Abstract
Egle Robineau-Desvoidy is a modest-sized genus of small blackish anthomyiids with a Holarctic and northern Oriental distribution. They are univoltine, interacting closely with the host plants of their seed-feeding larvae, mainly species of willow (Salix), but also poplar (Populus) and false tamarisk (Myricaria) support some species. Summaries are given of adult and larval habits based on published and new observations. Adult mouth part structure in relation to pollen feeding is treated in some detail. A taxonomic revision with illustrated descriptions and an identification key to males and females is given for all known West Palearctic species and a few more from Greenland and Central Asia. Seven new species are described: Egle anderssoni, E. ignobilis and E. suwai from Denmark and Sweden, E. inermoides from Spain, E. groenlandica from Greenland, E. setiapicoides from Uzbekistan, and E. subarcticoides from Israel. Egle polychaeta Griffiths, 2003 is considered identical with E. steini Schnabl, 1911 (syn. nov.). Egle groenlandica sp. nov. is only known from females and may be the first known case of a calyptrate fly reproducing parthenogenetically.References
Ackland, D.M. (1970) Notes on the Palaearctic species of Egle R.-D. (Dipt., Anthomyiidae) with descriptions of two new species. Entomologist’s Monthly Magazine, 105, 185–192.
Ackland, D.M. (1977) Some Anthomyiidae from Southern Spain, with descriptions of two new species (Insecta, Diptera). Steenstrupia, 4, 195–209.
Ackland, D.M. (1989) Anthomyiidae (Dipt.) new to Britain, with a description of a new species of Botanophila Lioy. Entomologist’s Monthly Magazine, 125, 211–230.
Ackland, D.M. (2003) Egle lyneborgi Ackland & Griffiths, 2003 (Diptera, Anthomyiidae) new to the British list and E. inermis Ackland, 1970 is a good species. Dipterists Digest (Second Series), 10 (2), 84.
Ackland, D.M. & Merz, B. (2003) New records of Anthomyiidae (Diptera) from Switzerland. Mitteilungen der schweizerischen entomologischen Gesellschaft, 76, 207–219.
Ackland, D.M. & Pont, A.C. (1977) Family Anthomyiidae. In: Delfinado, M.D. & Hardy, D.E. (Eds.), A catalog of the Diptera of the Oriental Region. Volume III. Suborder Cyclorrhapha (excluding division Aschiza). The University Press of Hawaii, Honolulu, pp. 439–446.
Audcent, H. (1932) Bristol insect fauna: Diptera (part 5). Proceedings of the Bristol Naturalists’ Society, 7, 358–370.
Barták, M. & Rozkošný, R. (1984) Faunistic records from Czechoslovakia. Diptera, Anthomyiidae. Acta entomologica bohemoslovakia, 81, 74–75.
Barták, M., Michelsen, V. & Rozkošný, R. (1989) Faunistic records from Czechoslovakia. Diptera, Anthomyiidae. Acta entomologica bohemoslovakia, 86, 315–318.
Barták, M., Michelsen, V. & Rozkošný, R. (1990) New records of Anthomyiidae from Czechoslovakia, with revised check list of Czechoslovak species (Diptera). Scripta facultatis scientiarium naturalium universitatis purkyniensis brunensis, 20, 439–449.
Beuk, P.L.Th., Prijs, H.J. & Jong, H. de (2002) Anthomyiidae. In: Beuk, P.L.Th. (Ed.), Checklist of the Diptera of the Netherlands, 315–321. KNNV Uitgeverij, Utrecht.
Bezzi, M. & Stein, P. (1907) Cyclorrapha Aschiza. Cyclorrapha Schizophora: Schizometopa. In: Becker, T., Bezzi, M., Kertész, K. & Stein, P. (Eds.), Katalog der palaarktischen Dipteren, 3, 828 pp. Budapest.
Chandler, P. (Ed.) (1998) Anthomyiidae. In: Checklists of insects of the British Isles (New Series). Part 1: Diptera (Incorporating a List of Irish Diptera). Handbooks for the Identification of British Insects, 12, 165–171.
Collin, J.E. (1947) On the identification of the type of Anthomyia minuta Meigen (1826) with consequential changes in the names of three British Anthomyiidae. Proceedings of the Royal Entomological Society, London, (B) 16, 33–34.
Coquillett, D.W. (1910) The type-species of the North American genera of Diptera. Proceedings of the United States National Museum, 37, 499–647.
Darvas, B. (2001) Anthomyiidae. In: Papp, L. (Ed.), Checklist of the Diptera of Hungary, Hungarian Natural History Museum, Budapest. 386–403.
Darvas, B., Dely-Draskovits, Á., Ackland, D.M. & Jermy, T. (2001) Anthomyiidae (Diptera) new to Hungary. Folia entomologica hungarica, 62, 307–312.
Dely-Draskovits, Á. (1993) Family Anthomyiidae. In: Soós, Á. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera, Hungarian Natural History Museum, Budapest. 13, 11–102.
Draber-Mońko, A. (1991) Anthomyiidae. In: Razowski, J. (Ed.), Checklist of animals of Poland, Ossolineum Press, Wrocław. 2, 233–237.
Dušek, J. (1969) Präimaginale Stadien mitteleuropäischer Anthomyiiden (Diptera) I. Acta scientiarum naturalium academiae scientiarum bohemoslovacae, (n.s.) 3 (2), 1–37, 10 pls.
Dušek, J. (1970) Präimaginale Stadien mitteleuropäischer Anthomyiiden (Diptera) II. Acta scientiarum naturalium academiae scientiarum bohemoslovacae, (n.s.) 4 (1), 1–29, 7 + 4 pls.
Elberg, K.Yu. (1969) Fam. Anthomyiidae. In: Bei-Bienko, G.Ya. (Ed.), [Keys to the Fauna of the USSR], 5, 458–501. Nauka Publishers, Leningrad [= St. Petersburg]. [in Russian; English translation 1988, Amerind Publishing Co., New Delhi.]
Falcoz, L. (1930) Observations biologiques sur divers Insectes des environs de Vienne en Dauphiné (6e note) Bulletin de la Société Entomologique de France, 1930, 148–154.
Fan, Z.-d. (Ed.) (1988) Diptera: Anthomyiidae. Economic insect fauna of China, 37, 396 pp., 10 pls. Science Press, Beijing. [in Chinese with English summary of new taxa]
Frey, R. (1916) Diptera Brachycera. In: Poppius, B., Lundström, C. & Frey, R., (Eds.), Dipteren aus dem Sarekgebiet. Naturwissenschaftliche Untersuchungen des Sarekgebirkes in Schwedisch-Lappland, 4 (Zool.), 681–696.
Gorodkov, K., Pont, A.C. & Rozkošný, R. (1995) Diptera Muscoidea. In: Minelli, A., Ruffo, S., & La Posta, S. (Eds.), Checklist delle specie della fauna Italiana, Calderini, Bologna. 77, 18 pp.
Gosseries, J. & Ackland, D.M. (1991) Anthomyiidae. In: Grootaert, P., De Bruyn, L. & De Meýer, M. (Eds.), Catalogue of the Diptera of Belgium, 184–186. Documents de travail de l’Institut royal des Sciences naturelles de Belgique, Brussels.
Griffiths, G.C.D. (2003) Anthomyiidae [part]. In: Griffiths, G.C.D. (Ed.), Flies of the Nearctic Region, 8 (2), 14, 2289–2484. E. Schweizerbart, Stuttgart.
Grossmann, A. (1998) Egle myricariae spec. nov. (Diptera: Anthomyiidae), eine neue Egle-Art auf Tamariske (Myricaria germanica). Studia dipterologica, 5, 323–334.
Grossmann, A. (2000) Anthomyiidae. In: Ziegler, J. & Menzel, F. (Eds.), Die historische Dipteren-Sammlung Carl Friedrich Ketel. Revision einer zwischen 1884 und 1903 angelegten Sammlung von Zweiflüglern (Diptera) aus Mecklenburg-Vorpommern. Nova supplementa entomologica, 14, 173–180.
Gäbler, H. (1939) Schäden an den weiblichen Weidenkätzchen durch Egle (Hylemyia) parva Rob.-Desv. (Dipt.). Mitteilungen der Deutschen Entomologischen Gesellschaft, 9, 27–33.
Gäbler, H. (1953) Beitrag zur Kennnis von tierischen Forstsamenschädlingen. Beiträge zur Entomologie, 3, 479–487.
Gäbler, H. (1958) Beiträge zur Kenntnis der Schadinsekten der Blüten und Samen von Forstgehölzen. Archiv für Forstwesen, 7, 786–827.
Hackman, W. (1980) A check list of the Finnish Diptera II. Cyclorrhapha. Notulae entomologicae, 60, 117–162.
Hennig, W. (1967a) Anthomyiidae [part]. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 7 (1), Lief. 271, 97–144. E. Schweizerbart, Stuttgart.
Hennig, W. (1967b) Anthomyiidae [part]. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 7 (1), Lief. 272, 145–192. E. Schweizerbart, Stuttgart.
Hennig, W. (1972) Anthomyiidae [part]. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 7 (1), Lief. 294, 425–472. E. Schweizerbart, Stuttgart.
Hennig, W. (1976) Anthomyiidae [part]. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 7 (1), Lief. 315, I–LXXVIII. E. Schweizerbart, Stuttgart.
Hua, L.-z. (2006) List of Chinese insects, 4, 2+6+540 pp., 7 pls. Sun Yat-sen University Press, Guangzhou.
Huckett, H.C. (1921) On the morphology of the ovipositor of certain anthomyian genera. Annals of the Entomological Society of America, 14, 290–320, pls. XX–XXVII.
Huckett, H.C. (1924) A systematic study of the Anthomyiinae of New York, with special reference to the male and female genitalia. Memoirs. Cornell University Agricultural Experiment Station, 77, 91 pp.
Huckett, H.C. (1928) Little known anthomyid flies that commonly occur on the catkins of willow (Muscidae, Diptera). Bulletin of the Brooklyn Entomological Society, 23, 70–83.
Huckett, H.C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 369 pp.
Huckett, H.C. (1971) The Anthomyiidae of California exclusive of the subfamily Scatophaginae (Diptera). Bulletin of the California Insect Survey, 12, 121 pp.
Jin, Z.-y., Ge, F.-x. & Fan, Z.-d. (1981) Descriptions of five new species of Anthomyiidae from South Gansu, China (Diptera). Entomotaxonomia, 3, 87–92. [in Chinese with English summary]
Karl, O. (1928) Zweiflügler oder Diptera II: Muscidae. In: Dahl, F. (Ed.), Die Tierwelt Deutschlands und der angrenzenden Meeresteile nach ihren Merkmalen und nach ihrer Lebensweise, Gustav Fischer, Jena. 13, iv + 232 pp.
Karl, O. (1934) Ergänzungen und Berichtigungen zu meiner Arbeit über die Musciden. (Prof. Dr. Fr. Dahl, Die Tierwelt Deutschlands, Teil 13.) Teil IV. Zoologischer Anzeiger, 107, 90–93.
Karl, O. (1937) Die Fliegenfauna Pommerns. Diptera Brachycera. (Fortsetzung und Schluss.) Stettiner entomologische Zeitung, 98, 125–159.
Karl, O. (1943) Ergänzungen und Berichtigungen zu meiner Arbeit über die Musciden. (Prof. Dr. Fr. Dahl, Die Tierwelt Deutschlands, Teil 13.) Teil VI. Stettiner entomologische Zeitung, 104, 64–77.
Karl, O. (1944) Ergänzungen und Berichtigungen zu meiner Arbeit: Die Fliegenfauna Pommerns. Diptera Brachycera (Stett. Ent. Ztg. 1935–37). Stettiner entomologische Zeitung, 105, 82–84.
Komzáková, O. (2006) Anthomyiidae Latreille, 1829. In: Jedlička, L., Stloukalová, V. & Kúdela, M. (Eds.), Checklist of Diptera of the Czech Republic and Slovakia. Electronic version 1. Comenius University, Bratislava. Available from:
http: //zoology.fns.uniba.sk/diptera + CD-ROM: ISBN 80-969629-0-6
Kramer, H. (1917) Die Musciden der Oberlausitz. Abhandlungen der naturforschende Gesellschaft zur Görlitz, 28, 257–352.
Kröber, O. (1931) Dipterenfauna von Schleswig-Holstein und den benachbarten westlichen Nordseegebieten. 3. Teil: Diptera Brachycera, Braulidae–Larvivoridae nebst Nachträgen zum 1. Teil. Verhandlungen des Vereins für naturwissenschaftliche Heimatforschung zu Hamburg, 23, 63–113.
Lutovinovas E. (2004) Fly (Diptera: Brachycera) species recorded in Lithuania for the first time. New and Rare for Lithuania Insect Species. Records and Descriptions, 16, 62–73.
Lyneborg, L. (1965) On Muscidae and Anthomyiidae (Diptera) from Iceland. With descriptions of two new species. Opuscula entomologica, 30, 211–226.
Malloch, J.R. (1920a) Descriptions of new North American Anthomyiidae (Diptera). Transactions of the American Entomological Society, 46, 133–196, 3 pls.
Malloch, J.R. (1920b) Scientific results of the Katmai Expeditions of the National Geographic Society. XII. Descriptions of Diptera of the families Anthomyidae and Scatophagidae. Ohio Journal of Science, 20, 267–288, pls. 2–4.
Meigen, J.W. (1826) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten, 5, xii + 412 pp. Hamm.
Meijere, J.C.H. de (1907) Eerste supplement op de Nieuwe Naamlijst van Nederlandsche Diptera. Tijdschrift voor Entomologie, 50, 151–195, 1 pl. [in Dutch]
Meijere, J.C.H. de (1919) Derdee supplement op de Nieuwe Naamlijst van Nederlandsche Diptera. Tijdschrift voor Entomologie, 62, 161–195. [in Dutch]
Meixnerová, O. & Rozkošný, R. (1999) Anthomyiidae. In: Rozkošný, R. & Vaňhara, J. (Eds.), Diptera of the Pálava Reserve of UNESCO, II. Folia facultatis scientiarium naturalium universitatis masarykianae brunensis, Biologia, 100, 379–386.
Merz, B., Bächli, G. & Haenni, J.-P. (2006) Zweiter Nachtrag zur Checkliste der Diptera der Schweiz. Mitteilungen der Entomologischen Gesellschaft Basel, 56, 135–165.
Michelsen, V. (1976) Scatophagidae, Anthomyiidae, Fanniidae og Muscidae fra Anholt (Diptera, Muscoidea). Flora og Fauna, 82, 71–78. [in Danish]
Michelsen, V. (1979) Notes on the identity and type-material of the Anthomyiidae, Fanniidae, and Muscidae (Diptera) described by Fabricius, J.C., Steenstrupia, 5, 181–196.
Michelsen, V. (1985) A revision of the Anthomyiidae (Diptera) described by Zetterstedt, J.W., Steenstrupia, 11, 37–65.
Michelsen, V. (1988) A world revision of Strobilomyia gen.n.: the anthomyiid seed pests of conifers (Diptera: Anthomyiidae). Systematic Entomology, 13, 271–314.
Michelsen, V. (2004) Anthomyiidae. In: Pape, T. (Ed.), Diptera Brachycera. Fauna Europaea version 1.1. Available from: http://www.faunaeur.org.
Michelsen, V. (2006) Annotated catalogue of the Anthomyiidae, Fanniidae, Muscidae and Scathophagidae (Diptera: Muscoidea) of Greenland. Steenstrupia, 29, 105–126.
Michelsen, V. & Báez, M. (2002) Anthomyiidae. In: Carles-Tolrá Hjorth-Andersen, M. (Ed.), Catálogo de los Diptera de España, Portugal y Andorra (Insecta). Monografías S.E.A., 8, 200–203.
Michelsen, V. & Barták, M. (2001) Anthomyiidae. In: Barták, M. & Vaňhara, J. (Eds.), Diptera in an industrially affected region (North-Western Bohemia, Bílina and Duchcov environs) II. Folia facultatis scientiarium naturalium universitatis masarykianae brunensis, Biologia, 105, 455–62.
Michelsen, V. & Merz, B. (1998) Anthomyiidae. In: Merz, B., Bächli, G., Haenni, J.-P. & Gonseth, Y. (Eds.), Diptera – Checklist. Fauna helvetica, 1, 313–317.
Ólafsson, E. (1991) A checklist of Icelandic insects. Fjölrit Náttúrufræðistofnunar, 17, 69 pp. [in Icelandic with English summary]
Pandellé, L. (1900) Études sur les Muscides de France. 3e partie. [Part]. Revue d’Entomologie, 18 (Suppl.), 221–292.
Petersen, F.T. (2001) Anthomyiidae. In: Petersen, F.T. & Meier, R. (Eds.), A preliminary list of the Diptera of Denmark. Steenstrupia, 26, 182–186.
Pont, A.C. (1960) Egle parvaeformis Schnabl (Dipt., Muscidae), a species new to Britain. Entomologist’s Record & Journal of Variation, 72, 148.
Ringdahl, O. (1914) Bidrag till kännedomen om våra anthomyider. Entomologisk Tidskrift, 33, 142–154. [in Swedish]
Ringdahl, O. (1918) Neue nordische Anthomyiden. Entomologisk Tidskrift, 39, 148–194.
Ringdahl, O. (1930) Entomologische Ergebnisse der schwedischen Kantchatka-Expedition 1920–1922. 30. Diptera Brachycera 3. Fam. Muscidae. Arkiv för Zoologi, 21A (20), 16 pp.
Ringdahl, O. (1931) Flugor – Diptera brachycera. In: Sjöstedt, Y. (Ed.), Insektfaunan inom Abisko Nationalpark. III. K. Svenska Vetenskapsakademiens skrifter i naturskyddsärenden, 18, 1–32. [in Swedish]
Ringdahl, O. (1933) Översikt av i Sverige funna Hylemyia-arter. Entomologisk Tidskrift, 54, 1–35. [in Swedish]
Ringdahl, O. (1939a) Diptera Brachycera i Regio alpina. Entomologisk Tidskrift, 60, 37–50. [in Swedish]
Ringdahl, O. (1939b) Diptera der Fam. Muscidae, (die Gattungen Aricia und Anthomyza) von Zetterstedt in “Insecta Lapponica” und “Diptera Scandinaviae” beschrieben. Opuscula entomologica, 4, 137–159.
Ringdahl, O. (1951) Fliegen von den Fjeldgegenden der Provinzen Lappland, Jämtland und Härjedalen. Opuscula entomologica, 16, 113–186. [in Swedish with German summary]
Ringdahl, O. (1952) Catalogus insectorum sueciae. XI. Diptera Cyclorrapha: Muscaria Schizometopa. Opuscula entomologica, 17, 129–186.
Ringdahl, O. (1957) Neue isländische Dipteren nebst Beschreibung einer neuen Pegomyia-Art. Entomologiske Meddelelser, 28, 101–103.
Ringdahl, O. (1958) Dipterologiska notiser. 16. Norrländska Diptera Brachycera. 17. Värmländska Diptera Brachycera. Opuscula entomologica, 23, 90–94. [in Swedish with German summary]
Ringdahl, O. (1959) Tvåvingar. Diptera Cyclorapha Schizophora Schizometopa. 1. Fam. Muscidae. Häfte 3. Svensk Insektfauna, 11, 197–334. Stockholm. [in Swedish]
Ringdahl, O. (1960) Flugfaunan på Kullaberg och Hallands Väderö. Kullabergs Natur, 2, 1–40. [in Swedish]
Robineau-Desvoidy, J. B. (1830) Essai sur les myoidaires. Mémoires présentés par divers Savans à l’Académia Royale des Sciences de l’Institut de France 2 (2), 1–813.
Rondani, C. (1871) Diptera Italica non vel minus cognita descripta aut annotata. Fasc. IV. Addenda Anthomyinis Prodr. Vol. VI. Bollettino della Società Entomologica Italiana, 2 (1870), 317–338.
Rondani, C. (1877) Species Italicae ordinis dipterorum ordinatim dispositae, methodo analitica distinctae, et novis vel minus cognitis descriptis. Pars quinta. Stirps XVII-Anthomyinae. 304 pp. Parmae [= Parma].
Rottenberg, A., Zohary, D. & Nevo, E. (1999) Patterns of isozyme diversity and vegetative reproduction of willows in Israel. International Journal of Plant Science, 160, 561–566.
Rozkošný, R. & Barták, M. (1984) Faunistic records from Czechoslovakia. Diptera, Anthomyiidae. Acta entomologica bohemoslovakia, 81, 465–466.
Schnabl, J. & Dziedzicki, H. (1911) Die Anthomyiden. Nova Acta. Abh. der Kaiserl. Leop.-Carol. Deutschen Akademie der Naturforscher, 95 (2), 53–358, pls. 3–37.
Séguy, E. (1923) Diptères Anthomyides. Faune de France, 6, i–xi +1–393. Paris.
Séguy, E. (1934) Contribution a l’etude des mouches phytophages de l’Europe occidentale. II. Encyclopédie Entomologique, (B II) Diptera, 7, 231–264.
Séguy, E. (1937) Diptera. Fam. Muscidae. In: Wytsman, P. (Ed.), Genera insectorum, 205, 1–604, 9 pls.
Silén, F. (1906) Blombiologiska iakttagelser i Kittilä Lappmark. Meddelanden af Societas pro fauna et flora fennica, 31, 80–99. [in Swedish]
Stein (1901) Die Walker’schen aussereuropäischen Anthomyiden in der Sammlung des British Museum zu London. Zeitschrift für systematische Hymenopterologie und Dipterologie, 1, 185–221.
Stein, P. (1907) Zur Kenntnis der Dipteren von Central-Asien. II. Cyclorrhapha schizophora schizometopa. Die von Roborowsky und Kozlov in der Mongolei und Tibet gesammelten Anthomyiiden. L’Annuaire du Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 12, 318–372.
Stein, P. (1916) Die Anthomyiden Europas. Tabellen zur Bestimmung der Gattungen und aller mir bekannten Arten, nebst mehr oder weniger ausführlichen Beschreibungen. Archiv für Naturgeschichte, 81 A (1) (‘1915’), 1–224.
Suwa, M. (1974) Anthomyiidae of Japan (Diptera). Insecta matsumurana (n.s.), 4, 1–247.
Suwa, M. (1999) Japanese records of anthomyiid flies (Diptera: Anthomyiidae). Insecta matsumurana (n.s.), 55, 203–244.
Suwa, M. & Darvas, B. (1998) Family Anthomyidae. In: Papp, L. & Darvas, B. (Eds.), Contributions to a Manual of Palaearctic Diptera, 3, 571–616. Science Herald, Budapest.
Teschner, D. (1999) Anthomyiidae. In: Schumann, H., Bährmann, R. & Stark, A. (Eds.), Checkliste der Dipteren Deutschlands. Studia dipterologica, Supplement, 2, 130–133.
Tiensuu, L. (1936) Die bisher aus Finland bekannten Musciden. Acta societatis pro fauna et flora fennica, 58 (4), 56 pp.
Tiensuu, L. (1938) Beiträge zur Kenntnis der Musciden (Dipt.) Finnlands. 1. Annales entomologici fennici, 4, 21–33.
Urban, C. & Kopelke, J.-P. (2004) The composition and function of insect communities living in catkins of willows (Salix spp.) (Insecta: Coleoptera; Diptera; Lepidoptera; Thysanoptera; Hemiptera; Hymenoptera). Senckenbergiana biologica, 84, 81–95.
Villeneuve, J. (1923) Diptères nouveaux. Bulletin du Muséum National d’Histoire Naturelle, Paris, 29, 91–96.
Walker, F. (1849) List of the specimens of dipterous insects of the British Museum. 4, 689–1172. London.
Wei, L.-m., Xue, W.-q. & Cui, C.-y. (1998) Anthomyiidae. In: Xue, W.-q. & Chao, C.-m. (Eds.), Flies of China, 1 (‘1996’), 634–808. Liaoning Science & Technology Press, Shenyang. [in Chinese with English summary]
Yang, Q. & Gaskin, J. (2007) Tamaricaceae. In: Wu, Z.-g., Raven, P.H. & Hong, D.-y. (Eds.), Flora of China. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis. 13, 66–69.
Zangheri, P. (1950) Fauna di Romagna. Ditteri. 2a parte. Memorie della Società entomologica Italiana, 29, 68–95.
Zetterstedt, J.W. (1838) Dipterologis Scandinaviae. Sect. 3: Diptera. In: Zetterstedt, J.W., Insecta Lapponica, 477–868. Lipsiae [= Leipzig].
Zetterstedt, J.W. (1845) Diptera Scandinaviae disposita et descripta, 4, 281–1738. Lundae [= Lund].
Zheng, S.-s. & Fan, Z.-d. (1990) Descriptions of some new species of family Anthomyiidae from western China (Diptera). Contributions of the Shanghai Institute of Entomology, 9, 181–187. [in Chinese with English summary]