Abstract
The species belonging to the subgenus Molopsis Schatzmayr, 1943 of genus Tapinopterus Schaum, 1858 are revised. Thestudy is based on 372 specimens and includes, for each taxon, diagnosis, description (only for the new species), references,new distributional records and illustrations. Morphological characters of the male and female genitalia are widely used todelimit the separate species, the significance of some non-gonapophyseal genital structures of the females for the system-atics of Molopsis is explicitly emphasized. For the first time, the male genitalia of T. molopiformis, T. molopinus, T. wie-demanni, and T. rebellis, including also newly described species, as well as female genitalia of Molopsis are described andillustrated. All available type material is listed and represented by habitus photographs.As a result, 9 monotypic species of Molopsis are recognized. Tapinopterus rebellis (Chaudoir, 1868) and T. rebelliskumanensis (Reitter, 1884), formerly assigned to Molopsis are recognized as outgroup-taxa. The following new speciesare described: T. (Molopsis) aenigmaticus sp.nov. (Asian Turkey, „Armenia“ imprecise locality), T. (Molopsis) chaudoirisp.nov. (Asian Turkey, Bursa Villayet, Uludağ Mt., Sakarya (Adapazari) Villayet, Gökdağ Mt.), T. (Molopsis) oyukluensissp.nov. (Asian Turkey, Konya Villayet, Fasihan Pass), and T. (Molopsis) relegatus sp.nov. (Asian Turkey, Bolu Villayet,Mengen env., Dorukhan Pass). The following nomenclatural acts are also proposed: Tapinopterus (Molopsis) molopinus (Chaudoir, 1868) = Tapinopterus (Molopsis) dipojranus brussanus Straneo, 1984, syn.nov.; Tapinopterus (Molopsis)References
Allen, R.T. & Ball, G.E. (1980) Synopsis of Mexican taxa of the Loxandrus series (Coleoptera: Carabidae: Pterostichini). Transactions of the American Entomological Society, 105, 481–576.
Apfelbeck, V. (1904) Die Käferfauna der Balkanhalbinsel, mit Berücksichtigung Klein-Asien und der Insel Kreta. Erstes Band: Familienreiche Caraboidea. Berlin, R. Friedländer & Sohn, IX + 1–422.
Ball, G.E. & Shpeley, D. (1983) The species of eucheloid Pericalina: classification and evolutionary considerations (Coleoptera: Carabidae: Lebiini). Canadian Entomologist, 115, 743–806.
Bodemeyer, E. (1905) Quer durch Klein-Asien in den Bulghar-Dagh. Druck und Verlag, Emmendingen, V + 1–169.
Bousquet, Y. (1999) Supraspecific classification of the Nearctic Pterostichini (Coleoptera: Carabidae). Association des Entomologistes Amateurs du Québec, Supplement 9, 1–292.
Bousquet, Y. (2003) Pterostichini. In: Löbl & Smetana (eds.). Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata – Myxophaga – Adephaga. Stenstrup: Apollo Books, pp. 469–521.
Brandmayr, P. (1977) Ricerche etologiche e morfofunzionali sulle cure parentali in Carabidi Pterostichini (Coleoptera: Carabidae, Pterostichinae). Redia, 60, 275–316.
Brandmayr, P. (1978) Revisione del gen. Abax Bonelli (Coleoptera, Carabidae) su base morfo-funzionale, etologica ed ecologica. Atti del Congresso Nazionale Italiano di Entomologia, 11, 65–72.
Brandmayr, P. & Zetto Brandmayr, T. (1978) Parental ethology of pterostichine carabids (Coleoptera): The relationships between ovipositor morphology and the evolution of reproductive behaviour. Monitore Zoologico Italiano (N.S.), 12, 61–62.
Brandmayr, P. & Zetto Brandmayr, T. (1979) The evolution of parental care phenomena in Pterostichinae ground beetles with special reference to the genera Abax and Molops (Coleoptera, Carabidae). Miscellaneous Papers Landbouwhogeschool Wageningen, 18, 135–149.
Breit, J. (1914) Neue Coleopterenformen aus Süd-Europa. Coleopterologische Rundschau, 3, 50–62.
Casale, A. & Ribera, I. (2008) Are Molopina of the Euro-Mediterranean region related to the Madagascan, South African and Australian Pterostichini? (Coleoptera, Carabidae). Biogeographia, 29, 33–44.
Casale, A. & Vigna Taglianti, A. (1999) Caraboid beetle (excl. Cicindelidae) of Anatolia, and their biogeographical significance (Coleoptera, Caraboidea). Biogeographia, 20, 277– 406.
Chaudoir, M. de (1850) Supplément á la faune des carabiques de la Russie. Bulletin de la Société Impériale des Nauralistes de Moscou, 23, 62–206.
Chaudoir, M. de (1868) Description d´espéces nouvelles ou peu connues de Feronia d´Europe et du bassin méditerranéen. L´Abeille, Mémoires d’Entomologie, 5, 219–260.
Chaudoir, M. de (1876) Synonymische Bemerkungen. Deutsche Entomologische Zeitschrift, 20, 345–348.
Csiki, E. (1930) Carabidae: Harpalinae IV (pars 112). In: Junk W. & Schenkling S. (eds.): Coleopterorum Catalogus. Vol. II. Carabidae II. Berlin: W. Junk, pp. 529–737.
Fairmaire, L. (1859) Miscellanea Entomologica. Troisiéme Partie. Annales de la Société Entomologique de France (3) 7, 21–64.
Fairmaire, L. (1866) Notice sur les Coléoptéres récoltés par M. J. Lédérer sur le Bosz-Dagh (Asie Mineure). Annales de la Société Entomologique de France (4) 6, 249–280.
Ganglbauer, L. (1889) Berichte über von E. v. Oertzen im Jahre 1887 in Griechenland u. Klein-Asien gesammelten Coleopteren. V. Carabidae (Tapinopterus, Ditomus), Lamellicornia, Buprestidae, Throscidae, Elateridae, Meloidae, Oedemeridae, Cerambycidae. Deutsche Entomologische Zeitschrift, 1889, 49–57.
Gemminger, M. & Harold, E. von. (1868) Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae-Carabidae. Monachii: E.H. Gummi, xxxvi + 424 + 8 upp. pp.
Giachino, P.M., Picciau, L., Vailati, D. & Casale, A. (in press) Taxonomic revision, phylogeny and biogeography of the European and Aegean species of Tapinopterus Schaum, 1858 (Coleoptera Carabidae). Memoirs on Biodiversity.
Giachino, P.M. & Casale, A. (1983) Sistematica e morfologia genitale di alcuni Pterostichus italiani (Coleoptera, Carabidae). Bolletino del Museo Regionale di Scienze Naturali Torino, 1 (2), 417–450.
Giachino, P.M. & Sciaky, R. (1991) Valore sistematico delle strutture genitali femminili in Pterostichinae (Coleoptera: Carabidae). Atti XVI Congresso nazionale italiano Entomologia., Bari-Martina Franca (TA), 23/18, 885–892.
Guéorguiev, B. (2007) Annotated catalogue of the carabid beetles of Albania (Coleoptera: Carabidae). Pensoft Series Faunistica No 64, Sofia-Moscow, 243 pp.
Guéorguiev, B. & Lobo, J. M. (2006) Adephagous beetles (Insecta: Coleoptera: Adephaga) in the Western Rhodopes (Bulgaria and Greece). In: Beron P. (ed). Biodiversity of Bulgaria. 3. Biodiversity of Western Rhodopes (Bulgaria and Greece) I. Pensoft & Nat. Mus. Natur. Hist., Sofia, pp. 283–346.
Guéorguiev, B. & Lohaj, R. (2008) Studies on genus Speluncarius, with description of a new subgenus and notes on the systematic position of S. (Hypogium) albanicus (Coleoptera: Carabidae: Pterostichini). In: L. Penev, T. Erwin & T. Assmann (Eds). 2008. Back to the Roots and Back to the Future. Towards a New Synthesys amongst Taxonomic, Ecological and Biogeographical Apporoaches in Carabidology. Proceedings of the XIII European Carabidologists Meeting, Blagoevgrad, August 20–24, 2007. Pensoft, Sofia-Moscow, pp. 125–141.
Habu, A. (1981a) Female genitalia of Pterostichini species mainly from Japan (I) (Coleoptera, Carabidae). Entomological Review of Japan, 35, 77–99.
Habu, A. (1981b) Female genitalia of Pterostichini species mainly from Japan (II) (Coleoptera, Carabidae). Entomological Review of Japan, 36, 33–53.
Heyden, L. von., Reitter, E. & Weise, J. (1891) Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. Familie Carabidae. Berlin, R. Friedländer & Sohn, viii + 1–420.
Heyden, L. von., Reitter, E. & Weise, J. (1906) Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. Editio secunda. Berlin, R. Friedländer & Sohn; Pascau, Edmund Reitter; Caen, Revue d’Entomologie. 4 p. + 744 columns.
Horne, P.A. (1990) Parental care in Notonomus Chaudoir (Coleoptera: Carabidae: Pterostichinae). Australian Entomological Magazine, 17, 65–69.
Hůrka, K. (1996) Carabidae of the Czech and Slovak Republics. Kabourek, Zlín, 565 pp.
Jakobson, G.G. (1907) Fasc. 5. In: Zhuki Rossii I Zapadnoi Evropy. Sankt-Petersburg: A.F. Devrien, 1024 pp. + lxxxiii pl., pp. 321–400.
Jeanne, C. (2005) Un Molopsis Schatzmayr nouveau d’Anatolie méridionale (Coleoptera, Carabidae). Nouvelle Revue d´Entomologie (N.S.), 21, 383–385.
Jedlička, A. (1963) Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau, 40–41, 16–22.
Kirschenhofer, E. (1991) Zwei neue Carabiden aus Kurdistan: Tapinopterus jaechi sp. nov. und Stenochlaenius jaechi sp. nov. (Coleoptera: Carabidae). Koleopterologische Rundschau, 61, 1–10.
Kraatz, G. (1875) Drei neue europäische Pterostichus Bon. Deutsche Entomologische Zeitschrift, 19, 417–419.
Lorenz, W. (1998) Nomina Carabidarum – a directory of the scientific names of ground beetles (Insecta, Coleoptera “Geadephaga”: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Tutzing: W. Lorenz, iv + 937 pp.
Lorenz, W. (2005) Systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae & Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. Tutzing: Lorenz Publishing, 530 pp.
Lutshnik, V.N. (1922) O Platysmatina (Coleoptera. Carabidae). Trudy Stavropolskogo Selskokhozaystvennogo Instituta, 1, 67–79.
Mařan, J. (1940) Nové formy podrodu Tapinopterus Schaum. De novis parumque cognitis subgeneris Tapinopterus Schaum formis. Coleoptera – Carabidae. Sborník entomologického Oddělení Národního Musea v Praze, 18, 56–59.
Marseul, S.A. de (1880) Nouveau répertoire contenant les descriptions des espèces de coléoptères de l’ancien-monde publiées isolément ou en langues étrangères, en dehors des monographies ou traités spéciaux et de l’Abeille. L’Abeille, Journal d’Entomologie, 19, 1–526.
Marseul, S.A. de (1882) Nouveau répertoire contenant les descriptions des espèces de coléoptères de l’ancien-monde publiées isolément ou en langues étrangères, en dehors des monographies ou traités spéciaux et de l’Abeille. L’Abeille, Journal d’Entomologie, 20, 1–196.
Mlynář, Z. (1977) Revision der Arten und Unterarten der Gattung Molops Bon. (s.str.) (Coleoptera, Carabidae). Folia Entomologica Hungarica. Rovartani Közlemények (N.S., suppl.), 30, 3–150.
Müller, J. [G.] (1923) Risultati scientifici della spedizione Ravasini-Lona in Albania. V. Carabidae. Bolletino della Societa Entomologica Italiana, 55, 105–112.
Müller, J [G.] (1932) Neue und wenig bekannte Carabiden von Leo Weirather auf der Balkanhalbinsel und in Kleinasien gesammelt. Koleopterologische Rundschau, 17, 209–223.
Müller, J. [G.] (1937) Note su alcuni carabidi della Balcania e della regione mediteranea. Atti del Museo Civico di Storia Naturale di Trieste, 13, 119–134.
Reiche, L. & Saulcy, F. (1855) Espèces nouvelles ou peu connues de coléoptères, recueillies par M. F. de Saulcy, membre de l’Institut, dans son voyage en Orient, et décrites par M. M. L. Reiche et Félicien de Saulcy. Annales de la Société Entomologique de France, 3, 561–645.
Schatzmayr, A. (1929) I Pterostichus italiani. Memoire della Societa Entomologica Italiana, 8, 145–339.
Schatzmayr, A. (1942) Bestimmungs-Tabellen europäischer Käfer. (8 Stück - II. Teil) II. Fam. Carabidae. Subfam. Pterostichinae. 65. Gattungen: Pterostichus Bon. u. Tapinopterus Schaum. Bestimmungstabellen der europäischen und nordafricanischen Pterostichus- und Tapinopterus-Arten. (I. Teil und Schluß). Wien: Zoologisch-botanische Gessellschaft, 1–80.
Schatzmayr, A. (1943) Bestimmungs-Tabellen europäischer Käfer. (8 Stück - II. Teil) II. Fam. Carabidae. Subfam. Pterostichinae. 65. Gattungen: Pterostichus Bon. u. Tapinopterus Schaum. Bestimmungstabellen der europäischen und nordafricanischen Pterostichus- und Tapinopterus-Arten. (II. Teil und Schluß). Wien: Zoologisch-botanische Gessellschaft, 81–144.
Seidlitz, G.C.M. von (1887) Fauna Baltica. Die Käfer (Coleoptera) der deutschen Ostseeprovinzen Russlands. Zweite neu bearbeitete Auflage. Königsberg: Hartungsche Verlagsdruckerei, xl + 16 [Gattungen] + 96 pp. [Arten].
Seidlitz, G.C.M. von (1888) Fauna Transsylvanica. Die Käfer (Coleoptera) Siebenbürgens. (1888–1891). Königsberg: Hartung, xvi + 1–914.
Straneo, S.L. (1936) Note sui Platysma paleartici. 3a. Annali del Museo Civico di Storia Naturale “Giacomo Doria”, 59, 145–157.
Straneo, S.L. (1986) Nuovi Tapinopterus dell´Anatolia (Coleoptera, Carabidae). Fragmenta Entomologica, 19, 119–127.
Sturm, J. (1843) Catalog der Käfer-Sammlung. Nűrnberg, xii + 386 pp + 6 pl.
Thiele, H.U. (1977) Carabid beetles in their environment. Zoophysiology and Ecology, 10. New York, Springer-Verlag, 369 pp.