Abstract
The wolf spider genus Anomalosa Roewer, 1960 is revised with Anomalosa kochi (Simon, 1898) as type species. Anomalosa includes a further Australian species, A. oz sp. nov. Representatives of Anomalosa are small, elongated lycosids with a longitudinal light median band on the dorsal shield of the prosoma and on the opisthosoma, the latter being particularly distinct in males. They are closely related to Venonia Thorell, 1894. Similar to Venonia, males have a bipartite prolateral tegular lobe on the pedipalp, but it is much larger than in Venonia and, in contrast to Venonia, larger than the membranous tegular apophysis. Anomalosa kochi has only been found in Queensland, whereas the distribution of A. oz sp. nov. includes New South Wales, South Australia and Victoria. This allopatric distribution coincides with the McPherson Range as a biogeographical border. Although most males and females of Anomalosa have been caught in pitfall traps or running freely in moderately moist habitats, such as near creeks and dams, there is evidence that representatives of this genus build sheet-webs similar to Venonia. This behaviour is supported by morphological evidence as species of Anomalosa have elongated posterior spinnerets. The original description of A. harishi (Dyal, 1935) from Panjab, India, does not match the diagnosis of Anomalosa. Consequently, I reject the inclusion of A. harishi in Anomalosa and re-transfer it to its original genus Anomalomma Simon, 1890, Anomalomma harishi Dyal, 1935, pending a systematic revision of this genus.References
Bonnet, P. (1955) Bibliographia Araneorum, Tome II (1re partie: A–B). Douladoure, Toulouse, 918 pp.
Coddington, J.A. (1990) Ontogeny and homology in the male palpus of orb-weaving spiders and their relatives, with comments on phylogeny (Araneoclada: Araneoidea, Deinopoidea). Smithsonian Contributions to Zoology, 49, 1–52.
Crisp, M.D., Linder, H.P. & Weston, P.H. (1995) Cladistic biogeography of plants in Australia and New Guinea: congruent pattern reveals two endemic tropical tracks. Systematic Biology, 44, 457–473.
Dahl, F. (1908) Die Lycosiden oder Wolfspinnen Deutschlands und ihre Stellung im Haushalt der Nature. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germaniae Naturea Curiosum, 88, 175–678.
Dondale, C.D. (1986) The subfamilies of wolf spiders (Araneae: Lycosidae). Actas X Congreso Internacional de Aracnología, Jaca, España, 1, 327–332.
Dyal, S. (1935) Spiders of Lahore. Bulletin of the Department of Zoology, Panjab Univiversity, 1, 117–252.
Ford, J. (1987) Hybrid zones in Australian birds. Emu, 87, 158–178.
Framenau, V.W. (2002) Review of the wolf spider genus Artoria Thorell (Araneae, Lycosidae). Invertebrate Systematics, 16, 209–235.
Framenau, V.W. (2005) The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae). Records of the Western Australian Museum, 22, 265–292.
Framenau, V.W. (2006) Revision of the wolf spider genus Diahogna Roewer, 1960 (Araneae, Lycosidae). Journal of Natural History, 40, 273–292.
Framenau, V.W., Gotch, T.B. & Austin, A.D. (in press) The wolf spiders of artesian springs in arid South Australia, with a revalidation of Tetralycosa (Araneae, Lycosidae). Journal of Arachnology.
Hasselt, A.W.M. van (1890) Araneae ex Archipelago Malayano. In: Weber, M., Zoologische Ergebnisse einer Reise in Niederländisch Ost-Indien. Leiden, 1, pp. 193–210.
Hippa, H. & Lehtinen, P.T. (1983) The zantheres group of Zoicinae (Araneae, Lycosidae) and a relimitation of the subfamily. Annales Zologici Fennici, 20, 151–156.
International Commission on Zoological Nomenclature (ICZN) (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, 306 pp.
James, C.H. & Moritz, C. (2000) Intraspecific phylogeography in the sedge frog Litoria fallax (Hylidae) indicates pre-Pleistocene vicariance of an open forest species from eastern Australia. Molecular Ecology, 9, 349–385.
Keogh, S.J., Scott, I.A.W., Fitzgerald, M. & Shine, R. (2003) Molecular phylogeny of the Australian venomous snake genus Hoplocephalus (Serpentes, Elapidae) and conservation genetics of the threatened H. stephensii. Conservation Genetics, 4, 57–65.
Koch, C.L. (1847) Die Arachniden. Vierzehnter Band. Zeh’sche Buchhandlung: Nürnberg, pp. 89–210.
Koch, L. (1877) Die Arachniden Australiens, nach der Natur beschrieben und abgebildet. Bauer and Raspe, Nürnberg, pp. 889–968.
Lehtinen, P.T. (2005) Review of the wolf spider genus Passiena (Araneae, Lycosidae). Journal of Arachnology, 33, 398–407.
Lehtinen, P.T. & Hippa, H. (1979) Spiders of the Oriental-Australian region. I. Lycosidae: Venoniinae and Zoicinae. Annales Zoologici Fennici, 16, 1–22.
McKay, R.J. (1973) The wolf spiders of Australia (Araneae: Lycosidae): 1. The bicolor group. Memoirs of the Queensland Museum, 16, 375–398.
McKay, R.J. (1979) The wolf spiders of Australia (Araneae: Lycosidae): 10. A new species of the genus Flanona Simon. Memoirs of the Queensland Museum, 19, 231–235.
McKay, R.J. (1985) Lycosidae. In: Walton, D.W. (Ed.), Zoological Catalogue of Australia, Vol. 3. Arachnida, Mygalomorphae, Araneomorphae in Part, Pseudoscorpionida, Amblypygida, Palpigradi. Australian Government Publishing Service, Canberra, 73–88.
Murphy, N.P., Framenau, V.W., Donellan, S.C., Harvey, M.S., Park, Y.-C. & Austin, A.D. (2006) Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12S rRNA, 28 rRNA and NADH1 genes: implications for classification, biogeography and the evolution of web building behavior. Molecular Phylogenetics and Evolution, 38, 583–602.
Platnick, N.I. (1993) Advances in Spider Taxonomy, 1988–1991. With Synonymies and Transfers 1940–1980. New York Entomological Society in association with the American Museum of Natural History, New York, 846 pp.
Platnick, N.I. (1998) [imprint date 1997]. Advances in Spider Taxonomy, 1992–1995. With Redescriptions 1940–1980. New York Entomological Society in association with the American Museum of Natural History, New York, 976 pp.
Rainbow, W.J. (1911) A census of Australian Araneidae. Records of the Australian Museum, 9, 107–319.
Roewer, C.F. (1955) [imprint date 1954] Katalog der Araneae von 1758 bis 1940. Vol. 2a. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 923 pp.
Roewer, C.F. (1960) [imprint date 1959]. Araneae Lycosiformae II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l’Upemba – Mission GF de Witte, 55, 519–1040.
Roewer, C.F. (1961) Über Namen der Gattungen und Arten der Lycosidae (Araneae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 37, 1–19.
Sheehan, D.C. & Hrapchak, B.B. (1980) Theory and Practice of Histotechnology. Second Edition. Battelle Press, Columbus, Ohio, 481 pp.
Simon, E. (1898a) Histoire Naturelle des Araignées. Roret, Paris, pp. 193–380.
Simon, E. (1898b) Descriptions d'arachnides nouveaux des familles des Agelenidae, Pisauridae, Lycosidae et Oxyopidae. Annales de la Société Entomologique de Belgique, 42, 5–34.
Sundevall, J.C. (1833) Conspectus Arachnidum. C.F. Berling, Lund, 39 pp.
Thorell, T. (1877) Studi sui Ragni Malesi e Papuani. I. Ragni di Selebes raccolti nel 1874 dal Dott. O. Beccari. Annali di Museo civico di storia naturale ‘Giacomo Doria’, Genova, 10, 341–637.
Thorell, T. (1894) Decas aranearum in ins. Singapore a Cel. Th. Workman inventarum. Bollettino della Societa Entomologica Italiana, 26, 321–355.
Vink, C.J. (2001) A revision of the genus Allotrochosina Roewer (Lycosidae: Araneae). Invertebrate Taxonomy, 15, 461–466.
Yoo, J.-C. & Framenau, V.W. (in press) Systematics and biogeography of the wolf spider genus Venonia (Araneae: Lycosidae). Invertebrate Systematics.
Zyuzin, A.A. (1993) Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum, 33, 693–700.