Abstract
Of the 12 skipper taxa described by Jinhaku Sonan (1892-1984) from Taiwan and Hainan Island, 2 are recognized as valid species-level names, 4 as valid subspecies-level names, whereas the remaining 6 are considered junior subjective synonyms of other taxa. A revised status is established for a Hainan taxon, Tagiades japetus hainana Sonan, stat. rev., and Polytremis kiraizana (Sonan) is formally confirmed as a species endemic to Taiwan. The valid species-level names are Ochlodes niitakanus (Sonan) and Polytremis kiraizana (Sonan). The valid subspecies-level names are Lobocla bifasciata kodairai Sonan, Pseudocoladenia dan sadakoe Sonan & Mitono, Notocrypta feisthamelii arisana Sonan, and Tagiades japetus hainana Sonan. Tagiades menaka kotoshona Sonan is considered a junior subjective synonym of T. trebellius martinus Plötz, Ampittia maro matsumurai Sonan a junior subjective synonym of Ampittia dioscorides etura Mabille, Parnara kotoshona Sonan a junior subjective synonym of Parnara guttata (Bremer & Grey), Parnara ranrunna Sonan a junior subjective synonym of Caltoris cahira austeni (Moore), Gangara thyrsis hainana Sonan a junior subjective synonym of Gangara thyrsis thyrsis (Fabricius), and Telicota palmarum hainanum Sonan a junior subjective synonym of Cephrenes acalle oceanica (Mabille) in the present study.References
Anonymous. (1984) Obituary: Jinhaku Minamikawa. Japanese Journal of Applied Entomology and Zoology, 28, 187. [In Japanese]
Bridges, C. A. (1983) Lepidoptera: Hesperiidae, Notes on Species-Group Names. Published by author, Urbana, 274 pp.
Chiba, H & Eliot, J. N. (1991) A revision of the genus Parnara Moore (Lepidoptera, Hesperiidae), with special reference to the Asian species. Tyô to Ga, 42, 179–194.
Chiba, H., Hsu, Y. F. & Shirôzu, T. (1992) Hesperiidae. In: Heppner, J. N., Inoue, H. (Eds.) Lepidoptera of Taiwan, Vol. 1, part 2: checklist. Association for Tropical Lepidoptera, Gainesville, pp. 130–132.
Chiba, H & Tsukiyama, H. (1996) A review of the genus Ochlodes Scudder, 1872, with special reference to the Eurasian species (Lepidoptera: Hesperiidae). Butterflies, 14, 3–16. [In Japanese with English abstract]
Chu, Y. I. (2005) Insects into Focus: Entomology in Taiwan, 1864–1945. Yushan Publisher, Taipei, 614 pp. [In Chinese]
de Jong, R. & Treadaway C. G.. (1993) The Hesperiidae (Lepidoptera) of the Philippines. Zoologische Verhandelingen, 288, 1–139.
Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History). The British Museum (Natural History), London, xix + 502 pp.
Hamano, E. (1986) Ecological Encyclopedia of Taiwanese Butterflies. Kodansha, Tokyo, 474 pp. [In Japanese]
Hsu, Y. F. & Li, C. F. (1989) The revised taxonomic position of Notochrypta from Orchid Is. (Lepidoptera, Hesperiidae). Journal of the Taiwan Museum, 42, 9–13.
Hsu, Y. F. & Wang, S. M. (2005) Notes on immature biology of a rare skipper Caltoris bromus yanuca Fruhstorfer (Lepidoptera: Hesperiidae), with a discussion on the systematic status of Parnara ranrunna Sonan. Pan-Pacific Entomologist, 80, 70–81.
Hsu, Y. F., Chen, S. C. & Lu, C. C. (2007) On the immature biology of Pseudocoladenia dan sadakoe (Sonan & Mitono), with a discussion of the systematic status of this rare taxon (Lepidoptera: Hesperiidae). Pan-Pacific Entomologist , 83, 19–31.
Hsu, Y. F., Chiba, H., Yang, Y. C., Tsukiyama, H., Yang, P. S. & Wang S. M. (2006) A Taxonomic review of the Taiwanese skippers described by Shonen Matsumura (Lepidoptera: Hesperiidae). Zoological Studies, 45, 322–338.
Kawazoé, A. & Wakabayashi, M. (1976) Coloured Illustrations of the Butterflies of Japan. Hoikusha, Osaka, VII + 422 pp.
Maruyama, K. (1991) Butterflies of Borneo, 2 (2), Hesperiidae. Tobishima Corporation, Tokyo, xiii + 82 pp. [Japanese]; ix + 77 pp. [English]; 40 pls.
Morishita, K. (1978) Notocrypta. Yadoriga, 95/96: 15–33. [In Japanese]
Shih, H. T., Ho, C. C. & Chen, S. P. (2002) Insect and mite collection of Taiwan Agricultura; Research Institute: History and Characteristics. Co-work Museum Exhibit. National Museum of Natural Science, Taichung, pp. 65–68. [In Chinese]
Shih, H. T., Ho, C. C., Shih, C. W. & Feng, H. F. (2005) The establishment of digital museum on the type collection of the insect and mite collection of Taiwan Agriculture Research Institute. 2005 International workshop on integrated biodiversity and natural specimens databases and forum of species 2000 Asia-Oceania, National Museum of Natural Science, Taichung, pp. 91–107. [In Chinese with English summary]
Shimonoya, T. & Murayama, S. (1976) Remarks on some Formosan Rhopalocera with descriptions of two new species, and the revision of a few scientific names. Tyô to Ga, 27, 43–48. [In Japanese with English summary.]
Shirôzu, T. (1943) A revision of the genus Tagiades Hübner occurring in the Japanese Empire (Lepidoptera, Hesperiidae). Transactions of Kansai Entomological Society, 13, 9–24. [In Japanese]
Shirôzu, T. (1944) Notes on some Formosan butterflies (I). (Records of three species new to the fauna of Formosa). Transactions of Kansai Entomological Society, 14, 30–42. [In Japanese]
Shirôzu, T. (1960) Butterflies of Formosa in Colour. Hoikusha, Osaka, 481 pp. [In Japanese]
Sonan, J. (1930a) Some butterflies from Hainan Island. Transactions of the Natural History Society of Formosa, 20, 31–37. [In Japanese]
Sonan, J. (1930b) Notes on some butterflies from Formosa. Zephyrus, 2, 165–176, 1 pl. [In Japanese]
Sonan, J. (1936) Notes on some butterflies from Formosa. Zephyrus, 6, 205–216, 3 pls. [In Japanese]
Sonan, J. (1938a) A list of the butterflies of Hainan, with descriptions of two new sub-species of Hesperiidae. Transactions of the Natural History Society of Formosa, 28, 348–372, 2f. [In Japanese]
Sonan, J. (1938b) Notes on some butterflies from Formosa (5). Zephyrus, 7, 250–275, 1 pl. [In Japanese]
Sonan, J. & Mitono, T. (1936) On a new species of Hesperiidae, Coladenia sadakoe Sonan et Mitono. Zephyrus, 6,184–185. [In Japanese.]
Sugiyama, H. (1999) New butterflies from western China (VI). Pallarge 7: 1–14.
Tsukiyama, H. (1980) Miscellaneous notes on skippers of the Southeast Asia (2). Gekkan-Mushi, 118, 11–16. [In Japanese]
Tsukiyama, H., Chiba, H. & Fujioka, T. (1997) Hesperiidae. In: Fujioka, T. (Ed.), Japanese Butterflies and Their Relatives in the World, Vol. I. Shuppan Geijyutsu Sha, Tokyo, pp. 29–138. [In Japanese]
Uchida, H. (1991) Charms of Formosa, Island of Everlasting Summer: Butterflies of Taiwan – Nature and Man. Published by author, Numazu, 216 pp. [In Japanese.]
Uchida H. (1995) Everlasting Island with Beautiful Butterflies – Nature and Man. Published by author, Numazu, 208 pp. [In Japanese.]
Umeno, A. (1937) On the butterflies of Koushun Gun, Formosa, mainly around Kuraru area (IV). Zephyrus, 7, 142–149. [In Japanese]
Wu, Y. H. (1996) Forgotten Japanese Zoologists Who Contributed to the Fauna of Taiwan. Morning Star Publisher Inc., Taichung, 320 pp. [In Chinese]
Yamanaka, M. (1980) Distribution of Formosan butterflies. Tyô to Ga 30, suppl.1, 1–143. [In Japanese]