Skip to main content Skip to main navigation menu Skip to site footer
Type: Articles
Published: 2009-09-02
Page range: 35–58
Abstract views: 35
PDF downloaded: 2

The Marjuman trilobite Cedarina Lochman: thoracic morphology, systematics, and new species from western Utah and eastern Nevada, USA

Department of Geoscience, 121 Trowbridge Hall, University of Iowa, Iowa City, Iowa 52242, USA
Department of Geology & Geophysics, University of Wisconsin-Madison, 1215 W. Dayton St., Madison, Wisconsin 53706, USA
Oklahoma Museum of Natural History and School of Geology and Geophysics, University of Oklahoma, Norman, Oklahoma 73072, USA
Trilobita Cedariidae Remopleuridoidea <i>Cedarina</i> taxonomy Cambrian

Abstract

Cedarina schachti n. sp. from the Marjuman (Cedaria Zone) Weeks Formation of western Utah, USA, provides the first information on thoracic morphology within the genus. Its thorax is radically different from those of species of Cedaria Walcott, with which Cedarina Lochman has been classified in Cedariidae Raymond, but strikingly similar to those of plesiomorphic remopleuridoideans grouped in the paraphyletic Richardsonellinae Raymond. If Cedarina and the remopleuridoideans are genuinely related it follows that 1) Cedariidae as traditionally conceived is paraphyletic; 2) Cedarina is a plesiomorphic sister taxon of the remopleuridoideans; and 3) the remopleuridoideans are not a component of the Order Asaphida. Silicified material of a second new species, C. clevensis from the Marjuman (Crepicephalus Zone) Lincoln Peak Formation of eastern Nevada, confirms the presence of a long thoracic axial spine and provides the first information on ontogenetic development and ventral morphology within the genus.

References

  1. Adrain, J.M. & Westrop, S.R. (2004) A Late Cambrian (Sunwaptan) silicified trilobite fauna from Nevada. Bulletins of American Paleontology, 365, 1–51.

    Adrain, J.M. & Westrop, S.R. (2005) Late Cambrian ptychaspidid trilobites from western Utah: implications for trilobite systematics and biostratigraphy. Geological Magazine, 142, 377–398.

    Angelin, N.P. (1854) Palaeontologica Scandinavica. Pars II, Crustacea formationis transitionis. Academiae Regiae Scientarum Suecanae (Holmiae), 25–92.

    Barrande, J. (1872) Systême silurien du centre de la Bohême, Ière Partie; Recherches paléontologiques, I. Supplément. Prague and Paris, 647 pp.

    Bordonaro, O. (2003) Review of the Cambrian stratigraphy of the Argentine Precordillera. Geologica Acta, 1, 11–21.

    Borello, A. (1965) Sobre el desarrollo bioestratigráfico del Cámbrico de la Precordillera. Acta Geológica Lilloana, 7, 39.

    Burmeister, H. (1843) Die Organisation der Trilobiten aus ihren lebenden Verwandten entwickelt; nebst einer systematischen Uebersicht aller zeither beschriebenen Arten. Reimer, Berlin, 147 pp.

    Černyševa, N.E. (1960) Osnovy paleontologii; spravočnik dlâ paleontologov i geologov SSSR. 8, Členistonogie, trilobitoobraznye i rakoobraznye. Akademiâ Nauk SSSR, Moscow, 515 pp. (in Russian)

    Černyševa, N.E. (1968) [Middle Cambrian trilobites of the north Caucasus]. Paleontologičeskij Žurnal, 1968, 71–80. (in Russian)

    Dean, W.T. (2006) Cambrian stratigraphy and trilobites of the Samur Dağ area, south of Hakkâri, southeastern Turkey. Turkish Journal of Earth Sciences, 15, 225–257.

    Deland, C.R. & Shaw, A.B. (1956) Upper Cambrian trilobites from western Wyoming. Journal of Paleontology, 30, 542–562.

    Drewes, H. & Palmer, A.R. (1957) Cambrian rocks of the southern Snake Range. Bulletin of the American Association of Petroleum Geologists, 41, 1–104.

    Egorova, L.I., Shabanov, Y.Y., Rozanov, A.Y., Stavitsky, V.E. & Černyševa, N.E. (1982) [Maya Stage of type locality (Middle Cambrian of Siberian Platform)]. Akademiâ Nauk SSSR, Ministerstvo Geologii, Mežvedomstvennyj Stratigrafičeskij Komitet SSSR, Trudy, 8, 1–146.

    Fortey, R.A. (1974) A new pelagic trilobite from the Ordovician of Spitsbergen, Ireland and Utah. Palaeontology, 17, 111–124.

    Fortey, R.A. (1975) The Ordovician trilobites of Spitsbergen. II. Asaphide, Nileidae, Raphiophoridae and Telephinidae of the Valhallfonna Formation. Norsk Polarinstitutt Skrifter, 162, 1–207.

    Fortey, R.A. (1979) Early Ordovician trilobites from the Catoche Formation (St. George Group), western Newfoundland. Geological Survey of Canada Bulletin, 321, 61–114.

    Fortey, R.A. (1980) The Ordovician trilobites of Spitsbergen. III. Remaining trilobites of the Valhallfonna Formation. Norsk Polarinstitutt Skrifter, 171, 1–163.

    Fortey, R.A. (1985) Pelagic trilobites as an example of deducing the life habits of extinct arthropods. Transactions of the Royal Society of Edinburgh: Earth Sciences, 76, 219–230.

    Fortey, R.A. (1994) Late Cambrian trilobites from the Sultanate of Oman. Neues Jahrbuch fuer Geologie und Palaeontologie. Abhandlungen, 194, 25–53.

    Fortey, R.A. (2005) Opipeuterella, a replacement name for the trilobite Opipeuter Fortey, 1974, preoccupied. Journal of Paleontology, 79, 1036.

    Fortey, R.A. & Chatterton, B.D.E. (1988) Classification of the trilobite suborder Asaphina. Palaeontology, 31, 165–222.

    Fortey, R.A. & Owens, R.M. (1987) The Arenig Series in South Wales. Bulletin of the British Museum (Natural History), Geology Series, 41, 69–305.

    Harrington, H.J. (1938) Sobre las faunas del Ordoviciano Inferior del Norte Argentino. Revista del Museo de La Plata. Sección Paleont. N. S., 1, 109–289.

    Harrington, H.J. & Leanza, A.F. (1957) Ordovician trilobites of Argentina. Special Publications, Department of Geology, University of Kansas, Lawrence, Kansas, 276 pp.

    Hawle, I. & Corda, A.J.C. (1847) Prodrom einer Monographie der böhmischen Trilobiten. J. G. Calve, Prague, 176 pp.

    Hintze, L.F. & Davis, F.D. (2003) Geology of Millard County, Utah. Utah Geological Survey Bulletin, 133, 1–305.

    Howell, B.F. & Duncan, D. (1939) Middle-Upper Cambrian transition fauna of North America. Bulletin of the Wagner Free Institute of Science, 14, 1–11.

    Hu, C.-H. (1971) Ontogeny and sexual dimorphism of lower Paleozoic Trilobita. Palaeontographica Americana, 7, 31–155.

    Hu, C.-H. (1983) The ontogenetic development of two upper Middle Cambrian trilobites from the Bonnetterre Dolomite, Missouri (with a discussion on the morphologic varieties of Cedarina vale Lochman). Memoir of the Geological Society of China, 5, 275–280.

    Hu, C.-H. & Li, I.-L. (1971) Ontogenies of four proparian trilobites from the Upper Cambrian, Montana, U.S., (with a description of Vernaculina lushihi n. sp.). Proceedings of the Geological Society of China, 14, 165–184.

    Hughes, N.C., Gunderson, G.O. & Weedon, M.J. (1997) Circumocular suture and visual surface of "Cedaria" woosteri (Trilobita, Late Cambrian) from the Eau Claire Formation, Wisconsin. Journal of Paleontology, 71, 103–107.

    Jago, J.B. & Cooper, R.A. (2005) A Glyptagnostus stolidotus trilobite fauna from the Cambrian of northern Victoria Land, Antarctica. New Zealand Journal of Geology and Geophysics, 48, 661–681.

    Jell, P.A. (1985) Tremadoc trilobites of the Digger Island Formation, Waratah Bay, Victoria. Memoirs of the Museum of Victoria, 46, 53–88.

    Jell, P.A. & Adrain, J.M. (2003) Available generic names for trilobites. Memoirs of the Queensland Museum, 48, 331–553.

    Kobayashi, T. (1933) Upper Cambrian of the Wuhutsui Basin, Liaotung, with special reference to the limit of Chaumitien (or Upper Cambrian), of eastern Asia, and its subdivision. Japanese Journal of Geology and Geography, 11, 55–155.

    Kobayashi, T. (1935a) The Cambro-Ordovician formations and faunas of South Chosen. Palaeontology. Part III. The Cambrian faunas of South Chosen with a special study on the Cambrian trilobite genera and families. Journal of the Faculty of Science, Tokyo University, Section 2, 4, 49–344.

    Kobayashi, T. (1935b) On the Kainella fauna of the basal Ordovician age found in Argentina. Japanese Journal of Geology and Geography, 12, 59–67.

    Kobayashi, T. (1953) On the Kainellidae. Japanese Journal of Geology and Geography, 23, 37–61.

    Laurie, J.R. & Shergold, J.H. (1996) Early Ordovician trilobite taxonomy and biostratigraphy of the Emanuel Formation, Canning Basin, Western Australia. Part 1. Palaeontographica Abteilung A, 240, 65–103.

    Linnarsson, J.G.O. (1869) Om Vestergötlands Cambriska och Siluriska aflagringar. Kungliga Svenska Vetenskapsakademiens Handlingar, 8, 1–89.

    Lochman, C. (1936) New trilobite genera from the Bonneterre Dolomite of Missouri. Journal of Paleontology, 10, 35–43.

    Lochman, C. (1940) Fauna of the basal Bonneterre Dolomite (Upper Cambrian) of southeastern Missouri. Journal of Paleontology, 14, 1–53.

    Lochman, C. (1950) Upper Cambrian faunas of the Little Rocky Mountains, Montana. Journal of Paleontology, 24, 322–349.

    Lochman, C. & Duncan, D. (1944) Early Upper Cambrian faunas of central Montana. Geological Society of America Special Papers, 54, 1–181.

    Lochman, C. & Duncan, D. (1949) Homonyms and synonyms in "Early Upper Cambrian faunas of central Montana". Journal of Paleontology, 23, 439.

    Lochman, C. & Hu, C.-H. (1961) Upper Cambrian faunas from the northwest Wind River Mountains, Wyoming. Part II. Journal of Paleontology, 35, 125–146.

    Lochman, C. & Hu, C.-H. (1962) Upper Cambrian faunas from the northwest Wind River Mountains, Wyoming, Part III. Journal of Paleontology, 36, 1–28.

    Lu, Y.-H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, N. S. B, 11, 1–463. (in Chinese and English)

    Lu, Y.-H. & Qian, Y.-Y. (1983) New zonation and correlation of the Upper Cambrian Changshanian Stage in North China. Acta Palaeontologica Sinica, 22, 134.

    Ludvigsen, R. (1982) Upper Cambrian and Lower Ordovician trilobite biostratigraphy of the Rabbitkettle Formation, western District of Mackenzie. Royal Ontario Museum Life Sciences Contributions, 134, 1–188.

    Ludvigsen, R. & Chatterton, B.D.E. (1991) The peculiar Ordovician trilobite Hypodicranotus from the Whittaker Formation, District of Mackenzie. Canadian Journal of Earth Sciences, 28, 616–622.

    Ludvigsen, R. & Westrop, S.R. (1985) Three new Upper Cambrian stages for North America. Geology, 13, 139–143.

    Ludvigsen, R., Westrop, S.R. & Kindle, C.H. (1989) Sunwaptan (Upper Cambrian) trilobites of the Cow Head Group, western Newfoundland, Canada. Palaeontographica Canadiana, 6, 1–175.

    Marek, L. (1952) Contribution to the stratigraphy and fauna of the uppermost part of the Králův Dvůr Shales (Ashgillian). Sborník Ústředního Ústavu Geologckého, 19, 429–455. (in Czech, with Russian and English summaries)

    Öpik, A.A. (1967) The Mindyallan fauna of north-western Queensland. Bureau of Mineral Resources, Geology and Geophysics, Bulletin, 74, vol 1: 404 p. vol 2: 167 p.

    Osmólska, H. (1957) Trilobites from the Couvinian of Wydryszów (Holy Cross Mountains, Poland). Acta Palaeontologica Polonica, 2, 53–77.

    Palmer, A.R. (1955) The faunas of the Riley Formation in central Texas. Journal of Paleontology, 28, 709–786. (for 1954)

    Palmer, A.R. (1962) Glyptagnostus and associated trilobites in the United States. United States Geological Survey Professional Paper, 374-F, 1–49.

    Palmer, A.R. (1971) The Cambrian of the Great Basin and adjacent areas, western United States. In: Holland, C.H. (Ed.), Cambrian of the New World. Wiley-Interscience, London, New York, Sydney, Toronto, pp. 1–78.

    Palmer, A.R. (1998) A proposed nomenclature for stages and series of the Cambrian of Laurentia. Canadian Journal of Earth Sciences, 35, 323–328.

    Park, T.-Y. & Choi, D.K. (2008) A silicified Furongian trilobite fauna from Korea and its implications for monophyly of the Order Asaphida. In: Rábano, I., Gozalo, R. & García-Bellido, D. (Eds.), Advances in trilobite research. Cuadernos del Museo Geominero, 9, 305–306.

    Pegel, T.V. (2000) Evolution of trilobite biofacies in Cambrian basins of the Siberian Platform. Journal of Paleontology, 74, 1000–1019.

    Pegel, T.V. & Gogin, I.Y. (1995) [Upper Cambrian trilobites of southern Verkhoyanye]. Paleontologičeskij Žurnal, 1995, 67–74. (in Russian)

    Peng, S.-C. (1990) Tremadoc stratigraphy and trilobite faunas of northwestern Hunan. 2. Trilobites from the Panjiazui Formation and the Madaoyu Formation in Jiangnan Slope Belt. Beringeria, 2, 55–171.

    Peng, S.-C. (1992) Upper Cambrian biostratigraphy and trilobite faunas of the Cili-Taoyuan area, northwestern Hunan, China. Memoir of the Association of Australasian Palaeontologists, 1–119.

    Peng, S.-C., Babcock, L.E. & Lin, H.-L. (2004) Polymerid trilobites from the Cambrian of northwestern Hunan, China. Volume 2. Ptychopariida, Eodiscida, and undetermined forms. Science Press, Beijing, 355 pp.

    Poletaeva, O.K. & Romanenko, E.V. (1970) [Middle and Late Cambrian trilobites of the Altay]. Paleontologičeskij Žurnal, 1970, 72–83. (in Russian)

    Poulsen, C. (1927) The Cambrian, Ozarkian and Canadian faunas of northwest Greenland. Meddelelser om Grønland, 70, 235–343.

    Pratt, B.R. (1992) Trilobites of the Marjuman and Steptoean stages (Upper Cambrian), Rabbitkettle Formation, southern Mackenzie Mountains, northwest Canada. Palaeontographica Canadiana, 9, 1–179.

    Qian, Y.-Y. (1994) Trilobites from the Middle Upper Cambrian (Changshan Stage) of north and northeast China. Palaeontologia Sinica, 183, 1–190. (in Chinese with English summary)

    Rasetti, F. (1965) Upper Cambrian trilobite faunas of northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1–127.

    Raymond, P.E. (1924) New Upper Cambrian and Lower Ordovician trilobites from Vermont. Proceedings, Boston Society of Natural History, 37, 389–446.s

    Raymond, P.E. (1937) Upper Cambrian and Lower Ordovician Trilobita and Ostracoda from Vermont. Geological Society of America Bulletin, 48, 1079–1146.

    Rees, M.N. (1986) A fault-controlled trough through a carbonate platform: The Middle Cambrian House Range embayment. Geological Society of America Bulletin, 97, 1054–1069.

    Resser, C.E. (1938) Cambrian System (restricted) of the southern Appalachians. Geological Society of America Special Paper, 15, 1–140.

    Robison, R.A. (1988) Trilobites of the Holm Dal Formation (late Middle Cambrian), central North Greenland. Meddelelser om Grønland, Geoscience, 23–103.

    Rozova, A.V. (1964) [Biostratigraphy and description of Middle and Upper Cambrian trilobites from the north-west Siberian Platform]. Trudy Instituta Geologii i Geofiziki Akademiâ Nauk SSSR, 1–106. (in Russian)

    Saito, K. & Sakakura, K. (1936) Le Cambrien du côté sud-oriental de la Pli-faille inverse de Zedô, Heian-nando, Corée. Appendice: Description de deux nouvelles espèces de Trilobite. Journal of the Geological Society of Japan, 434, 104–117.

    Shergold, J.H. (1972) Late Upper Cambrian trilobites from the Gola Beds, western Queensland. Bureau of Mineral Resources, Geology and Geophysics, Australia, Bulletin, 112, 1–89. (for 1971)

    Shergold, J.H. (1975) Late Cambrian and Early Ordovician trilobites from the Burke River Structural Belt, western Queensland, Australia. Bureau of Mineral Resources, Geology and Geophysics, Australia, Bulletin, 153, 1–251.

    Shergold, J.H. (1980) Late Cambrian trilobites from the Chatsworth Limestone, western Queensland. Australian Bureau of Mineral Resources, Geology, and Geophysics Bulletin, 186, 1–111.

    Stitt, J.H. (1998) Trilobites from the Cedarina dakotaensis Zone, lowermost part of the Deadwood Formation (Marjuman Stage, Upper Cambrian), Black Hills, South Dakota. Journal of Paleontology, 72, 1030–1046.

    Stubblefield, C.J. (1959) Evolution in trilobites. An address to the Geological Society of London at its anniversary meeting on 29 April, 1959. Quarterly Journal of the Geological Society of London, 115, 145–162.

    Sun, Y.C. (1935) The Upper Cambrian trilobite faunas of North China. Palaeontologia Sinica, Series B, 2(2), 1–69.

    Sundberg, F.A. (2005) The Topazan Stage, a new Laurentian stage (Lincolnian Series - "Middle" Cambrian). Journal of Paleontology, 79, 63–71.

    Ulrich, E.O. & Resser, C.E. (1930) The Cambrian of the Upper Mississippi Valley, part 1, Trilobita; Dikelocephalinae and Osceolinae. Bulletin of the Public Museum of the City of Milwaukee, 12, 1–122.

    Walcott, C.D. (1884) The paleontology of the Eureka District, Nevada. United States Geological Survey Monograph, 8, 1–298.

    Walcott, C.D. (1916a) Cambrian Geology and Paleontology, III. No. 3 - Cambrian trilobites. Smithsonian Miscellaneous Collections, 64, 157–258.

    Walcott, C.D. (1916b) Cambrian Geology and Paleontology, III. No. 5 - Cambrian trilobites. Smithsonian Miscellaneous Collections, 64, 303–456.

    Walcott, C.D. (1924) Cambrian geology and palaeontology, V. No. 2 - Cambrian and Lower Ozarkian trilobites. Smithsonian Miscellaneous Collections, 75, 53–60.

    Walcott, C.D. (1925) Cambrian geology and paleontology, V. No. 3 - Cambrian and Lower Ozarkian trilobites. Smithsonian Miscellaneous Collections, 75, 61–146.

    Westrop, S.R. & Ludvigsen, R. (2000) The Late Cambrian (Marjuman) trilobite genus Hysteropleura Raymond from the Cow Head Group, western Newfoundland. Journal of Paleontology, 74, 1020–1030.

    Whitehouse, F.W. (1939) The Cambrian faunas of north-eastern Australia. Part 3. Memoirs of the Queensland Museum, 11, 179–282.

    Whitfield, R.P. (1882) Palaeontology. In: Geology of Wisconsin, Volume 4, Part 3. pp. 163–363.

    Whittington, H.B. (1952) A unique remopleuridid trilobite. Breviora, 4, 1–9.

    Whittington, H.B. (1959) Silicified Middle Ordovician trilobites: Remopleurididae, Trinucleidae, Raphiophoridae, Endymionidae. Bulletin of the Museum of Comparative Zoology, Harvard, 121, 371–496.

    Whittington, H.B. (1981) Paedomorphosis and cryptogenesis in trilobites. Geological Magazine, 118, 591–602.

    Wilson, J.L. (1954) Late Cambrian and early Ordovician trilobites from the Marathon Uplift, Texas. Journal of Paleontology, 28, 249–285.

    Yuan, J.-L. & Yin, G.-Z. (1998) New polymerid trilobites from the Chefu Formation in early Late Cambrian of eastern Guizhou. Acta Palaeontologica Sinica, 37, 137–172.

    Zhang, W.-T. (1963) A classification of the Lower and Middle Cambrian trilobites from the north and northeastern China, with descriptions of new families and new genera. Acta Palaeontologica Sinica, 11, 447–487. (in Chinese with English summary; Chang, W.-T.)

    Zhou, Z.-Y. & Zhang, J.-L. (1978) Cambrian-Ordovician boundary of the Tanshan area with descriptions of the related trilobite fauna. Acta Palaeontologica Sinica, 17, 1–26. (in Chinese with English summary)