Abstract
A new species of Cyrtodactylus described from Nyemo County, Xizang Autonomous Region, China is here referred to a new species, Cyrtodactylus zhaoermii sp. nov. It is distinguished from all other Cyrtodactylus by the following characters: dorsal surface of the body with fine granules intermixed with larger sub-conical tubercles arranged into 20 more or less regular rows; proximal subdigital lamellae transversely expanded; 19–20 subdigital lamellae on toe IV; 30–32 midbody ventral scales; most scales in femoral region small, granular, series of 12–14 enlarged femoral scales lacking pores; a series of precloacal pores (4) present in male; tail with distinct segments, large, posteriorly directed tubercles in whorls, numbering nine to ten per caudal annulus, three on each side and three to four on the dorsum, situated at the posterior edge of each annulus; subcaudals not transversely expanded. The new species is the fourth Cyrtodactylus known from Xizang Autonomous Region.References
Anderson, S.C. (1999) The Lizards of Iran. Society for the Study of Amphibians and Reptiles. Ithaca, New York, 442 pp.
Bauer, A.M. (2002) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from Myanmar. Proceedings of the California Academy of Sciences, 53, 75–88.
Bauer, A.M., Sumontha, M. & Pauwels, O.S.G. (2003) Two new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Thailand. Zootaxa, 1376, 1–18.
Cervenka, J., Kratochvil, L. & Frynta, D. (2008) Phylogeny and taxonomy of the Middle Eastern geckos of the genus Cyrtopodion and their selected relatives. Zootaxa, 1931, 25–36.
Darevsky, I.S. & Szczerbak, N.N. (1997) A new gecko of the genus Gonydactylus (Sauria: Gekkonidae) with a key to the species from Vietnam. Asiatic Herpetological Research, 7, 19–22.
Duda, P.L. & Sahi, D.N. (1978) Cyrtodactylus mansarulus n. sp. from Jammu. Indian Science Congress Association Proceedings, 65, 211; (C).
Grismer, L.L. (2005) New species of bent-toed gecko (Cyrtodactylus Gray 1827) from Pulau Aur, Johor, West Malaysia. Journal of Herpetology, 39, 424–432.
Hayden, C.J., Brown, R.M., Gillespie, G., Iqbal Setiadi, M., Linkem, C.W., Iskandar, D.T., Umilaela Bickford, D.P., Riyanto, A. Mumpuni & Mcguire, J.A. (2008) A new species of bent-toed gecko Cyrtodactylus Gray 1827 (Squamata: Gekkonidae) from the island of Sulawesi, Indonesia. Herpetologica, 64, 109–120.
Heidrich, A., Roesler, H., Thanh, V.N., Boehme, W. & Ziegler, T. (2007) Another new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha - Ke Bang National Park, central Truong Son, Vietnam. Zootaxa, 1445, 35–48.
Khan, M.S. (1980) A new species of gecko from northern Pakistan. Pakistan Journal of Zoology, 12, 11–16.
Khan, M.S. (1993) A new angular-toed gecko from Pakistan, with remarks on the taxonomy and a key to the species belonging to genus Cyrtodactylus (Reptilia: Sauria: Gekkonidae). Pakistan Journal of Zoology, 25, 67–73.
Khan, M.S. (2001) Taxonomic notes on angular-toed Gekkota of Pakistan, with description of a new species of genus Cyrtopodion. Pakistan Journal of Zoology, 33, 13–24.
Khan, M.S. (2003) Questions of generic designation of angular-toed geckos of Pakistan with descriptions of three new genera (Reptilia: Gekkonidae). Journal of Natural History and Wildlife, 2, 1–9.
Khan, M.S. & Baig, K.J. (1992) A new Tenuidactylus gecko from northeastern Gilgit Agency, North Pakistan. Pakistan Journal of Zoology, 24, 273–277.
Kluge, A.G. (1983) Cladistic relationships among gekkonid lizards. Copeia, 1983, 465–475.
Kluge, A.G. (2001) Gekkotan lizard taxonomy. Hamadryad, 26, 1–209.
Kraus, F. (2007) A new species of Cyrtodactylus (Squamata: Gekkonidae) from western Papua New Guinea. Zootaxa, 1425, 63–68.
Krysko, K.L., Rehman, H. & Auffenberg, K. (2007) A new species of Cyrtopodion (Gekkonidae: Gekkoninae) from Pakistan. Herpetologica, 63, 100–113.
Leviton, A.E. & Anderson, S.C. (1984) Description of a new species of Cyrtodactylus from Afghanistan with remarks on the status of Gymnodactylus longipes and Cyrtodactylus fedtschenkoi. Journal of Herpetology, 18, 270–276.
Linkem, C.W., Mcguire, J.A., Hayden, C.J., Iqbal Setiadi, M., Bickford, D.P. & Brown, R.M. (2008) A new species of bent-toe gecko (Gekkonidae: Cyrtodactylus) from Sulawesi island, eastern Indonesia. Herpetologica, 64, 224–234.
Macey, J.R., Ananjeva, N.B., Wang, Y.Z., & Papenfuss, T.J. (2000) Phylogenetic relationships among Asian gekkonid lizards formerly of the genus Cyrtodactylus based on cladistic analyses of allozymic data: monophyly of Cyrtopodion and Mediodactylus. Journal of Herpetology, 34, 258–265.
Ngo, V.T. & Bauer, A.M. (2008) Descriptions of two new species of Cyrtodactylus Gray 1827 (Squamata : Gekkonidae) endemic to southern Vietnam. Zootaxa, 1715, 27–42.
Rao, D.Q. (2000) A complementary survey of the Herpeto-fauna of Xizang Autonomous Region (Tibet) with discussion of their distribution and current status [in Chinese]. Sichuan Journal of Zoology, 19, 107–112.
Roesler, H. & Glaw, F. (2008) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa, 1729, 8–22.
Sahi, D.N. & Duda, P.L. (1985) A checklist and keys to the amphibians and reptiles of Jammu and Kashmir state, India. Bulletin of Chicago Herpetological Society, 20, 86–97.
Schleich, H.H. & Kästle, W. (1998) Description of Gonydactylus nepalensis spec. nov. from the Inner Terai of far west Nepal (Reptilia: Sauria: Gekkonidae). In: Schleich H. H. & Kästle W. (Eds.), Contributions to the herpetology of South Asia (Nepal, India). Fuhlrott-Museum, Wuppertal. Pp. 269-280.
Szczerbak, N.N. & Golubev, M.L. (1984) On generic assignment of the Palearctic Cyrtodactylus lizard species (Reptilia, Gekkonidae) [in Russian]. Vestnik Zoologii, 2, 50–56.
Szczerbak, N.N. & Golubev, M.L. (1986) Gekkony fauny SSSR i sopredelnych stran [in Russian]. Naukova Dumka, Kiev, 234 pp.
TETCAS (Tibetan Expedition Team of the Chinese Academy of Science) (1982) Tibetan Physical Geology [in Chinese]. Science Press, Beijing, 178pp.
Xuan, H.Q., Orlov, N.L., Ananjeva, N.B., Johns, A.G., Hoang Ngoc, T. & Dau Quang, V. (2007) Description of a new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from the karst of North Central Vietnam. Russian Journal of Herpetology, 14, 98–106.
Underwood, G. (1954) On the classification and evolution of geckos. Proceedings of the Zoological Society of London, 124, 469–492.
Zhang, D.D. (2001) Tectonically controlled Fluvial land forms in Yaluzangbu River and their applications for the evolution of the river. Mountain Research and Development, 21, 61–68.
Zhao, E.M. & Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Oxford, Ohio. 521pp.