Abstract
The genus name Liodes and the family name Liodidae have been the cause of much confusion in zoology. The genus name Liodes has been used in Acari, Lepidoptera, Opiliones, and Coleoptera, and has often been misused, misspelled, or misunderstood. This paper reviews the history of these names and the related names Leiodes and Neoliodes, and concludes that (1) the names Leiodes Latreille 1796 and Leiodidae Fleming 1821 are available for a genus and family of beetles; (2) the names Neoliodes Berlese 1888 and Neoliodidae Sellnick 1928 are available for a genus and family of mites; (3) the names Liodes and Liodidae are not available for any group of animals, and should not be used.References
Aoki, J.I. (1963) An observation on the prevalence of Liodes sp. (Oribatei) in a main hall of a Buddhist temple in northern Japan. Japanese Journal of Sanitary Zoology, 14, 183–185.
Aoki, J.-I. (2006) New and newly recorded oribatid mites (Arachnida, Acari, Oribatida) from the Ryukyu Islands, Japan. Bulletin of the National Science Museum, Tokyo, Series A, 32, 105–124.
Baranowski, R. (1993) Revision of the genus Leiodes Latreille of North and Central America (Coleoptera: Leiodidae). Entomologica Scandinavia Supplement, 42, 1–149.
Berlese, A. (1888) Acari Austro-Americani quos collegit Aloysius Balzan. Manipulus primus. Species novas circiter quinquaginta complectens. Bollettino della Società Entomologica Italiana, 20, 171–222 + Plates V–XIII.
Brown, W.J. (1928) New Silphidae and Melyridae in the Canadian National Collection. The Canadian Entomologist, 60, 141–148.
Colloff, M.J. & Halliday, R.B. (1998) Oribatid Mites: A Catalogue of the Australian Genera and Species. CSIRO Publishing, Melbourne, 222 pp.
Crawford, R.L. (1992) Catalogue of the genera and type species of the harvestman superfamily Phalangoidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History, 8, 1–60.
Daffner, H. (1983) Revision der paläarktischen Arten der Tribus Leiodini Leach (Coleoptera, Leiodidae). Folia Entomologica Hungarica, 44 (2), 9–163.
Daffner, H. (1986) Die Arten der Gattung Leiodes Latreille (Coleoptera, Leiodidae) aus Indien und Nepal. Revue Suisse de Zoologie, 93, 71–87.
Decelle, J.E. (1988) Le genre Dietta Sharp, 1876 (Coleoptera: Liodidae) en Afrique continentale. Revue Zoologique Africaines, 102, 53–59.
Emetz, V.M. (1975) Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei. Nr. 358. Silphidae und Liodidae (Coleoptera). Folia Entomologica Hungarica, 28, 57–71.
Erichson, W.F. (1845) Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Dritter Band. Lieferungen I, II. Nicolaische Buchhandlung, Berlin, 320 pp.
Ermilov, S.G. (2009) Morphology of oribatid mite Poroliodes farinosus (Oribatida, Neoliodidae). Zoologicheskii Zhurnal, 88, 914–921.
Fall, H.C. (1910) New Silphidae of the tribe Anisotomini. The Canadian Entomologist, 42, 4–8.
Fernandez, N.A., Marcangeli, J.A. & Martinez, P.A. (1995) Contribution à la connaissance de la famille Liodidae en Argentine. I. Liodes marplatensis et Liodes elongatus, espèces nouvelles. Acarologia, 36, 247–259.
Fleming, J. (1821) Insecta. In: Supplement to the Fourth, Fifth and Sixth Editions of the Encyclopædia Britannica, Volume 5. Constable, Edinburgh, pp. 41–56. [not seen]
Gottwald, J. (1967) Liodes (Oreosphaerula) nitida sedlicaensis ssp. n. aus der Tschechoslowakei (Coleoptera, Liodidae). Acta Entomologica Bohemoslovaca, 64, 232–234.
Grandjean, F. (1936) Les Oribates de Jean Frédéric Hermann et de son Père [Arachn. Acar.]. Annales de la Société Entomologique de France, 105, 27–110.
Guenée, A. (1857) Histoire naturelle des Insectes. Species Général des Lépidoptères, 10, 1–584 + Plates 1–24.
Gumppenberg, C. von (1887) Systema Geometrarum zonae temperatioris septentrionalis. Systematische Bearbeitung der Spanner der nördlichen gemässsigten Zone. Nova Acta Academia Caesar Leopold Carolus, 52, 125–255. [not seen]
Halliday, R.B. (1998) Mites of Australia: A Checklist and Bibliography. CSIRO Publishing, Melbourne, 317 pp.
Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter Kongelige Danske Videnskabernes Selskab, 15 (2), 1–108 + Plates I–XLV.
Hatch, M.H. (1929) The genera and subgenera of Leiodidae and Clambidae. Journal of the New York Entomological Society, 37, 1–6.
Heethoff, M., Helfen, L. & Norton, R.A. (2009) Description of Neoliodes dominicus n. sp. (Acari, Oribatida) from Dominican amber, aided by synchrotron x-ray microtomography. Journal of Paleontology, 83, 153–159.
Hemming, F. (1947) On the question whether the names Liodes Heyden, 1826 (Class Arachnida, Order Acarina) and Leiodes Latreille, 1796 (Class Insecta, Order Coleoptera) are of the same origin and meaning. Bulletin of Zoological Nomenclature, 1, 223–224.
Hemming, F. (1953) Copenhagen Decisions on Zoological Nomenclature. Additions to, and Modifications of, the Règles Internationales de la Nomenclature Zoologique. International Trust for Zoological Nomenclature, London, 135 pp.
Heyden, C.H.G. von (1826) Versuch einer systematichen Einteilung der Acariden. Isis, 18, 608–613.
Hlisnikovsky, J. (1972) Beitrag zur Kenntnis der Liodidae (Coleoptera) von Ceylon. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 45, 131–149.
Hochberg, M.E., Bertault, G., Poitrineau, K. & Janssen, A. (2003) Olfactory orientation of the truffle beetle, Leiodes cinnamomea. Entomologia Experimentalis et Applicata, 109, 147–153.
Horn, G.H. (1880) Synopsis of the Silphidae of the United States with reference to the genera of other countries. Transaction of the American Entomological Society, 8, 219–322.
International Commission on Zoological Nomenclature (1943) Opinion 147. On the principles to be observed in interpreting Article 34 of the International Code in relation to the rejection, as homonyms, of generic and subgeneric names of the same origin and meaning as names previously published. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 2 (14), 125–129.
International Commission on Zoological Nomenclature (1954) Opinion 218. Addition of the generic name Liodes Heyden, 1826 (Class Arachnida) to the Official Index of Rejected and Invalid Generic Names in Zoology. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 4 (8), 83–92.
International Commission on Zoological Nomenclature (1961) International Code of Zoological Nomenclature. First Edition. International Trust for Zoological Nommenclature, London, 176 pp.
International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, 306 pp.
Jacot, A.P. (1929) Concerning the genus Neoliodes (Oribatoidea-Acarina). Transactions of the American Microscopical Society, 48, 30–43.
Jacot, A.P. (1947) On the question whether Liodes Heyden, 1826 (Class Arachnida) is a homonym of Leiodes Latreille, 1796 (Class Insecta, Order Coleoptra). Bulletin of Zoological Nomenclature, 1, 223.
Koch, L. (1869) Beitrag zur Kenntniss der Arachnidenfauna Tirols. Zeitschrift des Ferdinandeums für Tirol und Volarberg. Series 3, 14, 149–206. [not seen]
Koch, L. (1872) Beitrag zur Kenntniss der Arachnidenfauna Tirols. Zweite Abhandlung. Zeitschrift des Ferdinandeums für Tirol und Volarberg. Series 3, 17, 239–328. [not seen]
Latreille, P.A. (1796) Précis des Caractères Génériques des Insectes, Disposés dans un Ordre Naturel. Brive, Bourdeaux, 201 pp.
Le Conte, J.L. (1850) General remarks upon the Coleoptera of Lake Superior. In: Agassiz L. & Cabot, J.E. (Eds) Lake Superior, its Physical Character, Vegetation, and Animals, Compared with those of Other Regions. Gould, Kendall and Lincoln, Boston, USA. pp. 201–242.
Le Conte, J.L. (1859) Additions to the Coleopterous fauna of northern California and Oregon. Proceedings of the Academy of Natural Sciences of Philadelphia, 11, 281–292.
Le Conte, J.L. (1866) Additions to the Coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Natural Sciences of Philadelphia, 18, 361–394.
Leng, C.W. (1920) Catalogue of the Coleoptera of America, North of Mexico. John D. Sherman Jr., Mount Vernon, USA, 470 pp.
Mahunka, S. (2006) Some interesting beetle mites from Pacific islands collected by Antonius van Harten (Acari: Oribatida). (Acarologica Genavensia CVIII). Revue Suisse de Zoologie, 113, 579–593.
Majka, C.G. & Langor, D. (2008) The Leiodidae (Coleoptera) of Atlantic Canada: new records, faunal composition, and zoogeography. ZooKeys, 2, 357–402.
Mannerheim, C.G. (1852) Zweiter Nachtrag zur Käfer-Fauna der Nord-Amerikanischen Laender des Russichen Reiches. Bulletin de la Société Impériale des Naturalistes de Moscou, 25 (2), 283–387. [not seen]
Mannerheim, C.G. (1853) Dritter Nachtrag zur Kaefer-Fauna der Nord-Amerikanischen Laender des Russischen Rieches. Bulletin de la Société Impériale des Naturalistes de Moscou 26 (3), 95–273. [not seen]
Marshall, V.G., Reeves, R.M. & Norton, R.A. (1987) Catalogue of the Oribatida of continental United States and Canada. Memoirs of the Entomological Society of Canada, 139, 1–418.
Marske, K.A., Leschen, R.A.B., Barker, G.M. & Buckley, T.R. (2009) Phylogeography and ecological niche modelling implicate coastal refugia and trans-alpine dispersal of a New Zealand fungus beetle. Molecular Ecology, 18, 5126–5142.
McQuillan, P.B. (1996) The Tasmanian geometrid moths associated with the genus Amelora auctorum (Lepidoptera: Geometridae: Ennominae). Invertebrate Taxonomy, 10, 433–506.
Melville, R.V. & Smith, J.D.D. (1987) Official Lists and Indexes of Names and Works in Zoology. International Trust for Zoological Nomenclature, London, 366 pp.
Newton, A.F. (1998) Phylogenetic problems, current classification and generic catalog of world Leiodidae (including Cholevidae). In: Giachino, P.M. & Peck, S.B. (eds.), Phylogeny and Evolution of Subterranean and Endogean Cholevidae (=Leiodidae Cholevinae). Proceedings of XX I.C.E., Firenze, 1996. Atti Museo Regionale di Scienze Naturali, Torino. pp. 41–178.
Norton, R.A. (2006) First record of Collohmannia (C. schusteri n. sp.) and Hermannia (H. sellnicki n. sp.) from Baltic amber, with notes on Sellnick's genera of fossil oribatid mites (Acari : Oribatida). Acarologia, 46, 111–125.
Oudemans, A.C. (1937) Kritisch Historisch Overzicht der Acarologie, Deerde Gedeelte, 1805-1850 Band E + F. (E. J. Brill, Leiden). pp 1999–2735.
Panzer, G.W.F. (1797) Faune Insectorum Germanicae initia oder Deutschlands Insecten, Heft 37. Felsecker, Nürnberg, Germany.
Park, S.-J. & Ahn, K.-J. (2007) The Korean species of the genus Leiodes Latreille with descriptions of five new species (Coleoptera: Leiodidae: Leiodinae). Journal of the Kansas Entomological Society, 80, 27–42.
Peck, S.B. (2000) Leiodidae Fleming, 1821. In: Arnett, R.H., Jr. & Thomas, M.C. (eds.) American Beetles, Volume 1: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 250–258.
Reitter, E. (1885) Bestimmungs-Tabellen der Europäischen Coleopteren, XII. Necrophaga (Platypsyllidae, Leptinidae, Silphidae, Anisotomidae und Clambidae). Verhandlungen des Naturforschenden Vereines Brünn, 23, 3–122. [not seen]
Schmidt, W.L.E. (1841) Revision der deutschen Anisotomen. In: Germar, E.F. (ed.) Zeitschrift für die Entomologie, Volume 3, Friedrich Fleischer, Leipzig, Germany, pp. 130–202.
Seago, A.E. & Newton, A.F. (2009) A new genus of Leiodid beetle from Chile, with generic key and species checklist of described Neopelatopini (Coleoptera: Leiodidae: Camarinae). Annales Zoologici, 59, 297–304.
Sellnick, M. (1928) Formenkreis: Hornmilben, Oribatei. Die Tierwelt Mitteleuropas, 3, 1–42.
Sellnick, M. (1959) Acarina from Southeastern Polynesia – II (Oribatei). Occasional Papers of Bernice P. Bishop Museum, 22, 109–152.
Troukens, W. (2005) Truffelkevers en schimmelkevers aan de westrand van Brussel (Coleoptera: Liodidae & Scaphidiidae). Phegea, 33, 111–114.
Wallwork, J.A. (1963) Some Oribatei from Ghana. XII. The family Liodidae. (Ist. series). Acarologia, 5, 102–114.
Westwood, J.O. (1838) The Entomologist's Textbook: An Introduction to the Natural History, Structure, Physiology, and Classification of Insects Including the Crustacea and Arachnida. Wm. S. Orr and Co., London, England. 429 pp.
Willmann, C. (1931) Moosmilben oder Oribatiden (Oribatei). Die Tierwelt Deutschlands, 22, 79–200.