. (1875) Descriptions of some new genera and species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 13, 33–54.
  517. Sharp, D. (1876) Descriptions of some new species of Scarabaeidae from tropical Asia and Malasia. Coleopterologische Hefte, 15, 65–90.
  518. Silvestre, G. (2002) Une nouvelle espèce de Xylotrupes des Philippines (Coleoptera, Dynastidae). Coléoptères, 8 (17), 247–251.
  519. Silvestre, G. (2006) Les Xylotrupes des Philippines et des îles voisines (Coleoptera, Dynastidae). Coléoptères, 12 (27), 361–375.
  520. Skelley, P.E. (2007) Generic limits of the Rhyparini with respect to the genus Termitodius Wasmann, 1894 (Coleoptera: Scarabaeidae: Aphodiinae). Insecta Mundi, 9, 1–9.
  521. Skelley, P.E. (2015) Saprovisca sarangay, new species, a second species in an unusual Philippine genus of Eupariini (Coleoptera: Scarabaeidae: Aphodiinae). Insecta Mundi, 0419, 1–6.
  522. Stebnicka, Z.T. (1986) Revision of the Aphodiinae of the Nepal-Himalayas (Coleoptera: Scarabaeidae). Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 397, 1–51.
  523. Stebnicka, Z.T. (1989) Revision of the Aphodiinae of the Western Himalayas (Coleoptera: Scarabaeidae). Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 441, 1–29.
  524. Stebnicka, Z.T. (1991) Aphodiinae from Sabah (Coleoptera: Scarabaeidae). Revue Suisse de Zoologie, 98 (1), 3–7.
  525. https://doi.org/10.5962/bhl.part.79773
  526. Stebnicka Z.T. (1992) Aphodiinae from Thailand (Coleoptera: Scarabaeidae). Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 481, 1–16.
  527. Stebnicka, Z.T. (1993) A new genus and species of Aphodiinae from Leyte, Philippines with notes on other taxa (Coleoptera: Scarabaeoidea). The Philippine Entomologist, 9 (1), 28–35.
  528. Stebnicka, Z.T. & Howden, H.F. (1997) Revision of the Australian species of Ataenius Harold (Coleoptera: Scarabaeoidea: Aphodiinae: Eupariini). Invertebrate Taxonomy, 11 (5), 735–821.
  529. https://doi.org/10.1071/IT95046
  530. Stebnicka, Z.T. & Lago, P.K. (2005) The new world species of Ataenius Harold, 1867. V. Revision of the A. strigatus group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi, 19 (1–2), 55–83.
  531. https://doi.org/10.3409/000000006783995364
  532. Stebnicka, Z.T. (2012) A revision of the Indonesian species of Saprosites Redtenbacher, 1858 (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, 55 (2), 13–45.
  533. https://doi.org/10.3409/azc.55_2.13
  534. Théry, T. & Bordat, P. (2012) Aphodiidae de Nouvelle-Calédonie: mise à jour des connaissances et descriptions de nouvelles espèces (Coleoptera, Scarabaeoidea). Bulletin de la Société Entomologique de France, 117 (3), 309–338.
  535. https://doi.org/10.3406/bsef.2012.29307
  536. Thomson, J. (1878) Typi Cetonidarum, suivis de typi Molommidarum et de Typi Nilionidarum Musaei Thomsoniani. Émile Deyrolle, Paris, 44 pp.
  537. Ueda, A., Dwibadra, D., Kahono, S., Sugiarto, Ochi, T. & Kon, M. (2022) Atlas of dung beetles collected in the Sungai Wain Protection Forest and its surroundings in the lowlands of Borneo. Bulletin of Forestry and Forest Products Research Institute, Kyushu Research Center, 21 (2), 165–192.
  538. Voirin, J.P. (1996) Six nouvelles espèces de Dynastinae (Col. Scarabaeoidea). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 69 (2), 215–228.
  539. Wada, K. (1989) Two New Species of the Genus Parastasia (Rutelini, Parastasiina) from Philippines. Lamellicornia, 5, 1–7.
  540. Wada, K. (1997) Two new species of the Genus Parastasia (Coleoptera, Scarabaeidae, Rutelinae) from the Philippines. Elytra, 25 (2), 419–423.
  541. Wada, K. (1998a) Three new species of the Genus Callistethus (Coleoptera, Scarabaeidae, Rutelinae) from Palawan and Borneo. Elytra, 26 (1), 145–153.
  542. Wada, K. (1998b) Two new species of the Genus Lutera (Coleoptera, Scarabaeidae, Rutelinae) from Borneo and Sumatra. Elytra, 26 (2), 359–367.
  543. Wada, K. (2002) Three new species of the genus Callistethus (Coleoptera, Scarabaeidae, Rutelinae) from Sulawesi and Mindanao. Elytra, 30 (1), 173–181.
  544. Wada, K. (2003) Notes on the genus Parastasia (Scarabaeidae, Rutelinae) from the Oriental region. IV Re-evaluation of the status of P. isidai Wada, 1989, with redescriptions of P. helleri Ohaus, 1898 and P. isidai. Kogane, 4, 1–5.
  545. Wada, K. (2007) Three new species of the genus Parastasia (Coleoptera, Scarabaeidae, Rutelinae) from Southeast Asia. Elytra, 35 (2), 497–505.
  546. Wada, K. (2008) Five new species of the genus Parastasia (Coleoptera, Scarabaeidae, Rutelinae) from the Oriental region. Elytra, 36 (1), 87–99.
  547. Wada, K. (2015) Systematic study of the genus Parastasia Westwood, 1842 (Coleoptera: Scarabaeidae). PhD thesis, Kyushu University, Fukuoka, 567 pp.
  548. Wada, K. (2016) A new species of the genus Parastasia (Coleoptera, Scarabaeidae) from Luzon Island, the Philippines. Elytra, New Series, 6 (1), 123–125.
  549. Wallace, A.R. (1867) A catalogue of the Cetoniinae of the Malayan Archipelago, with descriptions of the new species, brief diagnose of novelties. Proceedings of the Entomological Society of London, 3 (3), 92–97.
  550. Wallace, A.R. (1868) A catalogue of the Cetoniinae of the Malayan Archipelago, with descriptions of the new species. Transactions of the Entomological Society of London, 4 (3), 519–601.
  551. Waterhouse, C.O. (1884) New Coleoptera in the British Museum. The Annals and Magazine of Natural History, Series 5, 13 (77), 370–372.
  552. https://doi.org/10.1080/00222938409459256
  553. Waterhouse, G.R. (1841a) Descriptions of some new Coleopterous insects from the Philippine Islands, collected by H. Cuming, Esq. Journal of Proceedings of the Entomological Society of London, 1, 36–38.
  554. Waterhouse, G.R. (1841b) Euchirus quadrilineatus and Xylotrupes pubescens from the Philippine Islands. The Annals and Magazine of Natural History, 7, 538–539.
  555. Westwood, J.O. (1841) On the Australian genus Cryptodus, and upon Parastasia, the Asiatic representative of the Rutelidae. The Annals and Magazine of Natural History, 8, 303–304.
  556. Westwood, J.O. (1842) Insectorum novorum Centuria. Decadis secundae, ex ordine Coleopterorum Synopsis. The Annals and Magazine of Natural History, Series 1, 8 (50), 203–205. https://doi.org/10.1080/03745484109442753
  557. Westwood, J.O. (1874) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope, M. A., D.C.L., F.R.S., &c. With forty plates from drawings by the author. Clarendon Press, Oxford, 205 pp.
  558. https://doi.org/10.5962/bhl.title.14077
  559. White, A. (1856) Description of some coleopterous insects in the collection of the British Museum apparently unnoticed. Proceedings of the Zoological Society of London, 24, 8–17 + 406–410.
  560. Woodruff, R.E. (2006) The Asian mango flower beetle, Protaetia fusca (Herbst), and Euphoria sepulcralis (Fabricius) in Florida and the West Indies (Coleoptera: Scarabaeidae: Cetoniinae). Insecta Mundi, 20 (3–4), 227–232.
  561. Xu, H. & Zhao, M.Z. (2024) Synonymic notes in the Southeast Asian genus Lutera Westwood, 1875 (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa, 5523 (5), 531–542.
  562. https://doi.org/10.11646/zootaxa.5523.5.3
  563. Yamaya, S. (2006) A new species of the genus Papuana from Mindanao Is., the Philippines (Coleoptera, Dynastinae, Pentodontini). Kogane, 7, 49–51.
  564. Yamaya, S. (2009) Two new species of Dynastinae from the Philippines (Coleoptera, Scarabaeidae). Bulletin of the Nagaoka Municipal Science Museum, 44, 21–26.
  565. Yamaya, S. (2011) On the genus Neohyphus (Dynastinae, Oryctoderini) of the Philippines, with description of a new species. Bulletin of the Nagaoka Municipal Science Museum, 46, 3–8.
  566. Yang, Q.Z., Pathomwattananurak, W. & Zhao, M.Z. (2023) Two new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from the Philippines and Vietnam, with first description of E. quadratifovealis Yang & Pathomwattananurak, 2022 female. Zootaxa, 5249 (2), 290–300. https://doi.org/10.11646/zootaxa.5249.2.6
  567. Zídek, J. (2020) Catalogue of species-group names assigned to Copris Geoffroy, Coptodactyla Burmeister, Litocopris Waterhouse, Microcopris Balthasar, Paracorpis Balthasar, Pseudocopris Ferreira, Pseudopedaria Felsche, Sinocorpris Ochi, Kon and Bai, and Thyregis Blackburn (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini). Insect Mundi, 0756, 1–24.
  568. Zídek, J. (2021) Revision of Philacelota Heller, 1900, and Engertia Dalla Torre, 1912, with description of a new genus (Coleoptera: Scarabaeidae: Melolonthinae). In: Telnov, D., Barclay, M.V.L. & Pauwels, O.S.G. (Eds.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea. Vol. IV. Entomological Society of Latvia, Riga, pp. 397–411.
  569. Zorn, C. (2006) Taxonomic Revision of the Anomala cuprascens-Species Group of Sulawesi and the Papuan Region: The Species with Unidentate Protibiae (A. chlorotica-Subgroup) (Coleoptera: Scarabaeidae: Rutelinae). Arthropod Systematics & Phylogeny, 65 (1), 25–71.
  570. https://doi.org/10.3897/asp.65.e31665
  571. Zorn, C. (2011) Taxonomic revision of the Anomala cuprascens species group of Sulawesi and the Papuan Region. The species with a bidentate protibia (Coleoptera: Scarabaeidae: Rutelinae). In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea. Vol. 1. Entomological Society of Latvia, Riga, pp. 351–428.

How to Cite

Magallanes, J.G., Nuñeza, O.M. & Mondejar, E.P. (2026) First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa, 5746 (1), 1–148. https://doi.org/10.11646/zootaxa.5746.1.1