Abstract
A species list of phytoseiid mites (Acari: Mesostigmata: Phytoseiidae) in Japan is presented. The list was compiled from a survey of published literature, re-examination of accessible permanent slides, and from records obtained during field trips. A total of 97 taxa of phytoseiid mites, known to the species level, are so far reported as native to Japan. In addition, eight species are regarded as exotic, having been introduced into Japan. These phytoseiid mites are represented by the three subfamilies and 22 genera. The subfamily with the most known species is the Amblyseiinae Muma, with 64 species, among which 17 are in the genus Amblyseius Berlese, 15 in Neoseiulus Hughes, seven in Typhlodromips De Leon, four in Scapulaseius Karg & Oomen-Kalsbeek, three in each of Euseius Wainstein, Paraphytoseius Swirski & Schechter, and Proprioseiopsis Muma, two in each of Amblyseiulella Muma, Gynaeseius Wainstein, Okiseius Ehara, and Transeius Chant & McMurtry, and one in each of the genera, Amblydromalus (Garman & McGregor), Iphiseius Berlese, Phytoscutus Muma and Phytoseiulus Athias-Henriot. In the subfamily Phytoseiinae Berlese, there are 13 species of Phytoseius Ribaga, while the subfamily Typhlodrominae Wainstein, is represented by 28 species, of these, 23 in Typhlodromus and one species in each of the genera, Chanteius Wainstein, Galendromus Muma, Kuzinellus Wainstein, Paraseiulus Muma and Typhlodromina Muma. Identification keys to subfamilies, tribes, genera and species of the Phytoseiidae in Japan are presented.
References
- Abbasova, E.D. (1972) Phytoseiid mites (Parasitiformes: Phytoseiidae) of Azerbaijan. Avtoreferat Dissertatsii na Soiskanie Uchenoy Stepeni Kandidata Biologicheskikh Nauk. Akadrmiya Nauk Azerbaydzhanskoy SSR, Institut Zoologii, Baku, Azerbaijan, 34 pp. [in Russian]
- Amano, H. (1994) Collection record of predacious phytoseiid mites in Chiba Prefecture (Acari: Phytoseiidae). Faculty of Horticulture of Chiba University Technical Bulletin, 48, 237–240.
- Amano, H., Kawashima, M., Matsumura, M., Saito, M. & Toyoshima, S. (2011) Description of an unrecorded phytoseiid mite, Neoseiulus harrowi (Collyer), from Japanese Spinach Greenhouses. Journal of the Acarological Society of Japan, 20, 95–102. https://doi.org/10.2300/acari.20.95
- Amitai, S. & Swirski, E. (1981) A new species of Amblyseius (Acarina: Phytoseiidae) from the Far East. Israel Journal of Entomology, 15, 59–66.
- Athias-Henriot, C. (1957) Phytoseiidae et Aceosejidae (Acarina: Gamasina) d’Algerie. I. Genres Blattisocius Keegan, Iphiseius, Berlese, Amblyseius Berlese, Phytoseius Ribaga, Phytoseiulus Evans. Bulletin de la Societe d’Histore Natrelle de l’Afrique du Nord, 48, 319–352.
- Athias-Henriot, C. (1961) Mesostigmates (Urop. excl.) edaphiques Mediterraneens (Acaromorpha, Anactinotrichida). Acarologia, 3, 381–509.
- Athias-Henriot, C. (1962) Amblyseius swirskii, un nouveau phytoseiide voisin A. andersoni (Acariens anactinotriches). Annales de l’Ecole Nationale d’ Agriculture d’ Alger, 3, 1–7.
- Athias-Henriot, C. (1977) Nouvelles notes sur les Amblyseiini. III. Sur le genre Cydnodromus: Redefinition, composition (Parasitiformes, Phytoseiidae). Entomophaga, 22, 61–73. https://doi.org/10.1007/BF02372991
- Berlese, A. (1889) Acari, Myriapoda et Scorpiones hucusque in Italia seperta. Tipografia del Seminario, Padova, 4 pls. [pls. 6 + 7–9]
- Berlese, A. (1913) Acaroteca Italica. Tipografia di M. Ricci, Firenze, 221 pp.
- Berlese, A. (1914) Acari nuovi. Manipulus IX. Redia, 10, 113–150.
- Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12, 11–66.
- Berlese, A. (1921) Acari, Myriapoda et Pseudoscorpiones hucusque in Italia reperta. I. Indice sinonimico dei generi e delle species illustrate nei fascicoli 1 a 101. Redia, 14, 77–108.
- Blommers, L. (1974) Species of the genus Amblyseius Berlese, 1914, from Tamatave, east Madagascar (Acarina: Phytoseiidae). Bulletin Zoologisch Museum Universiteit van Amsterdam, 3, 143–155.
- Canestrini, G. & Fanzago, F. (1876) Nuovi acari italiani (Seconda Serie). Atti Societa Veneto-Trentina Scienze Naturali, 5, 130–142.
- Chant, D.A. (1957) Descriptions of some phytoseiid mites (Acarina: Phytoseiidae). The Canadian Entomologist, 189, 289–308. https://doi.org/10.4039/Ent89289-7
- Chant, D.A. (1959) Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of 38 new species. The Canadian Entomologist, 91, 1–166.
- Chant, D.A., Hansell, R.I.C. & Yoshida-Shaul, E. (1974) The genus Typhlodromus Scheuten (Acarina: Phytoseiidae) in Canada and Alaska. Canadian Journal of Zoology, 52, 1265–1291. https://doi.org/10.1139/z74-168
- Chant, D.A. & McMurtry, J.A. (1994) A review of the subfamilies Phytoseiinae and Typhlodrominae (Acari: Phytoseiidae). International Journal of Acarology, 20, 223–310. https://doi.org/10.1080/01647959408684022
- Chant, D.A. & McMurtry, J.A. (2003a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part I. Neoseiulini new tribe. International Journal of Acarology, 29, 3–46. https://doi.org/10.1080/01647950308684319
- Chant, D.A. & McMurtry, J.A. (2003b) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae). Part II. The tribe Kampimodromini Kolodochka. International Journal of Acarology, 29, 179–224. https://doi.org/10.1080/01647950308684331
- Chant, D.A. & McMurtry, J.A. (2004) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part III. The tribe Amblyseiini Wainstein, subtribe Amblyseiina n. subtribe. International Journal of Acarology, 30, 171–228. https://doi.org/10.1080/01647950408684388
- Chant, D.A. & McMurtry, J.A. (2005a) A review of the subfamily Amblyseiinae Muma (Acari; Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiini n. subtribe, and Ricoseiina n subtribe. International Journal of Acarology, 31, 187–224. https://doi.org/10.1080/01647950508684424
- Chant, D.A. & McMurtry, J.A. (2005b) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VII. Typhlodromipsini n. tribe. International Journal of Acarology, 31, 315–340. https://doi.org/10.1080/01647950508683673
- Chant, D.A. & McMurtry, J.A. (2006) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VIII. The tribes Macroseiini Chant, Denmark and Baker, Phytoseiulini n. tribe, Afroseiulini n. tribe and Indoseiulini Ehara & Amano. International Journal of Acarology, 32, 13–25. https://doi.org/10.1080/01647950608684439
- Chant, D.A. & McMurtry, J.A. (2007) Illustrated Keys and Diagnoses for the Genera and Subgenera of the Phytoseiidae of the World (Acari: Mesostigmata). Indira Publishing House, West Bloomfield, Michigan, pp. 220.
- Collyer, E. (1964) The occurrence of some mites of the family Phytoseiidae in New Zealand, and descriptions of seven new species. Acarologia, 6, 632–646.
- Corpuz, L.A. & Rimando, L. (1966) Some Philippine Amblyseiinae (Phytoseiidae: Acarina). The Philippine Agriculturist, 50, 114–136.
- De Leon, D. (1959) Two new genera of phytoseiid mites with a note on Proprioseius meridionalis Chant (Acarina: Phytoseiidae). Entomological News, 70, 257–262.
- De Leon, D. (1962) Twenty-three new phytoseiids, mostly from southeastern United States (Acarina: Phytoseiidae). The Florida Entomologist, 45, 11–27. https://doi.org/10.2307/3492899
- De Leon, D. (1965) A note on Neoseiulus Hughes 1948 and a new synonymy. Proceedings of the Entomological Society of Washington, 87, 23.
- Demite, P., McMurtry, J.A. & Moraes, G.J. de (2014). Phytoseiidae Database: a website for taxonomic and distributional information on phytoseiid mites (Acari). Zootaxa, 3795 (5), 571–577. https://doi.org/10.11646/zootaxa.3795.5.6
- Demite, P.R., Moraes, G.J. de, McMurtry, J.A., Denmark, H.A. & Castilho, R.C. (2025) Phytoseiidae Database. Available from: http://www.lea.esalq.usp.br/phytoseiidae/ (accessed 28 July 2025)
- Dosse, G. (1958) Uber einige neue Raubmilbenarten (Acar.: Phytoseiidae). Pflanzenschutz Berichte, 21, 44–61.
- Ehara, S. (1958) Three predatory mites of the genus Typhlodromus from Japan (Phytoseiidae). Annotationes Zoologicae Japonenses, 31, 53–57.
- Ehara, S. (1959) Some predatory mites of the genera Typhlodromus and Amblyseius from Japan (Phytoseiidae). Acarologia, 1, 285–295.
- Ehara, S. (1962) Notes on some predatory mites (Phytoseiidae and Stigmaeidae). Japanese Journal of Applied Entomology and Zoology, 6, 53–60. https://doi.org/10.1303/jjaez.6.53
- Ehara, S. (1964) Some mites of the families Phytoseiidae and Blattisociidae from Japan (Acarina: Mesostigmata). Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology, 15, 378–394.
- Ehara, S. (1966) A tentative catalogue of predatory mites of Phytoseiidae known from Asia, with descriptions of five new species from Japan. Mushi, 39, 9–30.
- Ehara, S. (1967a) Phytoseiid mites from Hokkaido (Acarina: Mesostigmata). Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology, 16, 212–233.
- Ehara, S. (1967b) Phytoseiid mites from Okinawa Island (Acarina: Mesostigmata). Mushi, 40, 67–82.
- Ehara, S. (1972) Some phytoseiid mites from Japan, with descriptions of thirteen new species (Acarina: Mesostigmata). Mushi, 46, 137–173.
- Ehara, S. (1977) A review of taxonomic studies on natural enemies of spider mites in Japan. Review of Plant Protection Research, 10, 29–48.
- Ehara, S. (1978) Two new species of phytoseiid mites from Hokkaido (Acarina: Phytoseiidae). Proceedings of the Japan Academy, Series B, 54, 446–450. https://doi.org/10.2183/pjab.54.446
- Ehara, S. (1981) Description of a new Typhlodromus from miso factories, with synonymy of T. bambusae Ehara (Acarina: Phytoseiidae). Japanese Journal of Sanitary Zoology, 32, 235–237. https://doi.org/10.7601/mez.32.235
- Ehara, S. (1982) Two new species of phytoseiid mites from Japan (Acarina: Phytoseiidae). Applied Entomology and Zoology, 17, 40–45. https://doi.org/10.1303/aez.17.40
- Ehara, S. (1985) Five species of phytoseiid mites from Japan with descriptions of two new species (Acarina, Phytoseiidae). Zoological Science, 2, 115–121.
- Ehara, S. & Amano, H. (1998) A revision of the mite family Phytoseiidae in Japan (Acari: Gamasina), with remarks on its biology. Species Diversity, 3, 25–73. https://doi.org/10.12782/specdiv.3.25
- Ehara, S. & Amano, H. (2002) Some Japanese phytoseiid mites (Acari: Phytoseiidae) mostly from Ishigaki and Taketomi Islands. Entomological Science, 5, 321–329.
- Ehara, S. & Amano, H. (2004) Checklist and keys to Japanese Amblyseiinae (Acari: Gamasina: Phytoseiidae). Journal of the Acarological Society of Japan, 13, 1–30. https://doi.org/10.2300/acari.13.1
- Ehara, S. & Gotoh, T. (Eds.) (2009) Colored Guide to the Plant Mites of Japan. Zenkoku Nôson Kyôiku Kyôkai, Tokyo, 349 pp. [in Japanese]
- Ehara, S. & Hamaoka, K. (1980) A new Typhlodromus from Japan with notes on four other species of phytoseiid mites. Acta Arachnologica, 29, 3–8. https://doi.org/10.2476/asjaa.29.3
- Ehara, S. & Kishimoto, H. (2007a) Description of a new species of Phytoscutus (Acari: Phytoseiidae) from Kyushu, Japan. International Journal of Acarology, 33, 111–113. https://doi.org/10.1080/01647950708684509
- Ehara, S. & Kishimoto, H. (2007b) The occurrence of Typhlodromus (Anthoseius) transvaalensis (Nesbitt) (Acari: Phytoseiidae) in Japan. Journal of the Acarological Society of Japan, 16, 139–143. https://doi.org/10.2300/acari.16.139
- Ehara, S. & Yokogawa, M. (1977) Two new Amblyseius from Japan with notes on three other species (Acarina: Phytoseiidae). Proceedings of the Japanese Society of Systematic Zoology, 13, 50–58.
- Ehara, S., Okada, Y. & Kato, H. (1994) Contribution to the knowledge of the mite family Phytoseiidae in Japan (Acari: Gamasina). Journal of the Faculty of Education, Tottori University, Natural Science, 42, 119–160.
- Ehara, S., Gotoh, T. & Amano, H. (2000) Two Japanese species of the genus Paraphytoseius Swirski and Shechter (Acari: Phytoseiidae). Journal of the Acarological Society of Japan, 9, 113–118. https://doi.org/10.2300/acari.9.113
- Ehara, S., Toyoshima, S. & Amano, H. (2006) Three new species of phytoseiid mites (Acari: Phytoseiidae) from Japan. International Journal of Acarology, 32, 345–350. https://doi.org/10.1080/01647950608684480
- Evans, G.O. (1952) On a new predatory mite of economic importance. Bulletin of Entomological Research, 43, 397–401. https://doi.org/10.1017/S0007485300040566
- Evans, G.O. (1953) On some mites of the genus Typhlodromus Scheuten, 1857, from S. E. Asia. Annals and Magazine of Natural History, Series 12, 6 (66), 449–467. https://doi.org/10.1080/00222935308654444
- Evans, G.O. & Macfarlane, D. (1962) (1961) A new mites of the genus Phytoseius Ribaga (Acari: Mesostigmata). Annual Magazine of Natural History, Series 13, 4 (46), 587–588. https://doi.org/10.1080/00222936108651183
- Funayama, K. (2024) Impact of occurrence of Amblyseius andersoni (Acari: Phytoseiidae) on two native phytoseiid (Mesostigmata) mites in a Japanese apple (Rosaceae) orchard. The Canadian Entomologist, 156, e18. https://doi.org/10.4039/tce.2024.20
- Garman, P. (1948) Mite species from apple trees in Connecticut. Bulletin of the Connecticut Agricultural Experimental Station, 520, 5–27.
- Garman, P. (1958) New species belonging to the genera Amblyseius and Amblyseiopsis with keys to Amblyseius, Amblyseiopsis, and Phytoseiulus. Annals of the Entomological Society of America, 51, 69–79. https://doi.org/10.1093/aesa/51.1.69
- Garman, P. & McGregor, E.A. (1956) Four new predaceous mites (Acarina: Phytoseiidae). Southern California Academy of Science Bulletin, 55, 7–13.
- Hada, H. (2012) Monitoring survey of predatory mite living in apple orchard in Iwate prefecture. Annual Report of Plant Protection of North Japan, 63, 215–217. [in Japanese]
- Hirschmann, W. (1962) Gangsystematik der Parasitiformes. Acarologie. Schriftenreihe fūr Vergleichende Milbenkunde, 5, 1–56.
- Hughes, A.M. (1948) The mites associated with stored food products. Ministry of Agriculture and Fisheries, H. M. Stationary Office, London, 168 pp.
- Kaneko, S., Shirotsuka, K., Shibao, M. & Yamanaka, S. (2018) Species composition and seasonal occurrence of phytoseiid mites on weeds around the eggplant field in Osaka prefecture. Annual Report of The Kansai Plant Protection Society, 60, 139–141. [in Japanese with English summary] https://doi.org/10.4165/kapps.60.139
- Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes): Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands und der angrenzenden Meeresteile. 59. Teil. VEB Gustav Fischer Verlag, Jena, 475 pp.
- Karg, W. (1989) Neue Raubmilbenarten der Gattuig Proprioseiopsis Muma, 1961 (Acarina, Parasitiformes) mit Bestimmungsschlusseln. Zoologische Jahrbucher Systematik, 116, 199–216.
- Karg, W. (1991) Die Raubmilbenarten der Phytoseiidae Berlese (Acarina) Mitteleuropas sowie angrenzender Gebiete. Zoologische Jahrbücher Systematik, 118, 1–64.
- Karg, W. & Oomen-Kalsbeek, F. (1987) Neue Raubmilbenarten der Gattung Amblyseius Berlese (Acarina, Parasitiformes, Phytoseiidae) - Antagonisten der unechten Spinnmilbe Brevipalpus phoenicis Geijskes. Zoologische Jahrbucher Systematik, 114, 131–140.
- Kato, M. (2021) Native predatory mites collected from undergrowth in commercial apple orchards in Iwate prefecture, northern Japan. Annual Report of Plant Protection of North Japan, 63, 215–217. [in Japanese]
- Koch, C.L. (1839) Deutschlands Crustaceen, Myriapoden und Arachniden. Vols. 5 & 6. Fascicle 25 & Fascicle 27. Friedrich Pustet, Regensburg, 3 tabulates. [Tabulat 22 & Tabulae 6 + 13]
- Kishimoto, H. (2002) Species composition and seasonal occurrence of spider mites (Acari: Tetranychidae) and their predators in Japanese pear orchards with different agrochemical spraying programs. Applied Entomology and Zoology, 37, 603–615. https://doi.org/10.1303/aez.2002.603
- Kishimoto, H., Teshiba, M., Kondoh, T., Miyazaki, T., Sugiura, N., Toda, S., Yamazaki, R., Wakatsuki, H., Motoyama, H. & Horie, H. (2007) Occurrence of Neoseiulus californicus (Acari: Phytoseiidae) on citrus in Kyushu district, Japan. Journal of the Acarological Society of Japan, 16, 129–137. [in Japanese with English summary] https://doi.org/10.2300/acari.16.129
- Koike, A., Nemoto, H. & Amano, H. (2000) New trap for survey of species structure and seasonal dynamics of phytoseiid mites on Japanese pear trees (Acari: Phytoseiidae). Japanese Journal of Applied Entomology and Zoology, 44, 35–40. [in Japanese with English summary] https://doi.org/10.1303/jjaez.2000.35
- Kolodochka, L.A. (1998) Two new tribes and the main results of a revision of Palearctic phytoseiid mites (Parasitiformes, Phytoseiidae) with the family system concept. Vestnik Zoologii, 32, 51–63. [in Russian]
- Komi, K., Arakawa, R. & Amano, H. (2008) Native phytoseiid mites (Acari: Phytoseiidae) occurring on greenhouse vegetable crops under the pest control programs with natural enemies in Kochi prefecture, Japan. Journal of the Acarological Society of Japan, 17, 23–28. [in Japanese with English summary] https://doi.org/10.2300/acari.17.23
- Kostiainen, T. & Hoy, M.A. (1996) The Phytoseiidae as biological control agents of pest mites and insects. A bibliography. Monograph 17. Agricultural Experiment Station, Institute of Food and Agricultural Sciences (IFAS), University of Florida, Gainesville, Florida, 355 pp.
- Kunimoto, Y., Nakama, T., Amano, H. & Takafuji, A. (2009) Species composition of predators of spider mites in persimmon orchards in Nara prefecture with reference to the effect of weed management. Journal of the Acarological Society of Japan, 18, 7–16. [in Japanese with English summary] https://doi.org/10.2300/acari.18.7
- Matthysse, J.G. & Denmark, H.A. (1981) Some phytoseiids of Nigeria (Acarina: Mesostigmata). The Florida Entomologist, 64, 340–357. https://doi.org/10.2307/3494585
- McGregor, E.A. (1954) Two new mites in the genus Typhlodromus (Acarina: Phytoseiidae). Southern California Academy of Science Bulletin, 53, 89–92.
- McMurtry, J.A., Moraes, G.J. de & Sourassou, N.F. (2013) Revision of the lifestyles of phytoseiid mites (Acari: Phytoseiidae) and implications for biological control strategies. Systematic and Applied Acarology,18, 297–320. https://doi.org/10.11158/saa.18.4.1
- Miyazaki, T., Kishimoto, H., Suga, Y. & Teramoto, T. (2012) The Occurrence of the native natural enemy of citrus red mite in citrus orchard in Nagasaki prefecture. Bulletin of the Nagasaki Agricultural & Forestry Technical Development Center, 3, 121–139. [in Japanese with English summary]
- Mochizuki, M. (2005) Release process of introduced phytoseiid mites in apple orchards and native phytoseiid mite fauna. Plant Protection, 59, 501–504. [in Japanese]
- Mochizuki, M. (2014) Seasonal occurrence and species composition of phytoseiid mites and phytophagous thrips on forage soybean with a view to conservation of phytoseiid mites in vineyards. Journal of the Acarological Society of Japan, 23, 79–89. [in Japanese with English summary] https://doi.org/10.2300/acari.23.79
- Moraes, G.J. de & Mesa, N.C. (1988) Mites of the family Phytoseiidae (Acari) in Colombia, with descriptions of three new species. International Journal of Acarology, 14, 71–88. https://doi.org/10.1080/01647958808683790
- Moraes, G.J. de, McMurtry, J.A. & Denmark, H.A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA Departamento de Difusão de Tecnologia, Brasília, 353 pp.
- Moraes, G.J. de, McMurtry, J.A., Denmark, H.A. & Campos, C.B. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1–494. https://doi.org/10.11646/zootaxa.434.1.1
- Muma, M.H. (1955) Phytoseiidae (Acarina) associated with citrus in Florida. Annals of the Entomological Society of America, 48, 262–272. https://doi.org/10.1093/aesa/48.4.262
- Muma, M.H. (1961) Subfamilies, genera and species of Phytoseiidae (Acarina: Mesostigmata). Bulletin of the Florida State Museum, 5, 267–302. https://doi.org/10.58782/flmnh.tqpo4380
- Muma, M.H. (1963) The genus Galendromus Muma, 1961 (Acarina: Phytoseiidae). The Florida Entomologist, Supplement 1, 15–41. https://doi.org/10.2307/3493355
- Narayanan, E.S., Kaur, R.B. & Ghai, S. (1960) Importance of some taxonomic characters in the family Phytoseiidae Berl. 1916 (predatory mites) with new records and descriptions of species. Proceeding of the National Institute of Science of India, 2613, 384–394.
- Negm, M.W. & Gotoh, T. (2020) A new species of Amblyseius Berlese (Acari: Phytoseiidae) from Japan. Biologia, 75, 1977–1981. https://doi.org/10.2478/s11756-020-00519-6
- Negm, M.W., Alatawi, F.J. & Aldryhim, Y.N. (2012a) Incidence of predatory phytoseiid mites in Saudi Arabia: new records and a key to the Saudi Arabian species (Acari: Mesostigmata: Gamasina: Phytoseiidae). Systematic & Applied Acarology, 17, 261–268. https://doi.org/10.11158/saa.17.3.6
- Negm, M.W., Alatawi, F.J. & Aldryhim, Y.N. (2012b) A new species of Neoseiulus Hughes, with records of seven species of predatory phytoseiid mites associated with date palm in Saudi Arabia (Acari: Phytoseiidae). Zootaxa, 3356 (1), 57–64. https://doi.org/10.11646/zootaxa.3356.1.3
- Negm, M.W., Matsuda, T., Kayukawa, T., Ho, C.-C., Hsu, Y.-T., Kongchuensin, M., Konvipasruang, P. & Gotoh, T. (2021) Morphological ontogeny and molecular analyses of geographic strains of two closely related Neoseiulus species (Acari: Phytoseiidae). Acarologia, 61, 432–452. https://doi.org/10.24349/acarologia/20214440
- Nesbitt, H. (1951) A taxonomic study of the Phytoseiinae (Family Laelaptidae) predacious upon Tetranychidae of economic importance. Zoologische Verhandelingen, 12, 1–64.
- Ohno, S., Gotoh, T., Miyagi, A., Ganaha-Kikumura, T., Kurima, M., Kijima, K. & Ooishi, T. (2012) Geographic distribution of phytoseiid mite species (Acari: Phytoseiidae) on crops in Okinawa, a subtropical area of Japan. Entomological Science, 15, 115–120. https://doi.org/10.1111/j.1479-8298.2011.00469.x
- Oudemans, A.C. (1915) Acarologische Aantekeningen. LVI. Entomologische Berichten, 4, 308–316. https://doi.org/10.5962/bhl.part.1128
- Oudemans, A.C. (1930) Acarologische Aantekeningen CIII. Entomologische Berichten, 8, 4–12.
- Prasad, V. (1968) Some phytoseiid mites from Hawaii. Annals of the Entomological Society of America, 61, 1459–1462. https://doi.org/10.1093/aesa/61.6.1459
- Ribaga, C. (1904) (1902) Gamasidi planticoli. Rivista di Patologia Vegetale, 10, 175–178.
- Pritchard, A.E. & Baker, E.W. (1962) Mites of the family Phytoseiidae from Central Africa, with remarks on genera of the world. Hilgardia, 33, 205–309. https://doi.org/10.3733/hilg.v33n07p205
- Ryu, M.O. (1993) A review of the Phytoseiidae (Mesostigmata: Acarina) from Korea. Insecta Koreana, 10, 92–137.
- Santoso, S., Takafuji, A. Amano, H. & Ozawa, A. (2004) Species composition of phytoseiid mites (Acari: Phytoseiidae) in tea fields with different management practices in Shizuoka prefecture, Japan. Journal of the Acarological Society of Japan, 13, 77–82. https://doi.org/10.2300/acari.13.77
- Sasaki, M., Anazawa, T. & Akai, H. (2010) Species composition of spider mites and phytoseiid mites in peach orchards in Fukushima prefecture. Tohoku Agricultural Research, 62, 119–120. [in Japanese]
- Schicha, E. (1975) A new predacious species of Amblyseius Berlese from strawberry in Australia, and A. longispinosus (Evans) redescribed (Acari: Phytoseiidae). Journal of the Australian Entomological Society, 14, 101–106. https://doi.org/10.1111/j.1440-6055.1975.tb02010.x
- Schicha, E. (1982) A new species of Amblyseius Berlese from the New Hebrides with notes on closely related species from Australia, Japan and China (Acari: Phytoseiidae). Australian Entomology Magazine, 9, 33–38.
- Schicha, E. (1983) New species, new records, and redescriptions of phytoseiid mites from Australia, Tahiti, and the African region (Acari: Phytoseiidae). International Journal of Entomology, 25, 103–126.
- Scheuten, A. (1857) Einiges über Milben. Archiv für Naturgeschichte, 23, 104–112.
- Schuster, R. & Pritchard, A. (1963) Phytoseiid mites of California. Hilgardia, 34, 191–285. https://doi.org/10.3733/hilg.v34n07p191
- Swirski, E. & Schechter, R. (1961) Some Phytoseiid mites (Acarina: Phytoseiidae) of Hong Kong, with a description of a new genus and seven new species. Israel Journal of Agricultural Research, 11, 97–117.
- Takahashi, F. & Chant, D.A. (1993) Phylogenetic relationships in the genus Phytoseiulus Evans (Acarina: Phytoseiidae). I. Geographic distribution. International Journal of Acarology, 19, 15–22. https://doi.org/10.1080/01647959308683534
- Toyoshima, S. (2003) A candidate of predatory phytoseiid mites (Acari: Phytoseiidae) for the control of the European red mite, Panonychus ulmi (Koch), (Acari: Tetranychidae) in Japanese apple orchards. Applied Entomology and Zoology, 38, 387–391. https://doi.org/10.1303/aez.2003.387
- Toyoshima, S. & Amano, H. (2012) Discrimination technique of Neoseiulus cucumeris (Oudemans) and Amblyseius swirskii Athias-Henriot from morphologically similar Japanese phytoseiid species. Japanese Journal of Applied Entomology and Zoology, 56, 176–179. https://doi.org/10.1303/jjaez.2012.176
- Toyoshima, S., Ihara, F. & Amano H. (2008) Diversity and abundance of phytoseiid mites in natural vegetation in the vicinity of apple orchards in Japan. Applied Entomology and Zoology, 43, 443–450. https://doi.org/10.1303/aez.2008.443
- Toyoshima, S., Kishimoto, H. & Amano, H. (2013) Phytoseiid Mite Portal. Available from: http://phytoseiidae.acarology-japan.org/ (accessed 10 February 2026)
- Toyoshima, S., Kishimoto, H., Morii, H. & Amano, H. (2014) Occurrence of Typhlodromips sessor (De Leon) (Acari: Phytoseiidae) on Mexican sunflower, Tithonia rotundifolia (Miller) (Asteridae: Asteraceae) planted around a tea plantation in Japan. Journal of the Acarological Society of Japan, 23, 29–33. https://doi.org/10.2300/acari.23.29
- Toyoshima, S., Kishimoto, H., Kaneko, M., H. & Amano, H. (2016) Occurrence of Amblyseius andersoni (Chant) (Acari: Phytoseiidae) in deciduous fruit tree orchards in Japan. Journal of the Acarological Society of Japan, 25, 37–43. https://doi.org/10.2300/acari.25.37
- Ueckermann, E.A. & Loots, G.C. (1985) Trochoseius Pritchard & Baker, a new subgenus of Amblyseius Berlese with notes on its former genus Iphiseius Berlese (Acari: Phytoseiidae). Phytophylactica, 17, 129–137.
- Ueckermann, E.A. & Loots, G.C. (1988) The African species of the subgenus Anthoseius De Leon and Amblyseius Berlese (Acari: Phytoseiidae). Entomology Memoir, Department of Agriculture and Water Supply, Republic of South Africa, 73, 1–168.
- Van der Merwe, G. (1965) South African Phytoseiidae (Acarina). I. Nine new species of the genus Amblyseius Berlese. Journal of the Entomological Society of South Africa, 28, 57–76.
- Van der Merwe, G. (1968) A taxonomic study of the family Phytoseiidae (Acari) in South Africa with contributions to the biology of two species. Entomology Memoirs, 18, 1–194.
- Wainstein, B.A. (1962) Revision du genre Typhlodromus Scheuten, 1857 et systematique de la famille des Phytoseiidae (Berlese, 1916) (Acarina: Parasitiformes). Acarologia, 14, 5–30.
- Wainstein, B.A. (1976) A new tribe of the family Phytoseiidae (Parasitiformes). Zoologicheskii Zhurnal, 55, 696–700.
- Wainstein, B.A. & Beglyarov, G.A. (1971) New species of the genus Amblyseius (Parasitiformes: Phytoseiidae) from the Primorsky Territory. Zoologicheskii Zhurnal, 50, 1803–1812. [in Russian]
- Wu, W.N. & Li, Z.Q. (1985) Three new species of the genus Amblyseius Berlese from south China (Acarina: Phytoseiidae). Acta Zootaxonomica Sinica, 10, 268–272. [in Chinese].
- Yamashiro, M. & Wakamasu, M. (2006) Distribution, seasonal occurrence and susceptibility to insecticides of phytoseiid mite on Japanese pear and the orchard ground cover in Tochigi prefecture. Annual Report of the Kanto-Tosan Plant Protection Society, 53, 129–135. [in Japanese]
- Yamazaki, D. & Itoyama, K. (2015) Seasonal occurrence of phytoseiid mites in urbanized Japanese pear orchards. Annual Report of the Kanto-Tosan Plant Protection Society, 62, 153–155. [in Japanese]
- Zhu, Z.-M. & Chen, S.M. (1983) A key to the species of genus Amblyseius from Jiangxi province with notes on four species (Acarina: Phytoseiidae). Natural Enemies of Insects, 5, 181–187. [in Chinese]
