Abstract
Two new cavernicolous pseudoscorpion species, Ephippiochthonius boritiensis sp. nov. from Kozmani Cave, and Ephippiochthonius trapezoides sp. nov. from Tsintskila Cave in Georgia (Caucasus) are described and illustrated. Furthermore, a key for all epigean and cavernicolous Ephippiochthonius species in Georgia is also provided.
References
- Antić, D., Margalitadze, A. & Šević, M. (2025) Four new cavernicolous species of the genus Acanthophorella Antić & Makarov, 2016 (Diplopoda, Chordeumatida, Anthroleucosomatidae) from Georgia, Caucasus. Zootaxa, 5609 (1), 41–69. https://doi.org/10.11646/zootaxa.5609.1.3
- Barjadze, Sh., Parimuchová, A., Raschmanová, N., Maghradze, E. & Kovač, Ľ. (2022) Two new species of Plutomurus Yosii (Collembola: Tomoceridae) from the Caucasus and central Europe. Zootaxa, 5169 (3), 252–266. https://doi.org/10.11646/zootaxa.5169.3.2
- Beier, M. (1963) Ordnung Pseudoscorpionidea (Afterskorpione). In: Bestimmungsbücher zur Bodenfauna Europas. Vol. 1. Akademie-Verlag, Berlin, pp. i–vi + 1–313. https://doi.org/10.1515/9783112748497
- Chamberlin, J.C. (1931) The arachnid order Chelonethida. Stanford University Publications. University Series. Biological Sciences. Vol. 7. No. 1. Stanford University Press, Stanford, California, 284 pp.
- Gardini, G. (2013) A revision of the species of the pseudoscorpions subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas. Zootaxa, 3655 (1), 1–151. https://doi.org/10.11646/zootaxa.3655.1.1
- Harvey, M.S. (2007) The smaller arachnid orders: diversity, descriptions and distributions from Linnaeus to the present (1758 to 2007). Zootaxa, 1668 (1), 363–380. https://doi.org/10.11646/zootaxa.1668.1.19
- Judson, M.L.I. (2007) A new and endangered species of the pseudoscorpion genus Lagynochthonius from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini (Arachnida, Chelonethi, Chthoniidae). Zootaxa, 1627 (1), 53–68. https://doi.org/10.11646/zootaxa.1627.1.4
- Kobakhidze, D. (1961) Die Standorte des Chthonius tetrachelatus (Preissler) in den verschiedenen Landschaftstypen der Georgischen SSR. Zoologischer Anzeiger, 167 (4/5), 166–169.
- Kobakhidze, D. (1964) Pseudoscorpione (Pseudoscorpiones). In: Animmal Kingdom of Georgia. Vol. II. Metsniereba, Tbilisi, pp. 30–47.
- Kolesnikov, V.B., Przhiboro, A.A. & Turbanov, I.S. (2023) The pseudoscorpions of the Caucasian Sphagnum bogs: part II. Description of Ephippiochthonius juliae sp.n. from Georgia, with remarks on the taxonomic status of some Caucasian species of the genus Ephippiochthonius Beier, 1930 (Arachnida: Pseudoscorpiones: Chthoniidae). Arthropoda Selecta, 32 (1), 65–74. https://doi.org/10.15298/arthsel.32.1.05
- Novák, J. & Gogshelidze, M. (2025) Three new cavernicolous Neobisium species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa, 5646 (4), 575–585. https://doi.org/10.11646/zootaxa.5646.4.6
- Parimuchová, A., Barjadze, S., Maghradze, E. & Kováč, Ľ. (2023) Two new species of the genus Deuteraphorura Absolon, 1901 (Hexapoda, Collembola, Onychiuridae) from Georgian caves with remarks on the subterranean biodiversity of the Caucasus Mountains. European Journal of Taxonomy, 879 (1), 64–82. https://doi.org/10.5852/ejt.2023.879.2161
- Sasmaz, A., Sasmaz, B. & Hein, J.R. (2020) Geochemical approach to the genesis of the Oligocene-stratiform manganese-oxide deposit, Chiatura (Georgia). Ore Geology Reviews, 128, 103910. https://doi.org/10.1016/j.oregeorev.2020.103910
- Schawaller, W. (1983) Pseudoskorpione aus dem Kaukasus (Arachnida). Stuttgarter Beiträge zur Naturkunde, 362 (A), 1–24.
- Schawaller, W. & Dashdamirov, S. (1988) Pseudoskorpione aus dem Kaukasus, Teil 2 (Arachnida). Stuttgarter Beiträge zur Naturkunde, 415 (A), 1–51.
- Turbanov, V.B. & Kolesnikov, I.S. (2023) First records of cave-dwelling pseudoscorpions of the genus Ephippiochthonius Beier, 1930 (Arachnida: Pseudoscorpiones: Chthoniidae) from the Caucasus, with description of three new species from Abkhazia and taxonomic comments on the genus. Zootaxa, 5325 (1), 41–62. https://doi.org/10.11646/zootaxa.5325.1.3
- World Pseudoscorpiones Catalog (2025) World Pseudoscorpiones Catalog. Natural History Museum Bern, Bern. Available from: http://wac.nmbe.ch (accessed 6 September 2025)
- Zaragoza, J.A. (2017) Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae). Zootaxa, 4246 (1), 1–221. https://doi.org/10.11646/zootaxa.4246.1.1
- Zaragoza, J.A., Novák, J., Gardini, G., Maghradze, E. & Barjadze, S. (2021) The taxonomic status of the Caucasian cave-dwelling pseudoscorpion Chthonius satapliaensis (Arachnida: Pseudoscorpiones). Zoology in the Middle East, 67, 356–364. https://doi.org/10.1080/09397140.2021.1965072
