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Type: Article
Published: 2003-11-17
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Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes

Division of Entomology, Natural History Museum, University of Kansas, Lawrence, KS 66045-7523, U.S.A. and Department of Entomology, St. Petersburg State University, Universitetskaya nab., 7/9, St. Petersburg 199034, Russia
Coleoptera Staphylinidae Aleocharinae Holarctic Nearctic taxonomy nomenclature synonymy

Abstract

Based on my revision of the types of Nearctic aleocharine staphylinids numerous nomenclatural and taxonomic changes are proposed. The following taxa are transferred: Acrimea acerba Casey, 1911a to Tinotus Sharp, 1883; Moluciba grandipennis Casey, 1911a to Oxypoda Mannerheim, 1830; Paradilacra Bernhauer, 1909 to subtribe Tachyusina Thomson, 1859 of tribe Oxypodini Thomson, 1859; Atheta angusticornis Bernhauer, 1907 to Boreophilia Benick, 1973; At. coriaria (Kraatz, 1856) (originally described in Homalota Mannerheim, 1830) to Dalotia Casey, 1910a; Homalota ambigua Erichson, 1839 to Strigota Casey, 1910a; Pseudousipalia microptera Lohse in Lohse et al., 1990 to Emmelostiba Pace, 1982; and Sableta brittoni Casey, 1911a to Thamiaraea Thomson, 1858. The following new synonymies are established: Acrimea Casey, 1911a with Tinotus Sharp 1883; Acri. fimbriata Casey, 1911a with Ti. trisectus Casey, 1906; Acri. resecta Casey, 1911a with Ti. acerbus (Casey, 1911a); Ancillota Casey, 1910a and Moluciba Casey, 1911a with Oxypoda Mannerheim, 1830; Anc. sollemnis Casey, 1910a, O. vetula Casey, 1911a, O. neptis Casey, 1911a and O. schaefferi Notman, 1920 with O. amica Casey, 1906; Atheta lanei Casey, 1910a and At. nomadica Casey, 1910a with At. graminicola (Gravenhorst, 1806); Homalota polita Melsheimer, 1844, At. disjuncta Casey, 1910a, At. replicans Casey, 1910a, At. spadix Casey, 1910a and At. bucolica Casey, 1910a with At. aemula (Erichson, 1839); At. innocens Casey, 1910a, At. achromata Casey, 1911a and At. profecta Casey, 1911a with At. keeni Casey, 1910a; Dimetrota dempsterensis Lohse in Lohse et al., 1990 with At. prudhoensis (Lohse in Lohse et al., 1990); At. aperta Casey, 1910a, At. wrangelica Casey, 1911a, At. morbosa Casey, 1911a, At. intacta Casey, 1911a and At. alaskana Casey, 1911a with At. picipennis (Mannerheim, 1843); At. leviceps Casey, 1910a, Dimetrota sectator Casey, 1910a, Dim. retrusa Casey, 1910a, Datomicra hebescens Casey, 1910a, Dat. insolida Casey, 1910a, Dat. pellax Casey, 1910a and Pseudota vana Casey, 1911a with At. hampshirensis Bernhauer, 1909; At. repexa Casey, 1911a with At. brumalis Casey, 1910a; At. querula Casey, 1910a, At. socors Casey, 1911a, Dimetrota resima Casey, 1910a, Dim. vigilans Casey, 1910a, Dim. immerita Casey, 1911a, Dim. incredula Casey, 1911a, Dim. opinata Casey, 1911a and Dim. cerebrosa Casey, 1911a with Atheta fenyesi Bernhauer, 1907; At. crassicornis virginica Bernhauer, 1907, At. rhodeana Casey, 1910a, At. capella Casey, 1910a, At. ducens Casey, 1910a, At. temperans Casey, 1910a, At. logica Casey, 1910a, At. tradita Casey, 1911a, At. fenisex Casey, 1911a, At. vierecki Casey, 1911a, At. auguralis Casey, 1911a and At. bifaria Casey, 1911a with At. modesta (Melsheimer, 1844); At. comitata Casey, 1910a, At. gnoma Casey, 1910a, At. elota Casey, 1910a, At. insidiosa Casey, 1910a, Pseudota puricula Casey, 1911a, At. candidula Casey, 1911a, At. diffisa Casey, 1911a, At. nata Casey, 1911a, At. modiella Casey, 1911a and At. vacillans Casey, 1911a with At. frosti Bernhauer, 1909; At. mollicula Casey, 1910a, Sableta phrenetica Casey, 1910a, At. callens Casey, 1911a, At. franklini Casey, 1911a and At. postulans Casey, 1911a with At. ventricosa Bernhauer, 1907; At. cephalina Casey, 1910a, At. nympha Casey, 1910a, At. discreta Casey, 1910a: 42 (nec Casey, 1893, nec Casey, 1910a: 79), Pseudota dissensa Casey, 1910a, At. villica Casey, 1911a and At. disca Moore & Legner, 1975 with At. klagesi Bernhauer,1909; At. citata Casey, 1910a, At. evecta Casey, 1910a, At. propitia Casey, 1911a, At. palpator Casey, 1911a, At. burra Casey, 1911a and At. nacta Casey, 1911a with At. annexa Casey, 1910a; At. sumpta Casey, 1911a with At. concessa Casey, 1911a; At. punctata Blatchley, 1910, Synaptina merica Casey, 1910a and Sy. consonens Casey, 1910a with At. festinans (Erichson, 1839); Boreostiba hudsonica Lohse in Lohse et al., 1990 with At. parvipennis Bernhauer, 1907; Boreophilia chillcotti Lohse in Lohse et al., 1990 with At. blatchleyi Bernhauer & Scheerpeltz, 1926; Datomicra decolorata Casey, 1910a, Dat. inopia Casey, 1910a, Dat. schematica Casey, 1910a and Dat. stilla Casey, 1910a with At. dadopora Thomson, 1867; Boreophilia caseyiana Lohse in Lohse et al., 1990 with Boreophilia nomensis (Casey, 1910a); Metaxya plutonica Casey, 1910a with Boreophilia angusticornis (Bernhauer, 1907); Boreostiba lamellifera Lohse in Lohse et al., 1990 with Boreostiba frigida (J. Sahlberg, 1880); At. laurentiana Blatchley, 1910 with Aloconota sulcifrons (Stephens, 1832); At. immigrans Easton, 1971 with Adota maritima (Mannerheim, 1843); Pseudota miscella Casey, 1910a, Dimetrota pectorina Casey, 1910a and Dim. crucialis Casey, 1910a with Dalotia coriaria (Kraatz, 1856); Dimetrota revoluta Casey, 1910a and Datomicra vaciva Casey, 1910a with Dochmonota rudiventris (Eppelsheim, 1886); At. insolens Casey, 1910a, Dimetrota resplendens Casey, 1910a and At. apposita Casey, 1911a with Liogluta nitens (Mäklin in Mannerheim, 1852); Achromata Casey, 1893 with Mocyta Mulsant & Rey, 1874a; Achromata fusiformis Casey, 1893, Dimetrota nuptalis Casey, 1910a, Acrotona lividula Casey, 1910a and Acro. adjuvans Casey, 1910a with Mocyta fungi (Gravenhorst, 1806); Acrotona digesta Casey, 1910a, Acro. severa Casey, 1910a, Acro. shastanica Casey, 1910a, Acro. prudens Casey, 1910a, Acro. ardelio Casey, 1910a, Acro. renoica Casey, 1910a and Acro. malaca Casey, 1910a with Mocyta breviuscula (Mäklin in Mannerheim, 1852); Eustrigota Casey, 1911a with Acrotona Thomson, 1859; Colpodota inceptor Casey, 1910a, C. abdicans Casey, 1910a, C. repentina Casey, 1910a, C. laxella Casey, 1910a, C. pupilla Casey, 1910a and Strigota seclusa Casey, 1911a with Acrotona sonomana (Casey, 1910a); Anaduosternum Notman, 1922 with Strigota Casey, 1910a; Strigota oppidana Casey, 1910a, St. gnava Casey, 1910a, St. verecunda Casey, 1910a, St. assueta Casey, 1910a, St. mediocris Casey, 1910a, St. vapida Casey, 1910a, St. inculta Casey, 1910a, St. placata Casey, 1910a, St. recta Casey, 1911a, Anaduosternum brevipennis Notman, 1922 and Atheta notmani Moore & Legner, 1975 with St. ambigua (Erichson, 1839); Pseudousipalia Lohse in Lohse et al., 1990 with Emmelostiba Pace, 1982; Fusalia Casey, 1911a with Thamiaraea Thomson, 1858; Th. lira Hoebeke, 1988 and Th. paralira Hoebeke, 1994 with Th. brittoni (Casey, 1911a); Drusilla cavicollis Casey, 1906 with Dr. canaliculata (Fabricius, 1787); Leptusa laticollis Notman, 1921 with Le. brevicollis Casey, 1893; Sipalia fontana Casey, 1911a and Pasilia virginica Casey, 1911a with Leptusa elegans Blatchley, 1910; and Pseudota cornicula Casey, 1911a with Placusa vaga Casey, 1911a. Tinotus pallidus Casey, 1911a is removed from synonymy with Ti. caviceps Casey, 1893 and is placed in synonymy with Ti. trisectus Casey, 1906. Atheta granulata (Mannerheim, 1846) (originally described in Homalota) is considered to be a synonym of At. graminicola (Gravenhorst, 1806), which has Holarctic (circumboreal) distribution. Atheta keeni Casey, 1910a is the valid name for At. vasta sensu Klimaszewski & Winchester, 2002. Boreostiba frigida (J. Sahlberg, 1880) is removed from synonymy with Boreostiba sibirica (Mäklin, 1880) and is considered to be a valid species. Leptusa obscura Blatchley, 1910 is removed from synonymy with Le. canonica Casey, 1906 and is considered to be a valid species. The following synonymies are confirmed: Devia congruens (Casey, 1893) with De. prospera (Erichson, 1839); Paradilacra persola Casey, 1910a, Pa. willametta Casey, 1910a, Pa. uintana Casey, 1910a, Pa. glenorica Casey, 1910a, Pa. symbolica Casey, 1911a, Pa. erebea Casey, 1911a, Pa. subaequa Casey, 1911a, Pa. sinistra Casey, 1911a, Pa. memnonia Casey, 1911a, Pa. vulgatulaCasey, 1911a and Pa. deserticola Casey, 1911a with Pa. densissima (Bernhauer, 1909); Atheta carlottae Casey, 1910a with At. picipennis (Mannerheim, 1843); At. maeklini Fenyes, 1820 (replacement name for Homalota moesta Mäklin in Mannerheim, 1852) with At. hampshirensis Bernhauer, 1909; At. fontis Casey, 1911a with At. pennsylvanica Bernhauer, 1907, Leptusa tricolor Casey, 1906, Le. nebulosa Casey, 1911a and Le. iowensis Casey, 1911a with Le. canonica Casey, 1906; Le. seminitens Casey, 1893 with Le. opaca Casey, 1893. Atheta picipennis (Mannerheim, 1843) (ex Homalota) is a nomen protectum and At. picipennis (Stephens, 1832) (ex Aleochara; a junior synonym of At. amicula (Stephens, 1832)) is a nomen oblitum. Lectotypes are designated for Acrimea fimbriata Casey, 1911a, Acri. acerba Casey, 1911a, Acri. resecta Casey, 1911a, Ancillota sollemnis Casey, 1910a, Oxypoda amica Casey, 1906, O. vetula Casey, 1911a, O. neptis Casey, 1911a, O. schaefferi Notman, 1920, O. prospera Erichson, 1839, O. congruens Casey, 1893, Atheta densissima Bernhauer, 1909, At. lanei Casey, 1910a, At. nomadica Casey, 1910a, At. disjuncta Casey, 1910a, At. replicans Casey, 1910a, At. spadix Casey, 1910a, At. keeni Casey, 1910a, At. innocens Casey, 1910a, At. achromata Casey, 1911a, At. profecta Casey, 1911a, At. carlottae Casey, 1910a, At. aperta Casey, 1910a, At. morbosa Casey, 1911a, At. alaskana Casey, 1911a, At. altaica Bernhauer, 1901, At. leviceps Casey, 1910a, At. hampshirensis Bernhauer, 1909, At. brumalis Casey, 1910a, At. repexa Casey, 1911a, At. fenyesi Bernhauer, 1907, At. querula Casey, 1910a, At. socors Casey, 1911a, At. crassicornis var. virginica Bernhauer, 1907, At. rhodeana Casey, 1910a, At. capella Casey, 1910a, At. ducens Casey, 1910a, At. temperans Casey, 1910a, At. logica Casey, 1911a, At. tradita Casey, 1911a, At. fenisex Casey, 1911a, At. vierecki Casey, 1911a, At. auguralis Casey, 1911a, At. bifaria Casey, 1911a, At. frosti Bernhauer, 1907, At. comitata Casey, 1910a, At. gnoma Casey, 1910a, At. elota Casey, 1910a, At. insidiosa Casey, 1910a, At. candidula Casey, 1911a, At. diffisa Casey, 1911a, At. nata Casey, 1911a, At. modiella Casey, 1911a, At. vacillans Casey, 1911a, At. ventricosa Bernhauer, 1907, At. mollicula Casey, 1910a, At. callens Casey, 1911a, At. franklini Casey, 1911a, At. postulans Casey, 1911a, At. klagesi Bernhauer, 1909, Atheta cephalina Casey, 1910a, At. nympha Casey, 1910a, At. discreta Casey, 1910a: 42 (nec Casey, 1893, nec Casey, 1910a: 79), At. citata Casey, 1910a, At. evecta Casey, 1910a, At. propitia Casey, 1911a, At. palpator Casey, 1911a, At. burra Casey, 1911a, At. nacta Casey, 1911a, At. concessa Casey, 1911a, At. punctata Blatchley, 1910, At. parvipennis Bernhauer, 1907, At. caviceps Blatchley, 1910, At. pennsylvanica Bernhauer, 1907, At. angusticornis Bernhauer, 1907, Atheta laurentiana Blatchley, 1910, At. insolens Casey, 1910a, At. apposita Casey, 1911a, Paradilacra persola Casey, 1910a, Pa. willametta Casey, 1910a, Pa. uintana Casey, 1910a, Pa. glenorica Casey, 1910a, Pa. symbolica Casey, 1911a, Pa. erebea Casey, 1911a, Pa. subaequa Casey, 1911a, Pa. sinistra Casey, 1911a, Pa. memnonia Casey, 1911a, Pa. deserticola Casey, 1911a, Aleochara graminicola Gravenhorst, 1806, Ale. nigritula Gravenhorst, 1802, Homalota aemula Erichson, 1839, H. polita Melsheimer, 1844, H. modesta Melsheimer, 1844, H. sodalis Erichson, 1837, H. festinans Erichson, 1839, H. ambigua Erichson, 1839, Dimetrota sectator Casey, 1910a, Dim. retrusa Casey, 1910a, Dim. resima Casey, 1910a, Dim. vigilans Casey, 1910a, Dim. incredula Casey, 1911a, Dim. opinata Casey, 1911a, Dim. cerebrosa Casey, 1911a, Dim. pectorina Casey, 1910a, Dim. crucialis Casey, 1910a, Datomicra hebescens Casey, 1910a, Dat. insolida Casey, 1910a, Dat. decolorata Casey, 1910a, Dat. inopia Casey, 1910a, Dat. schematica Casey, 1910a, Dat. stilla Casey, 1910a, Dat. vaciva Casey, 1910a, Pseudota vana Casey, 1911a, Ps. puricula Casey, 1911a, Ps. dissensa Casey, 1910a, Ps. miscella Casey, 1910a, Sableta phrenetica Casey, 1910a, Sa. brittoni Casey, 1911a, Synaptina merica Casey, 1910a, Sy. consonens Casey, 1910a, Metaxya plutonica Casey, 1910a, Acrotona lividula Casey, 1910a, Acro. adjuvans Casey, 1910a, Acro. digesta Casey, 1910a, Acro. severa Casey, 1910a, Acro. shastanicaCasey, 1910a, Acro. prudens Casey, 1910a, Acro. ardelio Casey, 1910a, Acro. renoica Casey, 1910a, Acro. malaca Casey, 1910a, Colpodota sonomana Casey, 1910a, C. inceptor Casey, 1910a, C. abdicans Casey, 1910a, C. repentina Casey, 1910a, C. laxella Casey, 1910a, C. pupilla Casey, 1911a, Strigota seclusa Casey, 1911a, St. oppidana Casey, 1910a, St. gnava Casey, 1910a, St. verecunda Casey, 1910a, St. assueta Casey, 1910a, St. mediocris Casey, 1910a, St. vapida Casey, 1910a, St. inculta Casey, 1910a, St. placata Casey, 1910a, St. recta Casey, 1911a, Leptusa seminitens Casey, 1893, Le. tricolor Casey, 1906, Le. nebulosa Casey, 1911a, Le. obscura Blatchley, 1910, Le. elegans Blatchley, 1910, Ulitusa pusio Casey, 1906 and Sipalia fontana Casey, 1911a. Oxypoda acuminata (Stephens, 1832) and Atheta dadopora Thomson, 1867 are reported from North America for the first time. North American records of Atheta altaica Bernhauer, 1901 are confirmed.

References

  1. Alexandrovitch, O.R., Lopatin, I.K., Pisanenko, A.D., Tsinkevitch, V.A. & Snitko, S.M. (1996) A Catalogue of Coleoptera (Insecta) of Belarus. FFR RB, Minsk, 103 pp.

    Aubé, C. (1850) Description de quelques insectes Coléoptères appartenents à l’Europe et à l’Algérie. Annales de la Société entomologique de France, 8, 299–346.

    Baudi di Selve, F. (1848) Alcune specie nuove di stafilini. Studi Entomologici, 1(2), 113–148.

    Benick, G. (1954) Revision der Untergattung Aloconota C.G.Thoms. (Gattung Atheta, Staph.). Entomologische Blätter, 50, 133–174.

    Benick, G. (1973) Die von Prof. Dr. Z. Kaszab in der Mongolei gesammelten Arten der Subfamilie Athetae (Col. Staphyl.). Nouvelle Revue d’Entomologie, 3(4), 211–217.

    Benick, G. (1982) Arten der Unterfamilie Athetae aus der Mongolei (Coleoptera, Staphylinidae), IV. Annales Historico-Naturales Musei Nationalis Hungarici, 74, 93–114.

    Benick, G. & Lohse, G.A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K.W. & Lohse, G.A. (Eds.), Die Käfer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 72–220.

    Bernhauer, M. (1901) Neue Staphyliniden aus Centralasien. Verhandlungen der k. k. zoologischbotanischen Gesellschaft in Wien, 51, 106–115.

    Bernhauer, M. (1905) 13. Folge neuer Staphyliniden der paläarktischen Fauna, nebst Bemerkungen. Verhandlungen der k. k. zoologisch-botanischen Gesellschaft in Wien, 55, 580–596.

    Bernhauer, M. (1906) Neue Aleocharinen aus Nordamerika. (II. Teil). Deutsche Entomologische Zeitschrift, 1906(2), 337–348.

    Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stück.). Deutsche Entomologische Zeitschrift, 1907(4), 381–405.

    Bernhauer, M. (1909) Neue Aleocharini aus Nordamerika. (Col.) (4.Stück.) Deutsche Entomologische Zeitschrift, 1909(4), 515–528.

    Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499–988.

    Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. U.S. National Museum Bulletin, 200, 1–483.

    Blackwelder, R. E. (1973) Checklist of the Staphylinidae of Canada, United States, Mexico, Central America and the West Indies. North American Beetle Fauna Project. Family no. 15, complete (Yellow Version). Biological Research Institute of America, Inc., Sienna College, Loudonville, New York, 165 pp.

    Blatchley, W.S. (1910) An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana. The Nature Publishing Co., Indianapolis, 1358 pp.

    Bordoni, A. (1973) I Coleotteri Stafilinidi delle isole circumsiciliane. Lavori della Società Italiana di Biogeografia, 3 [1972], 651–754.

    Brundin, L. (1940) Studien über die Atheta-Untergattung Oreostiba Ganglb. (Col. Staphylinidae). Entomologisk Tidskrift, 61(1–4), 56–130, Pl. 1–18.

    Burakowski, B., Mroczkowski, M. & Stefa½ska, J. (1981) Katalog fauny Polski — Catalogus faunae Poloniae. Cz“у 23, tom 8. Chrz_szcze — Coleoptera. Kusakowate — Staphylinidae, cz“у, 3 Aleocharinae. Pa½stwowe Widawnictwo Naukowe, Warszawa, 330 pp.

    Casey, T. L. (1893) Coleopterological Notices. V. Annals of the New York Academy of Sciences, 7, 281–606.

    Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16(6), 125–434.

    Casey, T. L. (1910a) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp.1–183.

    Casey, T. L. (1910b) Synonymical and other notes on Coleoptera. The Canadian Entomologist, 42(4), 105–114.

    Casey, T. L. (1911a) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, pp. 1–245.

    Casey, T. L. (1911b) Notes on the Coccinellidae with some general remarks and synonymy. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, pp. 246–254.

    Derunkov, A.V. & Melke, A. (2001) Familia Staphylinidae. In: Gutowski, J.M. & Jaroszewicz, B. (Eds.), Catalogue of the fauna of Bia»owieóa Primeval Forest. IBL, Warszawa, pp. 133 – 147.

    Dillwyn, L.W. (1829) Memoranda relating to coleopterous insects, found in the neighborhood of Swansea. Swansea, 75 pp.

    Easton, A.M. (1970) Atheta boletophila Thoms. (Col. Staphylinidae) new to Britain. The Entomologist's Monthly Magazine, 105(1262–1264) [1969], 197–198.

    Easton, A.M. (1971) A new British Atheta (Col., Staphylinidae) in a new subgenus. The Entomologist's Monthly Magazine, 107, 24–26.

    Eppelsheim, E. (1886) Neue Staphylinen vom Amur. Deutsche Entomologische Zeitschrift, 30(1), 33–46.

    Erichson, W.F. (1837) Die Käfer der Mark Brandenburg. Erster Band. Erste Abtheilung. F.H.Morin, Berlin, pp.i–viii, 1–384.

    Erichson, W.F. (1839) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp. i–viii, 1–400.

    Erichson, W.F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp. 401–954.

    Fabricius, J.C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom. 1. Christ. Gottl. Proft., Hafnia [Copenhagen], xx+348 pp.

    Fabricius, J.C. (1793) Entomologiae systematicae, emendatae et auctae. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tom I. Pars II. Christ. Gottl. Proft., Hafnia [Copenhagen], 538 pp.

    Fenyes, A. (1907) The Aleocharinae by Casey. Entomological News, 18(2), 60–61.

    Fenyes, A. (1918) Coleoptera: Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1–110.

    Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111–414.

    Fenyes, A. (1921) New genera and species of Aleocharinae with a polytomic synopsis of the tribes. Bulletin of the Museum of Comparative Zoology, 65, 17–36.

    FitzGerald, W. (1962) Casey locality code. Manuscript kept with Casey collection at the National Museum of National History, Washington, D.C.

    Fleming, J. (1821) Insecta. In: Supplement to the fourth, fifth and sixth editions of the Encyclopedia Britannica, with preliminary dissertations on the history of sciences. Vol. 5. Archibald Constable and Company, Edinburgh, pp. 41–56.

    Frank, J.H. (1980) Atheta coriaria (Kraatz) (Aleocharinae) and Sunius confluentus (Say) (Paederinae) in Florida (Coleoptera, Staphylinidae). The Coleopterists Bulletin, 34(4), 388.

    Ganglbauer, L. (1895) Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des französischen und italienischen Alpengebietes. Zweiter Band. Familienreihe Staphylinoidea. 1. Theil: Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, vi+881 pp.

    Gemminger, M. & Harold, E. von (1868) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom II. E.H.Gummi, Monachii [Munich], pp. 425–752 + 6pp.

    Gensicke, F. (1960) Zur Kenntnis der Nestfauna einiger Muridenarten in der Umgebung von Greifswald. Wissenschaftliche Zeitschrift der Ernst-Moeritz-Arndt-Universität Greifswald, 9, 189– 197.

    Gravenhorst, J.L.C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. C. Reichard, Brunswick, xxvi + 207 pp.

    Gravenhorst, J.L.C. (1806) Monographia Coleopterorum Micropterorum. Henrich Dieterich, Göttingen, xvi+248 pp.

    Gusarov, V.I. (2002a) A revision of Nearctic species of the genus Tomoglossa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa, 30, 1–19.

    Gusarov, V.I. (2002b) A revision of the genus Microlia Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Hoplandriini). Zootaxa, 34, 1–24.

    Gusarov, V.I. (2002c) A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa, 81, 1–88.

    Gusarov, V.I. (2002d) A revision of Nearctic species of the genus Earota Mulsant & Rey, 1874 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa, 92, 1–16.

    Gusarov, V.I. (2002e) A revision of Nearctic species of the genus Tropimenelytron Pace, 1983 (Coleoptera: Staphylinidae: Aleocharinae), a new genus for North America. Zootaxa, 114, 1– 24.

    Gusarov, V.I. (2003a) A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa, 142, 1–102.

    Gusarov, V.I. (2003b) A revision of the genus Goniusa Casey, 1906 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa, 164, 1–20.

    Gusarov, V.I. (2003c) A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa, 185, 1–35.

    Gusarov, V.I. (2003d) A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa, 239, 1–43.

    Gusarov, V.I. (in press, a) A revision of the Nearctic species of the genus Halobrecta Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa.

    Gusarov, V.I. (in press, b) A revision of the genus Lypoglossa Fenyes, 1918 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa.

    Gusarov, V.I. & Herman, L.H. (2003) Leptusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae): designation of the type species. Entomologische Blätter für Biologie und Systematik der Käfer, 98 [2002], 115–119.

    Gyllenhal, L. (1827) Insecta Suecica descripta. Classis I. Coleoptera sive Eleuterata. Tom I. Pars IV. Friederichum Fleischer, Lipsiae (Leipzig), viii+762 pp.

    Hammond, P.M. (2000) 16. Coastal Staphylinidae (rove beetles) in the British Isles, with special reference to saltmarshes. In: Sherwood, B.R., Gardiner, B.G. & Harris T. (Eds.), British Saltmarshes. Joint Symposium on British Saltmarshes organized between the Linnean Society of London, the Royal Society for the Protection of Birds and English Nature (2000: London). Forrest Text, Tresaith, Cardigan, Wales, pp. 247–302.

    Hanley, R. S. (2002) A new species of Mexican Tinotus from the refuse piles of Atta ants, including an annotated world catalog of Tinotus (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini). The Coleopterists Bulletin, 56(4), 453–471.

    Hansen, V., Klefbeck, E., Sjöberg, O., Stenius, G. & Strand, A. (1960) Catalogus Coleopterorum Fennoscandiae et Daniae. Entomologiska Sällskapet i Lund, 479 pp.

    Hanski, I. & Koskela, H. (1977) Niche relations among dung-inhabiting beetles. Oecologia, 28, 203–231.

    Hanski, I. & Koskela, H. (1978) Stability, abundance, and niche width in the beetle community inhabiting cow dung. Oikos, 31, 290–298.

    Hatch, M.H. (1957) The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Press, Seattle, ix+384 pp.

    Heer, O. (1839) Fauna coleopterorum Helvetica. Pars I, fasc. 2. Orelii, Fuesslini et Sociorum, Turici [Zurich], pp. 145–360.

    Hoebeke, E.R. (1988) Review of the genus Thamiaraea Thomson in North America (Coleoptera: Staphylinidae: Aleocharinae) with description of a new species. Journal of the New York Entomological Society, 96(1), 16–25.

    Hoebeke, E.R. (1994) Thamiaraea paralira, a new species from North America, and new distributional and habitat data for other Nearctic species of Thamiaraea (Coleoptera: Staphylinidae). Proceedings of the Entomological Society of Washington, 96(1), 1–7.

    ICZN (1957) Opinion 463. Designation under the Plenary Powers of a type species in harmony with accustomed usage for the genus Oxypoda Mannerheim, 1831 (Class Insecta, Order Coleoptera). Bulletin of Zoological Nomenclature, 16(2), 13–24.

    ICZN (1961) Opinion 600. Ischnopoda Stephens, 1835, and Tachyusa Erichson, 1837 (Insecta, Coleoptera); designations of type-species under the plenary powers. Bulletin of Zoological Nomenclature, 18(4), 241–243.

    ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature. xxix + 306 pp.

    Kahlen, M. (1987) Nachtrag zur Käferfauna Tirols. Erganzung zu den bisher erschienenen faunistischen Arbeiten über die Käfer Nordtirols (1950, 1971 und 1976) und Südtirols (1977). Im Selbstverlag des Tiroler Landesmuseum Ferdinandeum, Innsbruck, 288 pp.

    Keys, J.H. (1907) Exaleochara: a genus of Coleoptera new to science. The Entomologist's Monthly Magazine, 43, 102.

    Klimaszewski, J. & Peck, S.B. (1986) A review of the cavernicolous Staphylinidae (Coleoptera) of Eastern North America: Part I. Aleocharinae. Quaestiones Entomologicae, 22(2), 51–113.

    Klimaszewski, J. & Pelletier, G. (2002) Genus Tinotus (Coleoptera: Staphylinidae, Aleocharinae) from America north of Mexico: review of the types, distribution records, and key to species. The Canadian Entomologist, 134, 281–298.

    Klimaszewski, J., Pelletier, G., Germain, C., et al. (2001) Diversity of Placusa (Coleoptera: Staphylinidae, Aleocharinae) in Canada, with descriptions of two new species. The Canadian Entomologist, 133, 1–47.

    Klimaszewski, J. & Winchester, N.N. (with colour images by K. Bolte) (2002) Aleocharine rove beetles (Coleoptera Staphylinidae) of the ancient Sitka spruce forest on Vancouver Island, British Columbia, Canada. Mémoires de la Société royale belge d'Entomologie, 40, 3–126.

    Koch, K. (1968) Käferfauna der Rheinprovinz. Decheniana (Bonn), Beihefte 13, 1–382.

    Kraatz, G. (1856) Naturgeschichte der Insecten Deutschlands. Erste Abteilung. Coleoptera. Bd. 2. Lief. 1–2. Verlag der Nicolaischen Buchhandlung, Berlin, pp. 1–376.

    Kraatz, G. (1857) [New species]. In: Staudinger, O. Reise nachy Island zu entomologische Zwecken unternommen. Entomologische Zeitung. Stettin, 18, 284–287.

    Lohse, G. A. (1969) Vorschläge zur Änderung der Aleocharinensystematik (Coleoptera: Staphylinidae). In: Bericht über die 10. Wanderversammlung deutscher Entomologen, 15. bis 19. September 1965 in Dresden. Deutsche Akademie der Landwirtschaftswissenschaften zu Berlin, Berlin, pp. 169–175.

    Lohse, G.A. (1974a) Tribus: Schistogeniini. In: Freude, H., Harde, K.W. & Lohse, G.A. (Eds.), Die Käfer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, p. 221.

    Lohse, G.A. (1974b) Tribus: Zyrasini. In: Freude, H., Harde, K.W. & Lohse, G.A. (Eds.), Die Käfer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 222–229.

    Lohse, G.A. (1974c) 18. Tribus: Oxypodini. In: Freude, H., Harde, K.W. & Lohse, G.A. (Eds.), Die Käfer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 230–291.

    Lohse, G.A. (1985) Kritische Bemerkungen zur Staphyliniden in “Enumeratio Coleopterorum Fennoscandiae et Daniae” (Helsingfors 1979). Notulae Entomologicae, 65(2), 33–35.

    Lohse, G.A. (1988) Staphylinidenstudien II. Entomologische Blätter, 84(1–2), 41–50.

    Lohse, G.A. (1989) Ergänzungen und Berichtigungen zu Freude-Harde-Lohse “Die Käfer Mitteleuropas” Band 5 (1974), pp. 185–243 In: Lohse, G.A. & Lucht, W.H. (Eds.), Die Käfer Mitteleuropas. 1. Supplementband mit Katalogteil. Goecke & Evers Verlag, Krefeld, pp. 185–243.

    Lohse, G.A. (1991) Corrections to Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 45(1), 20.

    Lohse, G.A., Klimaszewski, J. & Smetana, A. (1990) Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 44(2), 121–202.

    Lohse, G.A. & Smetana, A. (1985) Revision of the types of species of Oxypodini and Athetini (sensu Seevers) described by Mannerheim and Mäklin from North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 39(3), 281–300.

    Lohse, G.A. & Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42(3), 269–278.

    Mäklin, F.G. (1880) Ytterligare diagnoser öfver några nya sibirska Coleopter-arter. Öfversigt af Finska Vetenskaps-Societetens Förhandlingar, 22, 79–86.

    Märkel, F. (1845) Beiträge zur Kenntniss der unter Ameisen lebenden Insekten. Zeitschrift für die Entomologie, 5 (1844), 193–271.

    Mannerheim, C.G. (1830) Précis d’un nouvel arrangement de la famille des Brachélytres, de l’ordre des Insectes Coléoptères. St. Pétersbourg, 87 pp.

    Mannerheim, C.G. (1843) Beitrag zur Kaefer-Fauna der Aleutischen Inseln, der Insel Sitkha. Bulletin de la Société Impériale des Naturalistes de Moscou, 16(1), 175–314.

    Mannerheim, C.G. (1846) Nachtrag zur Kaefer-Fauna der Aleutischen Inseln und der Insel Sitkha. Bulletin de la Société Impériale des Naturalistes de Moscou, 19(2), 501–516.

    Mannerheim, C.G. (1852) Zweiter Nachtrag zur Kaefer-Fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Société Impériale des Naturalistes de Moscou, 25(2), 283–387.

    Mannerheim, C.G. (1853) Dritter Nachtrag zur Kaefer-Fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Société Impériale des Naturalistes de Moscou, 26(3), 95–273.

    Melsheimer, F.E. (1844) Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 2(2), 26–43.

    Moore, I. & Legner, E.F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.

    Mulsant, M.E. & Rey, C. (1871) Histoire Naturelle des Coléoptères de France. Brévipennes. Aléochariens. [Huitième Branche, Bolitocharaires]. Deyrolle Fils, Paris, vi+321 pp., pl. 1–5.

    Mulsant, M.E. & Rey, C. (1873) Description de divers coléoptères brévipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147–189.

    Mulsant, M.E. & Rey, C. (1874a) Tribu des Brévipennes. Famille des Aléochariens. Septième branche: Myrmédoniaires. Annales de la Société d’Agriculture, Histoire Naturelle et Arts Utiles de Lyon, série 4. 6 [1873], 33–738.

    Mulsant, M.E. & Rey, C. (1874b) Histoire Naturelle des Coléoptères de France. Brévipennes. Aléochariens. Suite. Deyrolle, Paris, vi+695 pp., pl. 1–5.

    Muona, J. (1979a) The aleocharine types of Mulsant & Rey (Coleoptera, Staphylinidae) I. Annales Entomologici Fennici, 45(2), 47–58.

    Muona, J. (1979b) Staphylinidae. In: Silfverberg, H., Enumeratio Coleopterorum Fennoscandiae et Daniae. Helsingfors Entomologiska Bytesförening, Helsinki, pp. 14–28.

    Muona, J. (1984) Review of Palearctic Aleocharinae also occuring in North America (Coleoptera: Staphylinidae). Entomologica Scandinavica, 15, 227–231.

    Muona, J. (1987) Some aspects of Aleocharinae systematics - a response to Dr. G.A Lohse. Entomologische Blätter, 83(1), 19–24.

    Newton, A.F., Thayer, M.K., Ashe, J.S. & Chandler, D.S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R.H., Thomas, M.C. (Eds.), American Beetles. Vol.1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272–418.

    Notman, H. (1920) Coleoptera collected at Windsor, Broome Co., N. Y., 26 May to 5 June, 1918, with notes and descriptions. Journal of the New York Entomological Society, 28(2), 178–194.

    Notman, H. (1921) Some new genera and species of Coleoptera collected at Westfield, Chautauqua Co., N. Y. Journal of the New York Entomological Society, 29(3–4), 145–160.

    Notman, H. (1922) New species of Carabidae, Staphylinidae, and Elateridae. Bulletin of the Brooklyn Entomological Society, 17(4), 99–108.

    Palm, T. (1970) Svensk Insektfauna. 9. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Underfam. Aleocharinae (Atheta). Häfte 6. Entomologiska Föreningen i Stockholm, Stockholm, pp. 113–296, pl. I–XXI.

    Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89(2), 443–450.

    Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (II Serie), Sezione Scienze della Vita (A: Biologica), 8, 1–307.

    Pope, R.D. (1977). In: Kloet, G.S. & Hincks, W.D. (Eds) A Check List of British Insects. Second edition (completely revised). Part 3. Coleoptera and Strepsiptera. Handbooks for the Identification of British Insects, 11(3), pp. xiv+105.

    Sahlberg, J. (1876) Enumeratio coleopterorum brachypterorum Fenniae. I. Staphylinidae. Acta Societatis pro Fauna Flora Fennica, 1, 1–248.

    Sahlberg, J. (1880) Bidrag till Nordvestra Sibiriens Insektfauna. Coleoptera. Insamlade under Expeditionerna till obi och Jenessej 1876 och 1877. I. Cicindelidae, Carabidae, Dytiscidae, Hydrophilidae, Gyrinidae, Dryopidae, Georyssidae, Limnichidae, Heteroceridae, Staphylinidae och Micropeplidae. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 17(4), 1–115.

    Samouelle, G. (1819) The entomologist's useful compendium; or an introduction to the knowledge of British insects, comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linné, and the modern method of arranging the classes Crustacea, Myriapoda, spiders, mites, and insects from their affinities and structure, according to the views of Dr. Leach. Also an explanation of the terms used in entomology; a calendar of the times of appearance, and usual situations of near 3000 species of British insects; with instructions for collecting and fitting up objects for the microscope. Thomas Boys, London, 496 pp.

    Sawada, K. (1982) Atheta and its allies of Southeast Asia (Coleoptera; Staphylinidae). III. Oriental species described by V. von Motschulsky and G. Kraatz. Contributions from the Biological Laboratory, Kyoto University, 26(2), 141–187.

    Scheerpeltz, O. (1968) Coleoptera-Staphylinidae. Catalogus Faunae Austriae, 15 fa. Springer- Verlag, Wien, 279 pp.

    Scriba, W. (1870) 27. Leptusa tricolor Scriba nov. sp. In: Heyden, L. von, Entomologische Reise nach dem südlichen Spanien, der Sierra Guadarrama und Sierra Morena, Portugal und den Cantabrischen Gebirgen… Entomologischer Verein in Berlin, Berlin, p. 75.

    Seevers, C.H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp.

    Segers, R. (1986) Catalogus Staphylinidarum Belgicae (Coleoptera). Institut Royal des Sciences Naturelles de Belgique. Documents de Travail., No. 32, Bruxelles, 104 pp.

    Sharp, D. (1883) Pp. 145–312 [Staphylinidae]. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part II. Taylor & Francis, London, 824 pp.

    Sharp, D. (1887) Pp. 673–824 [Staphylinidae]. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part II. Taylor & Francis, London, 824 pp.

    Silfverberg, H. (1988) Lists of the insect types in the Zoological Museum, University of Helsinki. 8. Coleoptera: Staphyliniade. Acta Entomologica Fennica, 52, 15–40.

    Silfverberg, H. (1992) Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae. Helsingin Hyönteisvaihtoyhdistys, Helsinki, v+94 pp.

    Stephens, J.F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, time of appearance, localities, food, and economy, as far as practicable. Mandibulata, 5. Baldwin and Cradock, London, 1–447.

    Strand, A. & Vik, A. (1964) Die Genitalorgane der nordischen Arten der Gattung Atheta Thoms. (Col., Staphylinidae). Norsk Entomologisk Tidsskrift, 12(5–8), 327–335, Taf. I–XXI.

    Strand, A. & Vik, A. (1966) Die Genitalorgane der nordischen Arten der Gattung Oxypoda Mannh. (Col., Staphylinidae). Norsk Entomologisk Tidsskrift, 13, 169–175.

    Thomson, C.G. (1856) Några nya arter af Insekt-slägtet Homalota. Öfversigt af Kongl. Vetenskaps- Akademiens Förhandlingar, 13(4), 91–107.

    Thomson, C.G. (1858) Försök till uppställning af Sveriges Staphyliner. Öfversigt af Kongl. Vetenskaps- Akademiens Förhandlingar, 15, 27–40.

    Thomson, C.G. (1859). Skandinaviens Coleoptera, synoptiskt bearbetade. Tom 1. Berlingska Boktryckeriet, Lund, v+290 pp.

    Thomson, C.G. (1867) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom 9. Lundbergska Boktryckeriet, Lund, 408 pp.

    Tikhomirova, A.L. (1973) Morfoekologicheskiye osobennosti i filogenez stafilinid (s katalogom fauny SSSR). Nauka, Moscow, 191 pp.

    Topp, W. (1975) Morphologische Variabilität, Diapause und Entwicklung von Atheta fungi (Grav.) (Col., Staphylinidae). Zoologische Jahrbücher. Abteilung für Systematik, Ökologie und Geographie der Tiere, 102, 101–127.

    Uhlig, M. & Vogel, J. (1981) Zur Staphylinidenfauna der Umgebung von Waren /Müritz (Mecklenburg). Unter besonderer Berücksichtigung der Naturschutzgebiete “Ostufer der Müritz”, “Ostufer des Feisnecksees”, und des Flächennaturdenkmals “Wienpietschseen”. 5. Beitrag zur Faunistik der Staphylinidae (Coleoptera). Mitteilungen aus dem Zoologischen Museum in Berlin, 57(1), 75–168.

    Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25(1), 11–140.

    Zerche, L. (1991) Was ist Oxypoda lividipennis Mannerheim, 1831 (Coleoptera, Staphylinidae). Entomologische Blätter, 87(1–2), 79–82.

    Zetterstedt, J.W. (1828) Fauna Insectorum Lapponica. Pars I. Libraria Schulziana, Hammone, xx+563 pp.