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Type: Articles
Published: 2010-09-03
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An annotated synopsis of Taiwanese bark and ambrosia beetles, with new synonymy, new combinations and new records (Coleoptera: Curculionidae: Scolytinae)

161/2 Mu 5, Soi Wat Pranon, T.Donkaew, A.Maerim, Chiangmai 50180, Thailand
Department of Science Education, National Museum of Natural Sciences, One Guancian Road, Taichung, Taiwan 404 Present address: National Central Library, 20 Chungshan S. Rd., Taipei 10001, Taiwan
Coleoptera Curculionidae Scolytinae Taiwan faunal synopsis new synonyms new combinations new records dubious records

Abstract

The state of knowledge of Taiwanese bark and ambrosia beetles (Coleoptera: Curculionidae: Scolytinae) is summarised to serve as a baseline for future studies of the fauna, with a checklist including information on distribution, host trees, biology and taxonomy. Six faunal elements based on geographical distribution are discussed in relation to their breeding sites and habits, and host preferences. One hundred and thirty-three species are recorded from Taiwan, of which thirtyeight are recorded for the first time. The following new synonymy is proposed: Ambrosiodmus lewisi (Blandford) (= Ozopemon tuberculatus Strohmeyer n. syn.), Anisandrus hirtus (Hagedorn) (= Xyleborus taiwanensis Browne n. syn.), Cyclorhipidion fukiensis (Eggers) (= Xyleborus tenuigraphus Schedl n. syn.), Cyrtogenius luteus (Blandford) (=Orosiotes formosanus Schedl n. syn.), Diuncus haberkorni (Eggers) (=Xyleborus taichuensis Schedl n. syn.) , Dryocoetes hectographus Reitter (=Ozopemon ater Eggers n. syn., =Dryocoetes formosanus Nobuchi n. syn.), Hypothenemus taihokuensis (Schedl) (=Hypothenemus cosmoderoides Murayama n. syn.), Xyleborinus artestriatus (Eichhoff) (= Xyleborus beaveri Browne n. syn.), Xyleborus pinicola Eggers (= Xyleborus pinivorus Browne n. syn.). The synonymy of Webbia camphorae Eggers and Webbia medius Eggers with Arixyleborus rugosipes Hopkins is confirmed. The following new combinations are given: Ernoporus formosanus (Browne) from Ptilopodius Hopkins; Ambrosiophilus hunanensis (Browne), A.metanepotulus (Eggers), A. satoi (Schedl), A. subnepotulus (Eggers), Cyclorhipidion formosanus (Browne), C. fukiensis (Eggers), C. ohnoi (Browne), C. xyloteroides (Schedl), Microperus kirishimanus (Murayama), Planiculus minutus (Blandford), all originally described in Xyleborus Eichhoff. Previous records of 23 species are considered dubious and require confirmation.

References

  1. Al Adawi, A.O., Deadman, M.L., Al Rawahi, A.K., Al Maqbaliz, Y.M., Al Jahwari, A.A., Al Saadi, B.A., Al Amri, I.S. & Wingfield, M.J. (2006) Aetiology and causal agents of mango sudden decline disease in the Sultanate of Oman. European Journal of Plant Pathology, 116, 247–254.

    Alonso-Zarazaga, M.A. & Lyal, C.H.C. (2009) A catalogue of family and genus group names in Scolytinae and Platypodinae with nomenclatural remarks (Coleoptera: Curculionidae). Zootaxa, 2258, 1–134.

    Annila, E. (1977) Seasonal flight patterns of spruce bark beetles. Annales Entomologici Fennici, 43, 31–35.

    Beaver, R.A. (1976) The biology of Samoan bark and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae). Bulletin of Entomological Research, 65, 531–548.

    Beaver, R.A. (1979a) Non-equilibrium ‘island’ communities. A guild of tropical bark beetles. Journal of Animal Ecology, 48, 987–1002.

    Beaver, R.A. (1979b) Host specificity of temperate and tropical animals. Nature, 281, 139–141.

    Beaver, R.A. (1984) The biology of the ambrosia beetle, Sueus niisimai (Eggers) (Col., Scolytidae), in Fiji. Entomologist’s Monthly Magazine, 120, 99–102.

    Beaver, R.A. (1988) Biological studies on ambrosia beetles of the Seychelles (Col., Scolytidae and Platypodidae). Journal of Applied Entomology, 105, 62–73.

    Beaver, R.A. (1989) Insect-fungus relationships in the bark and ambrosia beetles. In: Wilding, N., Collins, N.M., Hammond, P.M. & Webber, J.F. (eds.) Insect-fungus interactions. Academic Press, London. pp. 121–143.

    Beaver, R.A. (2000) The ambrosia beetle genus Indocryphalus Eggers (Coleoptera: Scolytidae): a new species from Peninsular Malaysia, new synonymy and a key to species. Serangga, 5, 165–179.

    Beaver, R.A. (2010) A review of the genus Hadrodemius Wood, with new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa, 2444, 51–57.

    Beaver, R.A. & Browne, F.G. (1979) The Scolytidae and Platypodidae (Coleoptera) of Penang, Malaysia. Oriental Insects, 12, 575–624.

    Beaver, R.A. & Gebhardt, H. (2005) Notes on the tribe Hyorrhynchini (Col., Curculionidae, Scolytinae). Serangga, 9, 91–102.

    Beaver, R.A. & Gebhardt, H. (2006) A review of the Oriental species of Scolytoplatypus Schaufuss (Coleoptera: Curculionidae: Scolytinae). Deutsche Entomologische Zeitschrift, 53, 155–178.

    Beaver, R.A. & Liu, L.Y. (2007) A new species of Scolytoplatypus Schaufuss (Coleoptera: Curculionidae: Scolytinae) from Taiwan. Entomologist’s Monthly Magazine, 143, 227–231.

    Beaver, R.A. & Maddison, P.A. (1990) The bark and ambrosia beetles of the Cook Islands and Niue (Coleoptera: Scolytidae and Platypodidae). Journal of Natural History, 24, 1365–1375.

    Beaver, R.A. & Shih, H.T. (2003) Checklist of Platypodidae (Coleoptera: Curculionoidea) from Taiwan. Plant Protection Bulletin, 44, 75–90.

    Beaver, R.A., Kajimura, H. & Goto, H. (2008) Taxonomic changes and new records of Japanese bark and ambrosia beetles (Coleoptera, Curculionidae, Scolytinae). Elytra, Tokyo, 36, 231–239.

    Beeson, C.F.C. (1929) Platypodidae and Scolytidae. Insects of Samoa, Part 4, Coleoptera, Fasc. 4, 217–248.

    Beeson, C.F.C. (1930) The biology of the genus Xyleborus with more new species. Indian Forest Records, 14, 209–272.

    Beeson, C.F.C. (1939) New species and biology of Coccotrypes and Thamnurgides (Scolytidae, Col.). Indian Forest Records, Entomology, (N.S.) 5, 279–308.

    Beeson, C.F.C. (1961) The ecology and control of the forest insects of India and the neighbouring countries. Government of India, New Delhi. 1007 pp. + 36 pl.

    Biedermann, P.H.W., Klepzig, K.D. & Taborsky, M. (2009) Fungus cultivation by ambrosia beetles: behavior and laboratory breeding success in three xyleborine species. Environmental Entomology, 38, 1096–1105.

    Blandford, W.F.H. (1893) The Scolyto-platypini, a new subfamily of Scolytidae. Transactions of the Entomological Society of London, 1893, 425–442 + pl. XIV.

    Blandford, W.F.H. (1894a) The rhynchophorous Coleoptera of Japan. Part III. Scolytidae. Transactions of the Entomological Society of London, 1894, 53–141.

    Blandford, W.F.H. (1894b) Supplementary notes on the Scolytidae of Japan, with a list of species. Transactions of the Entomological Society of London, 1894, 575–580.

    Blandford, W.F.H. (1894c) Notes on Scolytidae and their food plants. Insect Life, 6, 260–265.

    Blandford, W.F.H. (1895) A list of the Scolytidae collected in Ceylon by Mr. George Lewis, with descriptions of new species. Annals and Magazine of Natural History, ser.6, 15, 315–328.

    Blandford, W.F.H. (1896) Descriptions of new Scolytidae from the Indo-Malayan and Austro-Malayan regions. Transactions of the Entomological Society of London, 1896, 191–228.

    Blandford, W.F.H. (1898) On some Oriental Scolytidae of economic importance, with descriptions of five new species. Transactions of the Entomological Society of London, 1898, 423–430.

    Brader, L. (1964) Étude de la relation entre le scolyte des rameaux du caféier, Xyleborus compactus Eichh. (X.morstatti Hag), et sa plante-hôte. Mededelingen van de Landbouwhogeschool te Wageningen, 64(7), 1–109.

    Bright, D.E. (2000) Scolytidae (Coleoptera) of the Gunung Mulu National Park, Sarawak, Malaysia, with ecological notes and descriptions of six new species. Serangga, 5, 41–85.

    Bright, D.E., Jr. & Skidmore, R.E. (1997) A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 1 (1990– 1994). NRC Research Press, Ottawa. vii + 368 pp.

    Bright, D.E., Jr. & Skidmore, R.E. (2002) A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 2 (1995– 1999). NRC Research Press, Ottawa. viii + 523 pp.

    Browne, F.G. (1955) Synonymy and descriptions of some Oriental Scolytidae and Platypodidae (Coleoptera). Sarawak Museum Journal, 6, 343–373.

    Browne, F.G. (1961) The biology of Malayan Scolytidae and Platypodidae. Malayan Forest Records, 22, 1–255.

    Browne, F.G. (1963) Taxonomic notes on Scolytidae (Coleoptera). Entomologische Berichten, Amsterdam, 23, 53–59.

    Browne, F.G. (1968) Pests and diseases of forest plantation trees. Clarendon Press, Oxford. 1330 pp.

    Browne, F.G. (1980a) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species, I. Kontyû, 48, 370–379.

    Browne, F.G. (1980b) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species, II. Kontyû, 48, 380–389.

    Browne, F.G. (1980c) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species, IV. Kontyû, 48, 490–500.

    Browne, F.G. (1981) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species, V. Kontyû, 49, 125–136.

    Browne, F.G. (1983) Three new species of Scolytidae (Coleoptera) from southern China. Entomologist’s Monthly Magazine, 119, 33–34.

    Browne, F.G. (1984) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species, VIII. Kontyû, 52, 150–158.

    Browne, F.G. (1986) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species, XV. Kontyû, 54, 661–671.

    Chapuis, F. (1869) Synopsis des Scolytides (Prodrome d’un travail monographique). J.Desoer, Liège. 61 pp.

    Chapuis, F. (1875) In: Chapuis, F. & Eichhoff, W.J. Scolytides recueillis au Japon par M. G.Lewis. Annales de la Société entomologique de Belgique, 18, 195–204.

    Chararas, C. (1962) Étude biologique des scolytides des conifères. Encyclopédie Entomologique, Ser.A–38. Lechevalier, Paris. 556 pp.

    Choo, H.Y. (1983) Taxonomic studies on the Platypodidae and Scolytidae (Coleoptera) from Korea. Ph.D. dissertation, Seoul National University.

    Choo, H.Y. & Woo, K.S. (1985) Taxonomic studies on the Platypodidae and Scolytidae (Coleoptera) from Korea. I. Platypodidae, Slolytoplatypinae [sic] and Scolytinae. Insecta Koreana, 5, 31–47.

    Chu, Y.I. & Hsiao, M.L. (1981) On the so-called “Shiraki collection”. Chinese Journal of Entomology, 1, 26–32. (In Chinese)

    Damon, A. (2000) A review of the biology and control of the coffee berry borer, Hypothenemus hampei (Coleoptera: Scolytidae). Bulletin of Entomological Research, 90, 453–465.

    Davis, M.A. & Dute, R.R. (1997) Fungal associates of the Asian ambrosia beetle, Xylosandrus crassiusculus. Southern Nursery Association Research Conference, 42, 106–112.

    Dole, S.A. & Cognato, A.I. (2010) Monographic revision of the genus Xylosandrus Reitter (Coleoptera: Curculionidae: Scolytinae). Proceedings of the California Academy of Sciences, (in press).

    Eggers, H. (1908) Fünf neue Borkenkäfer. Entomologische Blätter, 4, 214–217.

    Eggers, H. (1914) Bemerkungen zu Reitters Borkenkäferbestimmungstabellen. 2. Auflage. Entomologische Blätter, 10, 183–189.

    Eggers, H. (1920) 60 neue Borkenkåfer (Ipidae) aus Afrika, nebst zehn neue Gattungen, zwei Abarten. Entomologische Blätter, 16, 33–45.

    Eggers, H. (1923) Neue indomalayische Borkenkäfer (Ipidae). Zoologische Mededeelingen, 7, 129–220.

    Eggers, H. (1926) Japanische Borkenkäfer I. Entomologische Blätter, 22, 133–138,145–148.

    Eggers, H. (1927) Neue indomalayische Borkenkaefer (Ipidae). II. Nachtrag. Philippine Journal of Science, 33, 67–108.

    Eggers, H. (1930) Neue Xyleborus-Arten (Col., Scolytidae) aus Indien. Indian Forest Records, Entomology Series, 14, 177–208.

    Eggers, H. (1933) Borkenkäfer (Ipidae, Col.) aus China. Entomologisches Nachrichtenblatt, 7, 97–102.

    Eggers, H. (1936) Neue Borkenkäfer (Scolytidae, Col.) aus India. Annals and Magazine of Natural History, ser.10, 17, 626–636.

    Eggers, H. (1939a) Japanische Borkenkäfer II. Arbeiten über morphologische und taxonomische Entomologie aus Berlin-Dahlem, 6, 114–123.

    Eggers, H. (1939b) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Ipidae gesammelt von René Malaise. Arkiv för Zoologi, 31A(4), 1–14.

    Eggers, H. (1940) Neue indomalayische Borkenkäfer (Ipidae). III. Nachtrag. Tijdschrift voor Entomologie, 83, 132–154.

    Eggers, H. (1941) Neue Borkenkäfer (Ipidae, Col.) aus China. Entomologische Blätter, 37, 222–226.

    Eichhoff, W.J. (1868) Neue amerikanische Borkenkäfer-Gattungen und Arten. Zweites Stück. Berliner Entomologische Zeitschrift, 12, 145–152.

    Eichhoff, W.J. (1872) Neue exotische Tomiciden-Arten. Berliner Entomologische Zeitschrift, 15, 131–137.

    Eichhoff, W.J. (1875) [Tomicides]. In: Chapuis, F. & Eichhoff, W.J. Scolytides recueillis au Japon par M. G.Lewis. Annales de la Société entomologique de Belgique, 18, 195–204.

    Eichhoff, W.J. (1877) Japanische Scolytidae. Deutsche Entomologische Zeitschrift, 21, 117–128.

    Eichhoff, W.J. (1878a) Neue oder noch unbeschriebene Tomicinen. Entomologische Zeitung, Stettin, 39, 383–392.

    Eichhoff, W.J. (1878b) Ratio, descriptio, emendatio eorum Tomicinorum qui sunt in Dr.medic. Chapuisii et autoris ipsiae collectionibus et quos praeterea recognovit scriptor. Mémoires de la Société Royale de Sciences de Liège, 2e.Sér., 8, 1–531 + pl. I–V.

    Entwhistle, P.F. (1972) Pests of cocoa. Longman, London. 779 pp.

    Fabricius, J.C. (1775) Systema Entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Officina Libraria Kortii, Flensburgi et Lipsiae. 832 pp.

    Fabricius, J.C. (1801) Systema eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Vol.2. Bibliopoli Academici Novi, Kiliae. 687 pp.

    Ferrari, J.A. (1867) Die Forst- und Baumzuchtschädlichen Borkenkäfer (Tomicides Lac.) aus der Familie der Holzverderber (Scolytides Lac.), mit besonderer Berüchsichtigung vorzüglich der europäischen Formen, und der Sammling des k.k. zoologischen Kabinetes in Wien. Carl Gerold’s Sohn, Wien. 96 pp.

    Fischer, M. (1954) Untersuchungen über den kleinen Holzbohrer (Xyleborinus saxeseni Ratz.) Pflanzenschutzberichte, 12, 137–180.

    Fraedrich, S.W., Harrington, T.C., Rabaglia, R.J., Ulyshen, M.D., Mayfield III, A.E., Hanula, J.L., Eickwort, J.M. & Miller, D.R. (2008) A fungal symbiont of the redbay ambrosia beetle causes a lethal wilt in redbay and other Lauraceae in the southeastern United States. Plant Disease, 92, 215–224.

    Francke-Grosmann, H. (1966) Über Symbiosen von xylo-mycetophagen und phloeophagen Scolytoidea mit holzbewohnenden Pilzen. Material und Organismen, Beiheft 1, 503–522.

    Gebhardt, H., Weiss, M. & Oberwinkler, F. (2005) Dryadomyces amasae: a nutritional fungus associated with ambrosia beetles of the genus Amasa (Coleoptera: Curculionidae, Scolytinae). Mycological Research, 109, 687–696.

    Gibb, H., Hjältén, J., Ball, J.P., Atlegrim, O., Petterson, R.B., Hliszczański, J., Johansson, T. & Danell, K. (2006a) Effects of landscape composition and substrate availability on saproxylic beetles in boreal forests: a study using experimental logs for monitoring assemblages. Ecography, 29, 191–204.

    Gibb, H., Tetterson, R.B., Hjältén, J., Hilszczański, J., Ball, J.P., Johansson, T., Atlegrim, O. & Danell, K. (2006b) Conservation-oriented forestry and early successional saproxylic beetles: responses of functional groups to manipulated dead wood substrates. Biological Conservation, 129, 437–450.

    Gilbert, M., Nageleisen, L.-M., Franklin, A. & Grégoire, J.-C. (2005) Post-storm surveys reveal large scale spatial patterns and influences of site factors, forest structure and diversity in endemic bark-beetle populations. Landscape Ecology, 20, 35–49.

    Gray, B. (1972) Observations on Poecilips cardamomi (Schaufuss), the second species of Scolytidae to be found in bracken fern (Col.). Entomologisk Tidskrift, 74, 113–119.

    Gray, B. & Wylie, F.R. (1974) Forest tree and timber insect pests in Papua New Guinea. II. Pacific Insects, 16, 67–115.

    Haack, R. (2006) Exotic bark- and wood-boring Coleoptera in the United States: recent establishments and interceptions. Canadian Journal of Forest Research, 36, 269–288.

    Hagedorn, M. (1904) Enumeratio Scolytidarum e Sikkim et Japan naturum Musei historico-naturalis Parisiorum, quas Dominus J.Harmand annis 1890 et 1901 collegit descriptionibus specierum novarum adjectis. Bulletin du Muséum d’Histoire Naturelle, Paris, 1904, 122–126.

    Hanula, J.L., Mayfield III, A.E., Fraedrich, S.W. & Rabaglia, R.J. (2008) Biology and host asssociations of redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae), exotic vector of laurel wilt killing redbay trees in the Southeastern United States. Journal of Economic Entomology, 101, 1276–1286.

    Harrington, T.C. (2005) Ecology and evolution of mycophagous bark beetles and their fungal partners. In: Vega, F.E. & Blackwell, M. (Eds.) Insect-Fungal Associations. Ecology and Evolution. Oxford University Press, Oxford. pp. 257–291.

    Herfs, A. (1949) Studien an dem Steinnussborkenkäfer, Coccotrypes tanganus Eggers. I. Der Einfluss der Aussenfaktoren auf die Entwicklung. Höfchen-Briefe für Wissenschaft und Praxis, 2, 22–49.

    Herfs, A. (1950) Studien an dem Steinnussborkenkäfer, Coccotrypes tanganus Eggers. II. Die Soziologie von Coccotrypes tanganus. Höfchen-Briefe für Wissenschaft und Praxis, 3, 3–31.

    Herfs, A. (1958) Über den Steinnussborkenkäfer Coccotrypes dactyliperda F. Anzeiger für Schädlingskunde, 32, 1–4.

    Hoebeke, E.R. & Rabaglia, R.J. (2008) Xyleborus seriatus Blandford (Coleoptera: Curculionidae: Scolytinae), an Asian ambrosia beetle new to North America. Proceedings of the Entomological Society of Washington, 110, 470–476.

    Hijii, N., Kajimura, H. & Nishibe, Y. (1991) A note on the discoloration and fungal infiltration processes on wood tissues surrounding the gallery system of scolytid beetles. Bulletin of the Nagoya University Forest, 11, 31–38.

    Hopkins, A.D. (1915) Classification of the Cryphalinae, with decsriptions of new genera and species. United States Department of Agriculture, Office of the Secretary, Report, 99, 1–75 + pl. I–IV.

    Hornung, E.G. (1842) Über einige in den Betelnüssen vorkommende Käfer. Entomologische Zeitung, Stettin, 3, 115–117.

    Hosking, G.P. (1973) Xyleborus saxeseni, its biology and flight behaviour in New Zealand. New Zealand Journal of Forestry Science, 3, 37–53.

    Hsieh, C.F., Sun, I.F. & Yang, C.C. (2000) Species composition and vegetation pattern of a lowland rain forest at the Nanjenshan LTER site, southern Taiwan. Taiwania, 45, 107–119.

    Huang, S.H., Beaver, R., Shih, H.T. & Cheng, C.H. (2003) Insects of the Scolytidae and Platypodidae on Brownea capitella and Prunue [sic] mume in Taiwan. Formosan Entomologist, 23, 101–111 (In Chinese, English summary)

    Hulcr, J. (2010) Taxonomical changes in paleotropical Xyleborina (Coleoptera: Curculionidae: Scolytinae). Zookeys (in press).

    Hulcr, J. & Cognato, A.I. (2009) Three new genera of oriental Xyleborina (Coleoptera: Curculionidae: Scolytinae). Zootaxa, 2204, 19–36.

    Hulcr, J. & Cognato, A.I. (2010) New genera of Palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae) based on congruence between morphological and molecular characters. Zootaxa (in review).

    Hulcr, J., Dole, S.A., Beaver, R.A. & Cognato, A.I. (2007a) Cladistic review of generic taxonomic characters in Xyleborina (Coleoptera: Curculionidae: Scolytinae). Systematic Entomology, 32, 568–584.

    Hulcr, J., Mogia, M., Isua, B. & Novotny, V. (2007b) Host specificity of ambrosia and bark beetles (Col. Curculionidae: Scolytinae and Platypodinae) in a New Guinea rainforest. Ecological Entomology, 32, 762–772.

    Ito, M., Kajimura, H., Hamaguchi, K., Araya, K. & Lakatos, F. (2008) Genetic structure of Japanese populations of an ambrosia beetle, Xylosandrus germanus (Curculionidae: Scolytinae). Entomological Science, 11, 375–383.

    Johansson, T., Gibb, H., Hilszczański, J., Petterson, R.B., Hjältén, J, Atlegrim, O., Ball, J.P. & Danell, K. (2006) Conservation-oriented manipulations of coarse woody debris affect its value as a habitat for spruce-infesting bark and ambrosia beetles (Coleoptera: Scolytinae) in northern Sweden. Canadian Journal of Forest Research, 36, 174–185.

    Jordal, B.H., Normark, B.B. & Farrell, B.D. (2000) Evolutionary radiation of an inbreeding haplodiploid beetle lineage (Curculionidae, Scolytinae). Biological Journal of the Linnaean Society, 71, 483–499.

    Jordal, B.H., Beaver, R.A. & Kirkendall, L.R. (2001) Breaking taboos in the tropics: incest promotes colonization by wood-boring beetles. Global Ecology and Biogeography, 10, 345–357.

    Jordal, B.H. & Kirkendall, L.R. (1998) Ecological relationships of a guild of tropical beetles breeding in Cecropia petioles in Costa Rica. Journal of Tropical Ecology, 14, 153–176.

    Jordal, B.H., Normark, B.B., Farrell, B.D. & Kirkendall, L.R. (2002) Extraordinary haplotype diversity in haplodiploid inbreeders: phylogenetics and evolution of the bark beetle genus Coccotrypes. Molecular Phylogenetics and Evolution, 23, 171–188.

    Kajimura, H. & Hijii, N. (1992) Dynamics of the fungal symbiosis in the gallery system asnd the mycangia of the ambrosia beetle, Xylosandrus mutilatus (Blandford) (Coleoptera: Scolytidae) in relation to its life history. Ecological Research, 7, 107–117.

    Kajimura, H. & Hijii, N. (1994) Reproduction and resource utilization of the ambrosia beetle, Xylosandrus mutilatus, in field and experimental populations. Entomologia Experimentalis et Applicata, 71, 121–132.

    Kalshoven, L.G.E. (1959) Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. Second part. Tijdschrift voor Entomologie, 102, 135–173. + pl. 15–22.

    Kalshoven, L.G.E. (1961) A study of the twig borer, Xylosandrus morigerus Blandford, mainly based on observations in Java. Tijdschrift voor Entomologie, 104, 93–110.

    Kalshoven, L.G.E. (1962) Note on the habits of Xyleborus destruens Bldf., the near-primary borer of teak trees on Java. Entomologische Berichten, 22, 7–18.

    Kalshoven, L.G.E. (1964) The occurrence of Xyleborus perforans (Woll.) and X.similis in Java (Coleoptera, Scolytidae). Beaufortia, 11, 131–142.

    Kinuura, H. (1995) Symbiotic fungi associated with bark beetles. Japan Agricultural Research Quarterly, 29, 57–63.

    Kinuura, H. & Hijii, N. (1991) Life history and reproduction of the ambrosia beetle, Scolytoplatypus mikado Blandford (Coleoptera, Scolytidae). Japanese Journal of Entomology, 59, 763–773.

    Kinuura, H., Hijii, N. & Kanamitsu, K. (1991) Symbiotic fungi asssociated with the ambrosia beetle,Scolytoplatypus mikado Blandford (Coleoptera: Scolytidae)–succession of the flora and fungal phases in the gallery system and the mycangium in relation to the developmental stages of the beetle. Journal of the Japanese Forestry Society, 73, 197–205.

    Kinuura, H., Ohya, E, Makihara, H, Nagaki, A. & Fujioka, H. (1999) Infection rate of Monochamus alternatus Hope (Coleoptera: Cerambycidae) by an entomogenous fungus through mass-release of vector beetles Cryphalus fulvus Niijima (Coleoptera: Scolytidae) using an improved application device. Journal of the Japanese Forestry Society, 81, 17–21. (In Japanese, English summary)

    Kirisits, T. (2004) Fungal associates of European bark beetles with special emphasis on the Ophiostomatoid fungi. In: Lieutier, F., Day, K.R., Battisti, A, Grégoire, J-C. & Evans, H.F. (Eds) Bark and wood boring insects in living trees in Europe, a synthesis. Kluwer Academic, Dordecht. pp. 181–235.

    Kirkendall, L.R. (1993) Ecology and evolution of biased sex ratios in bark and ambrosia beetles. In: Wrensch, D.L. & Ebbert, M.A. (Eds.) Evolution and diversity of sex ratio in insects and mites. Chapman & Hall, New York. pp. 235–345.

    Kirkendall, L.R., Faccoli, M. & Ye, H. (2008) Description of the Yunnan shoot borer, Tomicus yunnanensis Kirkendall & Faccoli sp.n. (Curculionidae, Scolytinae), an unusually aggressive pine shoot beetle from southern China, with a key to the species of Tomicus. Zootaxa, 1819, 25–39.

    Klepzig, K.D. & Six, D.L. (2004) Bark beetle-fungal symbiosis: context-dependency in complex associations. Symbiosis, 37, 189–205.

    Koch, F.H. & Smith, W.D. (2008) Spatio-temporal analysis of Xyleborus glabratus (Coleoptera: Curculionidae: Scolytinae) invasion in eastern U.S.forests. Environmental Entomology, 37, 442–452.

    Kolařík, M., Kubátová, A., Hulcr, J. & Pažoutova, S. (2008) Geosmithia fungi are highly diverse and consistent bark beetle associates: evidence from their community structure in temperate Europe. Microbial Ecology, 55, 65–80.

    Kolk, A. & Starzyk, J.R. (1996) Atlas Skodliwych owadów lesnych. (The Atlas of Forest Insect Pests). Multico, Warszawa. 705pp. (In Polish, partial English translation by L.Sukatova et al. available from http://www.forestpests.org/poland/ ) (accessed 21 November 2009)

    Kovach, J. & Gorsuch, C.S. (1985) Survey of ambrosia beetle species infesting South Carolina peach orchards and a taxonomic key for the most common species. Journal of Agricultural Entomology, 2, 238–247.

    Krivolutskaya, G.O. (1973) Entomofauna of the Kuril Islands: Principal Features and Origin. Izdatel’stvo Nauka, Leningrad Division. 315 pp. (In Russian. English translation by E.B.Urdang available from http://www.washington.edu/burkemuseum/collections/ichthyology/okhotskia/ikip/Results/publications/entobook/title.htm )(accessed 18 November 2009).

    Kurenzov, A.I. (1941) Bark-beetles of the Far East, USSR. Akad.Nauk, USSR, Moscow-Leningrad. 234 pp. (In Russian)

    Kuschel, G. (1995) A phylogenetic classification of Curculionoidea to families and subfamilies. Memoirs of the Entomological Society of Washington, 14, 5–33.

    Långström, B. (1983) Life cycle and shoot feeding of the pine shoot beetles. Studia Forestalia Suecica, 163, 1–29.

    Lea, A.M. (1910) On Australian and Tasmanian Coleoptera, with descriptions of new species. Part I. Proceedings of the Royal Society of Victoria, 22 (N.S.), 113–152.

    Lee, C.F. & Sato, M. (2006) The Helotidae of Taiwan (Coleoptera: Cucujoidea). Zoological Studies, 45, 529–552.

    Le Pelley, R.H. (1968) Pests of coffee. Longmans, London and Harlow. 590 pp.

    Linnaeus, C. (1758) Systema Naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Ed. 10, T.1. Laurentii Salvii, Holmiae. 823 pp.

    Löbl, I. & Smetana, A. (Eds.). (2003) Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup. 819 pp.

    Lombardero, M.J., Vázquez-Mejuto, P. & Ayres, M.P. (2008) Role of plant enemies in the forestry of indigenous vs. nonindigenous pines. Ecological Applications, 18, 1171–1181.

    Lou, J., Zhang, T.L., Zhang, M.H. & Li, G.W. (2006) Occurrence and damage of bark beetles in Laotudingzi Nature Reserve. Forest Pest and Disease, 25, 32–35. (In Chinese)

    Lu, M., Miller, D.R. & Sun, J.H. (2007) Cross-attraction between an exotic and a native bark beetle: a novel invasion mechanism. PLoS ONE, 2(12), e1302.

    Maiti, P.K. & Saha, N. (1986) Contributions to the knowledge of the bark and timber beetles (Scolytidae: Coleoptera) of the Andaman and Nicobar islands. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 86, 1–182.

    Maiti, P.K. & Saha, N. (2004) Fauna of India and the adjacent countries. Scolytidae: Coleoptera (Bark- and Ambrosia-Beetles) Volume 1 (Part 1). Introduction and Tribe Xyleborini. Zoological Survey of India, Kolkata. 268 pp.

    Mandelshtam, M.Y., Petrov, A.V., Barclay, M.V.L., Knížek, M. & Beaver, R.A. (2008) Taxonomic changes in Scolytinae (Coleoptera: Curculionidae) from Eastern Asia. Russian Entomological Journal, 16, 459–464.

    Markalas, S. & Kalapanida, M. (1997) Flight pattern of some Scolytidae attracted to flight barrier traps baited with ethanol in an oak forest in Greece. Anzeiger für Schädlingskunde, Pflanzenschutz, Umweltschutz, 70, 55–57.

    Martikainen, P., Siitonen, J., Kaila, L. & Punttila, P. (1996) Intensity of forest management and bark beetles in non-epidemic conditions: a comparison between Finnish and Russian Karelia. Journal of Applied Entomology, 120, 257–264.

    Marvaldi, A., Sequeira, A.S., O’Brien, C.W. & Farrell, B.D. (2002) Molecular and morphological phylogenetics of weevils (Coleoptera: Curculionoidea): Do niche shifts accompany diversification? Systematic Biology, 51, 761–785.

    Masood, A., Saeed, S., Sajjad, A. & Ali, M. (2009) Life cycle and biology of mango bark beetle, Hypocryphalus mangiferae (Stebbing), a possible vector of mango sudden death disease in Pakistan. Pakistan Journal of Zoology, 4, 281–288.

    Michalski, J. (1973) Revision of the Palearctic species of the genus Scolytus Geoffroy (Coleoptera, Scolytidae). Panstwowe Wydawnictwo Naukowe, Warszawa. 214 pp. + 49 pl.

    Miller, D.R. & Rabaglia, R.B. (2009) Ethanol and (−)-α-pinene: attractant kairomones for bark and ambrosia beetles in the southeastern US. Journal of Chemical Ecology, 35, 435–448.

    Miwa, Y. (1931) A systematic catalogue of Formosan Coleoptera. Report of the Government Research Institute, Department of Agriculture, Taihoku, Formosa. No. 55. 359 pp.

    Motschulsky, V. von (1863) Essai d’un catalogue des insectes de l’île Ceylan. (Suite). Bulletin de la Société Impériale des Naturalistes de Moscou, 36, 421–532.

    Motschulsky, V. von (1866) Essai d’un catalogue des insectes de l’île Ceylan. Bulletin de la Société Impériale des Naturalistes de Moscou, 39, 393–446.

    Murayama, J. (1930) Révision des familles des Ipides et des Platypides de Corée. Journal of Chosen Natural History Society, 11, 1–34 + Pl. I–II.

    Murayama, J. (1934a) On the Ipidae (Coleoptera) from Formosa with special reference to their food plants. Journal of the Society of Tropical Agriculture, 6, 505–512.

    Murayama, J. (1934b) Notes on the Ipidae (Coleoptera) from Kiushu. Annotationes Zoologicae Japonenses, 14, 287–300.

    Murayama, J. (1936) Notes sur les scolytides (Coléoptères) de Honshû et Kiushû, Japon. Tenthredo, 1, 121–149.

    Murayama, J. (1950a) Nouvelles espèces de Scolytidae (Coléopt.) de l’îsle Shikoku. Insecta Matsumurana, 17, 61–64.

    Murayama, J. (1950b) A new genus and some new species of Scolytidae from Japan (Coleoptera). Transactions of the Shikoku Entomological Society, 1, 49–53.

    Murayama, J. (1953) The insect fauna of Mt. Ishizuchi and Omogo Valley, Iyo, Japan. The Scolytidae and Platypodidae (Coleoptera). Transactions of the Shikoku Entomological Society, 3, 144–165.

    Murayama, J. (1954) Scolytid-fauna of the northern half of Honshu with a distribution table of all the scolytid-species described from Japan. Bulletin of the Faculty of Agriculture, Yamaguti University, 5, 149–212.

    Murayama, J. (1955) Supplementary notes on the scolytid-fauna of Japan. Bulletin of the Faculty of Agriculture, Yamaguti University, 6, 81–106 + pl.3–4.

    Murayama, J. (1957) Studies in the scolytid fauna of the Northern half of the Far East. III. Dryocoetinae. Bulletin of the Faculty of Agriculture, Yamaguti University, 8, 587–632.

    Murayama, J.J. (1961) Scolytid beetles from Niigata Prefecture, Japan. Akitu, 10, 23–32.

    Nakashima, T., Otomo, T., Owada, Y. & Iizuka, T. (1992) SEM observations on growing conditions of the fungi in the galleries of several ambrosia beetles (Coleoptera: Scolytidae and Platypodidae). Journal of the Faculty of Agriculture, Hokkaido University, 65, 239–273.

    Niijima, Y. (1942) Die japanischen Phloeosinus-Arten (Coleoptera, Ipidae) und ihre Frasspflanzen. Transactions of the Sapporo Natural History Society, 17, 69–76 + pl.I. [In earlier papers the author spelled his surname as Niisima. We have used the spelling given in the individual paper.]

    Niisima, Y. (1908) Über die japanischen Cryphalus-Arten. Verhandlungen der k.k. zoologisch-botanischen Gesellschaft, Wien, 58, 89–92.

    Niisima, Y. (1909) Die Scolytiden Hokkaidos unter Berücksichtigung ihrer Bedeutung für Forstschaden. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo, 3, 109–179 + Pl. III–IX.

    Nobuchi, A. (1959) Some species of Scolytidae from Ryukyu Islands. Bulletin of the Government Forest Experiment Station, 116, 21–26.

    Nobuchi, A. (1966) Studies on Scolytidae, VI. Bulletin of the Government Forest Experiment Station, 185, 51–56.

    Nobuchi, A. (1967) Formosan Scolytoidea (Coleoptera). Bulletin of the Government Forest Experiment Station, 207, 11–30 + pl. I.

    Nobuchi, A. (1973) Studies on Scolytidae: XI. Scolytus Geoffroy of Japan (Coleoptera). Bulletin of the Government Forest Experiment Station, 258, 13–28.

    Nobuchi, A. (1973) Studies on Scolytidae: XII. The bark beetles of the tribe Ipini in Japan (Coleoptera). Bulletin of the Government Forest Experiment Station, 266, 33–60.

    Nobuchi, A. (1975) Studies on Scolytidae XIII. Twenty-one new species of Cryphalini from Japan (Coleoptera). Bulletin of the Government Forest Experiment Station, 277, 41–60.

    Nobuchi, A. (1980) Studies on Scolytidae, XIX. Formosan Platypodidae and Scolytidae collected by Dr. Yan I-Chu (Coleoptera). Entomological Review, Japan, 34, 93–97.

    Nobuchi, A. (1981a) Studies on Scolytidae (Coleoptera) XXI. Three new genera and species from Japan. Kontyû, 49, 12–18.

    Nobuchi, A. (1981b) Studies on Scolytidae (Coleoptera) XXII. Six new species and two new females of the genus Xyleborus Erichson [sic] from Japan. Kontyû, 49, 143–154.

    Norris, D.M. (1976) Chemical interdependencies among Xyleborus spp. ambrosia beetles and their symbiotic microbes. Material und Organismen, Beiheft 3, 479–488.

    Ohno, S. (1990) The Scolytidae and Platypodidae (Coleoptera) from Borneo found in logs at Nagoya port. 1. Research Bulletin of the Plant Protection Service, Japan, 26, 83–94.

    Ohno, S., Yoshioka, K., Yoneyama, K. & Nakazawa, H. (1988) The Scolytidae and Platypodidae (Coleoptera) from Solomon Islands found in logs at Nagoya port. 1. Research Bulletin of the Plant Protection Service, Japan, 24, 91–95.

    Ohno, S, Yoshioka, K., Uchida, N., Yoneyama, K. & Tsukamoto, K. (1989) The Scolytidae and Platypodidae (Coleoptera) from the Bismarck Archipelago found in logs at Nagoya port. Research Bulletin of the Plant Protection Service, Japan, 25, 59–69.

    Ohtaka, N., Masuya, H., Kaneko, S. & Yamaoka, Y. (2002) Two Ophiostoma species associated with bark beetles in wave-regenerated Abies veitchii forests in Japan. Mycoscience, 43, 151–157.

    Oliver, J.B. & Mannion, C.M. (2001) Ambrosia beetle (Coleoptera: Scolytidae) species attacking chestnut and captured in ethanol-baited traps in middle Tennessee. Environmental Entomology, 30, 909–918.

    Olivier, A.G. (1795) Scolyte: Scolytus. In: Entomologie ou histoire naturelle des insectes, avec leur caractères génériques et spécifiques, leur descriptions, leur synonymie, et leur figure enluminée. Coléoptères. Vol. 4, pt. 78. Lanneau, Paris. pp. 1–14.

    Panzer, G.W.F. (1791) Beschreibung eines noch unbekannten sehr kleinen Kapuzkäfers. Naturforscher, 25, 35–38.

    Peer, K. & Taborsky, M. (2004) Female ambrosia beetles adjust their offspring sex ratio according to outbreeding opportunities for their sons. Journal of Evolutionary Biology, 17, 257–264.

    Peer, K. & Taborsky, M. (2005) Outbreeding depression but no inbreeding depression in haplodiploid ambrosia beetles with regular sib mating. Evolution, 59, 317–323.

    Peer, K. & Taborsky, M. (2007) Delayed dispersal as a potential route to cooperative breeding in ambrosia beetles. Behavioural Ecology and Sociobiology, 61, 729–739.

    Peltonen, M. & Helioväara, K. (1999) Attack density and breeding success of bark beetles (Coleoptera, Scolytidae) at different distances from forest-clearcut edge. Agricultural and Forest Entomology, 1, 237–242.

    Rabaglia, R.J, Dole, S.A. & Cognato, A.I. (2006) Review of American Xyleborina (Coleoptera: Curculionidae: Scolytinae) occurring North of Mexico, with an illustrated key. Annals of the Entomological Society of America, 99, 1034–1056.

    Ratzeburg, J.T.C. (1837) Die Forst-Insekten oder Abbildungen und Beschreibung der in den Waldern Preussens und der Nachbarstaten als schädlich oder nützlich bekannt gewordenen Insekten. Erster Theil. Die Käfer. Nicolai, Berlin. 202 pp. + 22 pl.

    Reitter, E. (1913) Bestimmungs-tabelle der Borkenkäfer (Scolytidae) aus Europa und den angrenzenden Ländern. Wiener Entomologische Zeitung, 32(Beiheft), 1–116.

    Roberts, H. (1976) Observations on the biology of some tropical rain forest Scolytidae (Coleoptera) from Fiji. I. Subfamilies Hylesininae, Ipinae (excluding Xyleborini). Bulletin of Entomological Research, 66, 373–388.

    Roeper, R.A., Treeful, L.M., Foote, R.A. & Bunce, M.A. (1980a) In vitro culture of the ambrosia beetle Xyleborus affinis (Coleoptera: Scolytidae). Great Lakes Entomologist, 13, 33–35.

    Roeper, R.A., Treeful, L.M., O’Brien, K.M., Foote, R.A. & Bunce, M.A. (1980b) Life history of the ambrosia beetle Xyleborus affinis (Coleoptera: Scolytidae) from in vitro culture. Great Lakes Entomologist, 13, 141–143.

    Saha, N. & Maiti, P.K. (1996) Insecta: Coleoptera: Scolytidae. In: Director, Zoological Survey of India (Ed.) Fauna of West Bengal, Part-6b (Insecta: Coleoptera). Zoological Survey of India, Calcutta. pp. 775–866.

    Sampson, F.W. (1919) Notes on Platypodidae and Scolytidae collected by Mr. G. E. Bryant and others. Annals and Magazine of Natural History, ser.9, 4, 105–114.

    Sampson, F.W. (1922) Previously undescribed Scolytidae and Platypodidae from the Indian area. Annals and Magazine of Natural History, ser.9, 10, 145–152.

    Sasakawa, M. (1975) Developmental stages, behavior and re-attack pattern of the minute pine bark beetle, Taenioglyptes fulvus (Niijima), under laboratory conditions. Japanese Journal of Applied Entomology and Zoology, 19, 237–242. (In Japanese, English summary)

    Sasakawa, M. & Negishi, T. (1973) Laboratory and field studies on behaviouristic response of the minute pine bark beetle, Cryphalus fulvus Niijima (Coleoptera: Scolytidae) to the female pheromone. Applied Entomology and Zoology, 8, 143–156.

    Sasakawa, M. & Katayama, J. (1975) The effeects of attack density on brood production in the minute pine bark beetle, Taenioglyptes fulvus (Niijima) (Coleoptera: Scolytidae). Applied Entomology and Zoology, 10, 172–184.

    Sasakawa, M. & Kawaguchi, Y. (1987) Initial attack of the minute pine bark beetle, Cryphalus fulvus Niijima (Coleoptera: Scolytidae). Scientific Reports of the Kyoto Prefectural University, Agriculture, 39, 12–19.

    Sasakawa, M. & Sasakawa, T. (1981) Stridulatory organs of the minute pine bark beetle, Cryphalus fulvus Niijima (Coleoptera, Scolytidae), and the role of male sounds in the aggregation behavior. Kontyû, 49, 461–469.

    Sasakawa, M., Ohta, F. & Negishi, T. (1976) Relationship between response to the aggregation pheromone trap of the minute pine bark beetle, Taenioglyptes fulvus (Niijima) (Coleoptera: Scolytidae) and regulation of the entrance hole density. Bulletin of the Kyoto University Prefectural University Forests, 20, 42–48. (In Japanese, English summary)

    Sauvard, D. (2004) General biology of bark beetles. In: Lieutier, F., Day, K.R., Battisti, A, Grégoire, J-C. & Evans, H.F. (Eds.) Bark and wood boring insects in living trees in Europe, a synthesis. Kluwer Academic, Dordecht. pp. 63–88.

    Schaufuss, C.F.C. (1905) Borkenkäferstudien. II. 4. Über einige Schädlinge von Kulturgewächsen. Insektenbörse, 22, 8.

    Schedl, K.E. (1935) Scolytidae and Platypodidae. New species from the Philippine islands and Formosa. Philippine Journal of Science, 57, 479–489.

    Schedl, K.E. (1942a) Forschungsberichte zur Scolytiden-Fauna der Malayischen Halbinsel, V. 80 Beitrag. Kolonialforstliche Mitteilungen 5, 169–218.

    Schedl, K.E. (1942b) Neue Scolytidae aur Java. 76 Beitrag. Tijdschrift voor Entomologie, 85, 1–49.

    Schedl, K.E. (1946) Bestimmungstabellen der palaearktischen Borkenkäfer. Teil I. Die Gattung Crypturgus Er. Zentralblatt für das Gesamtgebiet der Entomologie, 1, 1–15.

    Schedl, K.E. (1949) Tropical seed beetles of the genus Coccotrypes Eichhoff. 99. Contribution to the morphology and taxonomy of the Scolytoidea. Tijdschrift voor Entomologie, 91, 113–120.

    Schedl, K.E. (1951) Fauna Indomalayensis, I. Tijdschrift voor Entomologie, 93, 41–98.

    Schedl, K.E. (1952a) Formosan Scolytoidea, I. 111. Contribution to the morphology and taxonomy of the Scolytoidea. Philippine Journal of Science, 81, 61–65.

    Schedl, K.E. (1952b) Zur Synonymie der Borkenkäfer, I. 121 Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Blätter, 47/48, 158–164.

    Schedl, K.E. (1953a) Fauna Sinensis, I. 120 Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Blätter, 49, 22–30.

    Schedl, K.E. (1953b) Fauna Indomalayensis, III. Contribution 133 to the morphology and taxonomy of the Scolytoidea. Annals and Magazine of Natural History, ser.12, 6, 288–304.

    Schedl, K.E. (1958) Zur Synonymie der Borkenkäfer. II. 159. Beitrag zur Morphologie und Systematik der Scolytoidea. Tijdschrift voor Entomologie, 101, 141–155.

    Schedl, K.E. (1961) Scolytidae und Platypodidae Afrikas. Band I. Unterfamilie Hylesininae (Fortsetzung). Revista de Entomologia de Moçambique, 4, 335–742.

    Schedl, K.E. (1962) Zur Synonymie der Borkenkäfer, VI. 203. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Blätter, 58, 201–211.

    Schedl, K.E. (1963) Scolytidae und Platypodidae Afrikas. Band II. Familie Scolytidae (Fortsetzung), Unterfamilie Ipinae (Fortsetzung). Revista de Entomologia de Moçambique, 5, 1–593.

    Schedl, K.E. (1966a) Pin-hole borers and bark-beetles (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports. 241. Contribution to the morphology and taxonomy of the Scolytoidea. Kontyû, 34, 29–43.

    Schedl, K.E. (1966b) Checklist of the Scolytidae and Platypodidae from the Philippine Islands. 196. Contribution to the morphology and taxonomy of the Scolytoidea. Entomologische Abhandlungen der Staatlichen Museum für Tierkunde in Dresden, 35, 1–122.

    Schedl, K.E. (1967a) Bark-beetles and pin-hole borers (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports, II. 254. Contribution to the morphology and taxonomy of the Scolytoidea. Kontyû, 35, 119–129.

    Schedl, K.E. (1967b) Zur Synonymie der Borkenkäfer XVI. 242. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologisk Tidskrift, 88, 147–163.

    Schedl, K.E. (1969a) Bark beetles and pinhole borers (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports III. 258. Contribution to the morphology and taxonomy of the Scolytoidea. Kontyû, 37, 202–219.

    Schedl, K.E. (1969b) Indian bark and timber beetles V. 217. Contribution to the morphology and taxonomy of the Scolytoidea. Oriental Insects, 3, 47–70.

    Schedl, K.E. (1970) Bark beetles and pinhole borers (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports IV. 274. Contribution to the morphology and taxonomy of the Scolytoidea. Kontyû, 38, 353–370.

    Schedl, K.E. (1973) New Scolytidae and Platypodidae from the Papuan subregion. 299. Contribution to the morphology and taxonomy of the Scolytoidea. Papua New Guinea Agricultural Journal, 24, 87–97.

    Schedl, K.E. (1975) Bark and timber beetles of the Oriental region. Oriental Insects, 6, 451–460.

    Schlyter, F., Zhang, Q.H., Anderson, P., Byers, J., Wadhams, L.J., Löfqvist, J. & Birgersson, G. (2000) Electrophysiological and behavioural responses of Tomicus piniperda and Tomicus minor (Coleoptera: Scolytidae) to non-host leaf and bark volatiles. Canadian Entomologist, 132, 965–981.

    Schröder, L.M. (1988) Attraction of the bark beetle Tomicus piniperda and some other bark- and wood-living beetles to the host volatiles alpha-pinene and ethanol. Entomologia Experimentalis et Applicata, 46, 203–210.

    Shiao, S.-F. (2006) Insect Museum Digital Archives Project. Department of Entomology, National Taiwan University, Taipei. Available from http://www.imdap.entomol.ntu.edu.tw/ (Accessed 20 November 2009)

    Shibata, E., Waguchi, Y. & Yoneda, Y. (1994) Seasonal flight of an ambrosia beetle, Xyleborus validus Eichhoff, in Japanese cedar stands. Applied Entomology and Zoology, 29, 593–594.

    Six, D.L. (2003) Bark beetle-fungus symbioses. In: Bourtzis, K. & Miller, T.A. (eds.) Insect Symbiosis. CRC Press, Boca Raton. pp. 97–114.

    Six, D.L., Stone, W.D., de Beer, Z.W. & Woolfolk, S.W. (2009) Ambrosiella beaveri, sp.nov., associated with an exotic ambrosia beetle, Xylosandrus mutilatus (Coleoptera: Curculionidae, Scolytinae), in Mississippi, USA. Antonie van Leeuwenhoek, 96, 17–29.

    Solheim, H. & Långström, B. (2001) Blue-stain fungi associated with Tomicus piniperda in Sweden and preliminary observations on their pathogenicity. Annales des Sciences Forestières, 48, 149–156.

    Speyer, E.R. (1923) Notes upon the habits of Ceylonese ambrosia beetles. Bulletin of Entomological Research, 14, 11–23.

    Sreedharan, K., Balakrishnan, M.M., Samuel, S.D. & Bhat, P.K. (1991) A note on the association of wood boring beetles and a fungus with the death of silver oak trees on coffee plantations. Journal of Coffee Research, 21, 145–148.

    Stauffer, C., Kirisits, T., Nussbaumer, C., Pavlin, R. & Wingfield, M.J. (2001) Phylogenetic relationships between the European and Asian eight-spined larch bark beetle populations (Coleoptera, Scolytidae) inferred from DNA sequence and fungal associates. European Journal of Entomology, 98, 99–105.

    Stebbing, E.P. (1909) On some undescribed Scolytidae of economic importance from the Indian region. II. Indian Forest Memoirs, Forest Zoology Series, 1(2), 13–32.

    Stebbing, E.P. (1914) Indian forest insects of economic importance. Coleoptera. Eyre & Spottiswoode, London. 648 pp. + 63 pl.

    Stone, W.D. & Nebeker, T.E. (2007) Distribution and seasonal abundance of Xylosandrus mutilatus. Journal of Entomological Science, 42, 409–412.

    Stone, W.D., Nebeker, T.E. & Gerard, P.D. (2007) Host plants of Xylosandrus mutilatus in Mississippi. Florida Entomologist, 90, 191–195

    Strohmeyer, H. (1912) H.Sauter’s Formosa-ausbeute, Ipidae und Platypodidae (Col.). Entomologische Mitteilungen, 1, 38–42.

    Suzuki, W. (2002) Notes on some Shiraki specimens of the families Eucnemidae and Elateridae (Coleoptera) preserved in the Taiwan Agricultural Research Institute, Taichung. Coleopterists News 137, 1–4 (In Japanese)

    Takagi, K. (1967) The storage organ of symbiotic fungus in the ambrosia beetle, Xyleborus rubricollis Eichhoff (Coleoptera: Scolytidae). Japanese Journal of Applied Entomology and Zoology, 2, 168–170.

    Tang, W.Q. (2000) Biological characteristics of Xyleborus mutilatus and its control. Journal of Zhejiang Forestry College, 17, 417–420. (In Chinese, English summary)

    Thompson, R.T. (1992) Observations on the morphology and classification of weevils (Coleoptera, Curculionoidea) with a key to major groups. Journal of Natural History 26, 835–891.

    Ueda, A. (1995) Field study of two spermathophagous [sic] scolytid beetles, Coccotrypes graniceps Eichhoff and C.cardamomi (Shaufuss) [sic], in acorns of Quercus myrsinaefolia Blume. In: Hain, F.P. et al. (Eds.) Behavior, population dynamics and control of forest insects. Proceedings IUFRO Joint Conference, Maui, Hawai’i, February 1994. pp. 610–626.

    Ueda, A. (1997) Brood development of an inbreeding spermatophagous scolytid beetle, Coccotrypes graniceps (Eichhoff) (Coleoptera: Scolytidae). Japanese Journal of Entomology, 65, 677–687.

    Väisänen. R. & Helioväara, K. (1994) Assessment of insect occurrence in boreal forests based on satellite imagery and field measurements. Acta Forestalia Fennica, 243, 1–39.

    Weber, B.C. & McPherson, J.E. (1983a) World list of host plants of Xylosandrus germanus (Coleoptera: Scolytidae). The Coleopterists Bulletin, 37, 114–134.

    Weber, B.C. & McPherson, J.E. (1983b) Life history of the ambrosia beetle Xylosandrus germanus (Coleoptera: Scolytidae). Annals of the Entomological Society of America, 76, 455–462.

    Westwood, J.O. (1836) Description of a minute coleopterous insect, forming the type of a new subgenus allied to Tomicus, with some observations upon the affinities of the Xylophaga. Transactions of the Entomological Society of London, 1, 34–36 + Pl. VII.

    Wollaston, T.V. (1857) Catalogue of the coleopterous insects of Madeira in the collection of the British Museum. Taylor & Francis, London. 234 pp.

    Wood, S.L. (1977) Introduced and exported American Scolytidae (Coleoptera). Great Basin Naturalist, 37, 67–74.

    Wood, S.L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1–1359.

    Wood, S.L. (1986) A reclassification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs, 10, 1–126.

    Wood, S.L. (1989) Nomenclatural changes and new species of Scolytidae (Coleoptera). Part IV. Great Basin Naturalist, 49, 167–185.

    Wood, S.L. (1992) Nomenclatural changes and new species in Platypodidae and Scolytidae (Coleoptera). 2. Great Basin Naturalist, 52, 78–88.

    Wood, S.L. (2007) Bark and ambrosia beetles of South America (Coleoptera, Scolytidae). Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah. 900 pp. + CCXXX pl.

    Wood, S.L. & Bright, D.E. (1987) A catalog of Scolytidae and Platypodidae (Coleoptera), Part 1: Bibliography. Great Basin Naturalist Memoirs, 11, 1–685.

    Wood, S.L. & Bright, D.E. (1992) A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: Taxonomic index. Great Basin Naturalist Memoirs 13, 1–1553.

    Yang, K.C., Lin, J.K., Hsieh, C.F., Huang, C.L., Chang, Y.M., Kuan, L.H., Su, J.F. & Chiu, S.T. (2008) Vegetation pattern and woody species composition of a broad-leaved forest at the upstream basin of Nantzuhsienhsi in mid-southern Taiwan. Taiwania, 53, 325–337.

    Yie, S.T. & Tang, P.M. (1966) A preliminary study on the ecosystem of the American pine tree insects. Taiwan Agriculture, 2(4), 1–36.

    Yuan, L.G. (2008) Research on the spatial distribution of population and integrated control technology of the main garden pest, Phloeosinus perlatus Chapuis. Journal of Anhui Agricultural Sciences, 36, 76–79. (In Chinese)

    Zchori-Fein, E., Borad, C. & Harai, A.L. (2006) Oogenesis in the date stone beetle, Coccotrypes dactyliperda, depends on symbiotic bacteria. Physiological Entomology, 31, 164–169.

    Zetterstedt, J.W. (1828) Fauna Insectorum Lapponica. Schulz, Hammone. 563 pp.

    Zhao, J.N. & Cao, B. (1987) Bionomics and control of Phloeosinus perlatus Chapuis. Insect Knowledge, 24, 227–230. (In Chinese, English summary).